#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on July 12, 2025; 14:59 GMT TreeBASE (cc) 1994-2008 Study reference: Hermsen E., Nixon K., & Crepet W. 2006. The impact of extinct taxa on understanding the early evolution of angiosperm clades: an example incorporating fossil reproductive structures of Saxifragales. Plant Systematics and Evolution, 260: 141-169. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S1585] BEGIN TAXA; TITLE Simultaenous_analysis; DIMENSIONS NTAX=42; TAXLABELS Allonia Altingia Aphanopetalum Boykinia Cercidiphyllum Choristylis Chrysosplenium Corylopsis Crassula Daphniphyllum Disanthus Divisestylus_brevistamineus Divisestylus_longistamineus Dudleya Exbucklandia Haloragis Hamamelis Heuchera Itea Kalanchoe Lambertia Leea Liquidambar Micranthes Microaltingia Myriophyllum Mytilaria Paeonia_fused Peltoboykinia Penthorum Platanus Pterostemon Rhodoleia Ribes Saxifraga Sedum Sullivantia Tarahumara Tetracarpaea Tetracentron Trochodendron Vitis ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M1253] TITLE Appendix_1; LINK TAXA = Simultaenous_analysis; DIMENSIONS NCHAR=45; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7 8 9" MISSING=? GAP= -; CHARSTATELABELS 1 Habit / 'herbaceous (herbs)' 'woody (shrubs or trees)', 2 Plant_gender_distribution / hermaphroditic monoecious dioecious polygamodioecious trioecous gynomonoecious not_a_character_state gynodioecious, 3 'Leaf, type' / 'simple (not compound, incl. leaves with lobed-dissected margins)' pinnately_compound palmately_compound trifoliate, 4 'Leaf, petiole' / 'absent, leaves sessile' 'present, leaves petiolate', 5 'Leaf, petiole attachment' / 'marginal, leaves not peltate' peltate, 6 'Leaf, phyllotaxy' / alternate opposite whorled, 7 'Leaf, major venation pattern' / pinnate palmate trinerved 'uninerved non-pinnate', 8 'Leaf, nodal anatomy' / unilacunar trilacunar pentalacunar multilacunar, 9 'Stomata, type' / anomocytic paracytic laterocytic anisocytic cyclocytic actinocytic, 10 Flower / unisexual bisexual, 11 'Flower, pedicel' / 'absent, flowers sessile' 'present, flowers pedicellate', 12 'Calyx, sepals' / absent present, 13 'Calyx, number of sepals/sepal lobes' / Not_a_character_state 1. 2. 3. 4. 5. 6. 7. 8. 9_or_more, 14 'Calyx, sepal connation' / sepals_free sepals_connate, 15 Corolla / absent present, 16 'Corolla, number of petals' / Not_a_character_state 1. 2. 3. 4. 5. 6. 7. 8. 9_or_more, 17 'Corolla, circinate vernation of petals' / absent present, 18 'Corolla, fusion' / 'petals free (or free beyond region of hypanthium)' petals_fused_into_corolla_tube, 19 'Stamens, epipetalous' / stamens_not_epipetalous stamens_epipetalous, 20 'Stamens, arrangement' / whorls Not_a_character_state spirals decussate_pairs fascicles, 21 'Stamens, position' / opposite_the_sepals 'opposite the petal-alternisepalous' 'opposite the sepals and petals (opp. and alt. the sepals)', 22 'Anthers, fixation' / basifixed dorsifixed, 23 'Anthers, basal pit' / absent present, 24 'Anthers, dehiscence type' / longitudinal_slits valves, 25 'Anthers, number of sporangia' / not_a_character_state '1 per theca (disporangiate)' '2 per theca (tetrasporangiate)', 26 Nectariferous_tissue / absent continuous_ring 'scales-phyllomes (separate structures, not clearly staminodal)' outer_carpel_wall staminodal petal_bases, 27 Ovary_position / superior 'semi-inferior' inferior, 28 'Carpels, number per flower' / Not_a_defined_character_state 1. 2. 3. 4. 5. 6. 7. 8. 9_or_more, 29 'Carpels, stipitation' / 'absent, carpels sessile' 'present, carpels stipitate', 30 'Carpels, fusion' / free connate, 31 Placentation / 'axile-marginal' parietal apical basal, 32 'Ovules, number per carpel' / less_than_one 1. 2. 3. 4. 5. 6. 7. 8. '9 or more (including "many" or "numerous")', 33 Styles / 'absent, stigmas sessile' present, 34 'Styles, fusion' / free fused, 35 Stigmas_decurrent / 'absent, stigmas terminal' 'present, stigmas decurrent', 36 'Stigmas, fusion' / free fused, 37 'Ovules, integuments' / not_a_character_state unitegmic bitegmic, 38 'Ovules, position' / anatropous orthotropous hemitropous, 39 'Fruit, type' / 'dry dehiscent, capsule or follicle' baccate 'dry dehiscent, schizocarpic' dry_indehiscent fleshy_and_drupaceous, 40 'Seeds, ejection' / absent present, 41 'Seeds, winged' / absent present, 42 'Pollen, number of apertures' / inaperturate 1. 2. 3. 4. 5. 6. 7. 8. 9_or_more, 43 'Pollen, ectoaperture shape' / colpus pore, 44 'Wood, vessels' / 'absent, wood lacking vessels' present, 45 Resin_ducts / absent present, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 ] [ . . . . . . . . . . . . ] Allonia ??????????01?115?000200122???????????????30?? Altingia 110100{01}{12}1000--0----0-00{01}201201{01}91010200019111 Aphanopetalum 10010100011141{01}400{01}02{01}102{03}1401011100203003010 Boykinia 000{01}001{123}011151150000001021{12}201091{01}00200003010 Cercidiphyllum 1{12}0101{01}{01}0000--0----?-00{01}20011-091-1-200013010 Choristylis 1{05}0100011{01}1151150000010021{12}201091{01}01100002110 Chrysosplenium 000{01}0{01}100111410----020102{01}{012}2011{678}1000200003010 Corylopsis 10010001{01}111511{15}0000200122{012}2012310002{01}010{34}010 Crassula {01}00{01}01?{01}3111{3456789}{01}1{3456789}0{01}{01}00010220{345}0{01}{01}{123456789}1000200003010 Daphniphyllum 1{23}010{01}011{01}1{01}{23456}{01}0----?-000200{234}{01}1{12}{12}10102{02}400{34}010 Disanthus 10010011110151151000000025{01}2010{56}1000200003010 Divisestylus_brevistamineus ????????011151150000000?212201091001??00030?? Divisestylus_longistamineus ????????0?11511500000????12201091001??00030?? Dudleya {01}00001?{01}311151150110201022050{01}{01}?1000200003010 Exbucklandia 100100111100--{01}{2345}0000-0011{02}1201{01}{5678}10102000{01}3010 Haloragis {01}00{01}0{01}{03}?0111{234}1140000{01}000202{234}0122100020300{456}010 Hamamelis 10010001{01}10141141000200114120121101020010{34}010 Heuchera 0001001101115115000001102{01}{12}{23}01{12}91000200003010 Itea 100100011{01}1151150000010021{01}201091{01}01200002110 Kalanchoe {01}0{01}{01}0{12}0{01}311141140110{02}0102204010?1000200003010 Lambertia 100{01}0{02}?1110????????0?0002{012}010-221-0-210013110 Leea {01}0{013}10{01}{02}1{045}111{45}11{45}01101100200{3456789}01311101201003010 Liquidambar 1101001{12}1000--0----0-00{01}201{123456}01{01}91010200019111 Micranthes 000{01}00?10111511{578}00002010211{25}01091000100003010 Microaltingia ?????????000--0----??????{02}12010{89}1000?0000301? Myriophyllum 0{012}0{01}0{012}{03}?0{01}{01}1{234}1{01}{234}0000{01}000202{234}0121100020200{456}010 Mytilaria 10010013110151150010?001201201{01}61010200003011 Paeonia_fused {01}0{13}100{02}20111501{56789}000{24}-0?02{12}0{2345}00090-00200003010 Peltoboykinia 000{01}101{13}0111511500002010211201091{01}00200003010 Penthorum {01}00100000111{678}1{01}{1234567}00002010201{5678}01091000200003010 Platanus 110100{01}3{012}00{01}{123456}{01}{01}{3456}0{01}000001200{3456789}000{12}1010213003010 Pterostemon 1001000101115115000021002025010{456}1101200003010 Rhodoleia 100100{02}1110{01}?1{01}{12345}000000012212011910002000{01}3010 Ribes 1{02}0100110{01}11{45}1{01}{45}0000010021{12}20119110{01}20100{89}110 Saxifraga 0{07}010{01}1{0123}0{01}11{45}11{45}0000201021{012}201091000200003010 Sedum {01}{23}0{01}0{012}?{01}3{01}11{456789}{01}{01}{456789}0{01}{01}0{02}010220{45}{01}{01}{01}{123456789}10002000{01}{234}010 Sullivantia 000100110111511500000010211201091000200013010 Tarahumara ?????????01??????????????0240121100020200??1? Tetracarpaea 10010000011140140000{02}000200{45}1{01}{01}91000200013010 Tetracentron 10010011210????????3-0012314010{456}1010200013000 Trochodendron 1001000{0123}211????????2-001231{456789}01091010200013000 Vitis 1{34}{02}{01}001{123}0{01}11511501001100{12}{12}02013{12}1101201003010 ; END;