#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on July 12, 2025; 8:35 GMT TreeBASE (cc) 1994-2008 Study reference: Cossard G., Sannier J., Sauquet H., Damerval C., Ronse de craene L., Jabbour F., & Nadot S. 2016. Subfamilial and tribal relationships of Ranunculaceae: evidence from eight molecular markers. Plant Systematics and Evolution, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S18509] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=31; TAXLABELS Aconitum_anthora Aconitum_lycoctonum Aconitum_napellus Actaea_cordifolia Actaea_racemosa Anemone_blanda Anemone_bracteata Anemone_nemorosa Anemone_pulsatilla Anemonella_thalictroides Aquilegia_atrata Aquilegia_chrysantha Aquilegia_ecalcarata Callianthemum_anemonoides Clematis_apiifolia Clematis_terniflora Clematis_vitalba Consolida_regalis Coptis_quinquefolia Delphinium_cheilanthum Delphinium_pictum Dichocarpum_stoloniferum Ficaria_verna Isopyrum_thalictroides Nigella_damascena Ranunculus_repens Semiaquilegia_adoxoides Thalictrum_aquilegiifolium Thalictrum_minus Trollius_chinensis Trollius_europaeus ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M34694] TITLE Ranunculaceae_CYC2; LINK TAXA = Taxa; DIMENSIONS NCHAR=348; FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Aconitum_anthora GAGACTTCCCCTTGAAATCGCCCGCAGGTTTTTCGATCTTCAAGATAAGCTTGGCTTCGATAAGGCTAGTAA---AACTGTTGAGTGGCTGATGAATAAATCGGGAGAAGCGATTGATGAAGTCTATAAAGA-C-GGCCGGACCGGAGATAACAAGAGCTCA------ACTTCGACTTCTGAGTGTGATGCGGTGTCTGAAAATAAGGATCTGGGGCAA-AAGAATTTCTCTGAA----GCTGAGAAA------ACCAAA----------------------------------------ACTGTTAGAACTGCGCCATATCAGCTTGATAGGGATTCGAGGGAGAAG Aconitum_lycoctonum GAGACTTCCCCTTGAAATCGCCCGCAGGTTTTTCGATCTTCAAGATAAGCTTGGCTTCGATAAGGCTAGTAA---AACTGTTGAGTGGCTGATGAATAAATCGGGAGAAGCGATTGATGAAGTCTATAAAGA-C-GGCCGGACCGGAGATAACAAGAGCTCA------ACTTCTACTTCTGAGTGTGATGCGGTGTCTGAAAATAAGGATCTGGGGCAA-AAGGATTTCTCTGAA----GCTGACAAA------GCCAAA----------------------------------------ACTGTTAGAACTGCGCCATATCAGCTTGATAGGGATTCGAGGGAGAAG Aconitum_napellus GAGACTTCCCCTTGAAATCGCTCGCAGGTTTTTCGATCTTCAGGATAAGCTTGGTTTCCACAAGGCTAGTAA---AACTGTTGACTGGTTAATGAATAAATCGGGAGAAGCGATTGATGAGGTCTATAAAGG-C-GGCAGCACCGGAGATAACAAGAGCCCA------TCTTCGGTTTCAGAGTGTGATGTGGTGTCTGAAAATAAGGATGAGGGGCAA-AAGAGTTTCTCTAAA----GCTGAGAAA------ATGAAA----------------------------------------ACCATTAGAACTACGCCAAATCAGCTTGATAGGAATTCGAGGGAGAAG Actaea_cordifolia GAGACTCTCCCTTGAAGTCGCACGCAAGTTCTTTGATCTTCAAGATATGCTTGGTTTTGATAAAGCCAGTAA---GACCGTTGACTGGTTGTTGACGAAGTCGGAGGAAGCCATCAAGGAGCTCAGTAAA-----GGGTGCACTGGTGACGCCAAGAGTACTATA---TCTTCAACTTCTGAATGTGACGTGGTATCGGATATTGAT---GTGGGGAAG-AAAGGCTACTCCAAA----GGGAAATCT------TTAGATGGCTTTGTTTCG----AGAATGAAAATAATGAAGAAATCGTCGTGTTACACTACATCTCATCTTCTTGCAAGAGATTCGAGGCTGAAG Actaea_racemosa GAGACTCTCCCTTGAAGTCGCTCGCAAGTTCTTTGATCTTCAAGATATGCTTGGTTTTGATAAAGCCAGTAA---GACTGTTGACTGGTTGTTGACGAAGTCGGAGGAACCCATCAAGGAGCTCAGTAAA-----GGGTGCACTGATGACACCAAGAGTACTATA---TCTTTAACTTCTGAATGTGACGTAGTATCGAAAATTGAT---GAGGGGAAG-AAAGACTACTCCAAG----GGGAAATCT------TTAGATGGCTTTGTTTCG----AGAAAGAAAATAATGAAGAAACCGTCGTGTTACACTACATCTCATCTGCTTGCAAGAGATTCGAGGCTGAAG Anemone_blanda GAGACTTTCCCTTGAAATTGCTCGCAAATTCTTTAATCTTCAAGATATGCTTGGTTTTGATATAGCAAGTAA---GACCGTTGAGTGGTTGCTAAAGCAGTCCGAGCAAGCAATTCAAGAGCTTAGTCCA----TCGTGTACTAGCAAAACCAACAATGAGGCTTCTTCTTCTATTTTTGAATTCGGTGACGCCTCTAAAATTGCC-------------------------------------------------------TTGAGTGTGCCA---AAGAGCATGATGGCGCGGTCA------------------------------AGTAGAGGTTTGAGGGTACAA Anemone_bracteata GAGACTGTCACTTGATATTGCTCGAAAGTTCTTTAATCTTCAAGATATGCTTGGATTTGATATAGCAAGTAA---GACTGTCGAGTGGCTGCTGAAGCAATCGGAGGAAGCTATTCATGCGCTTAGTAAA----TCGTGCACTGACAAAATCAAGAGTGCTGTTTCTTCTTCTACTTCTGAATGTGGTAAAGCTGAT------------TTGGGACAC-AATCGT------------------------------------------GTGTCA---AAGGGCATGATGGTGAATAAATCGCCATCAAGTAGAGAT------------GCACGGGATTTGAGGGTACAG Anemone_nemorosa GAGACTTTCCCTTGAAATTGCTCGCAAATTCTTTAATCTTCAAGATATGCTTGGTTTTGATATAGCAAGTAA---GACTGTTGAGTGGTTGCTGAAGCAGTCCGAGCAAGCAATTCAAGAGCTTATTACA----TCGTGTACTAGCAAAACCAACAATGAGGCTTCTTCTTCTATTTCTGAATGCGGTGAGGCCTCTAAAATTGAC-------------------------------------------------------TTGAGTGTGCCA---AAGAGCATGATGGCGCGGTCA------------------------------AGTAGAGATTTGAGGGTACAA Anemone_pulsatilla TAGACTGTCACTTGATATTGCTCGCAGGTTTTTTAATCTTCAAGATATGCTTGGATTTGATATAGCAAGTAG---GACTGTTGAGTGGTTGCTGAAGCAGTCGGAGGAAGCCATTGATGAGCTTCGTAAA----TCGTGCATTGGCAAAACCAGGACTGGT------------------GAAGGTGGTGAAGCCTCTAAAACTGGT---TTGGGACAC-AATGGT------------------------------------------GTGTCG---AAGGGCATGACAGTAAATAAATCGCTCTCA---------------------AGTAGAGATTTGAGGGTACAA Anemonella_thalictroides GAGACTCTCCCTTGAAATCGCTCGCAAGTTCTTTGATCTGCAAGACATGCTTGGTTTTGACAAGGCAAGTAA---AACTGTTGAATGGTTGCTGGGGAAGTCGGAAGAAGCAATTAATGAGGTTAGTAAA----GGGTGCACTGGTGACACCAAAAGCACAGTATCTTCTTCTACTTCTGAATGTGAGGTAGTGTCAGAAATTGAT---GAGGGGCAG-AAAGGC-TTAACTAAA---GGGAAATCT--------GTAGATGGCATTGTTGTG---AGAGAGAAAAGGATAAATAAAATGTTGTGCAGAACTTCATGTCATATTCTCGCGAAAGACTCAAGGGTGAAG Aquilegia_atrata GAGACTCTCCCTTGAAATCGCTCGCAAGTTCTTTGATCTGCAAGACATGCTTGGTTTTGATAAGGCAAGTAA---AACTGTTGAATGGTTGCTGGGGAAGTCAGAAGAAGCAATTAATGAGCTCCATAAA----GGGTGCACTGGTGAAACCAAAAGCACAGTA---TCTTCTACTTCTGAATGTGATGTAGTGTCAGAAATTGAT---GAGGGGCAG-AAAGGC-TTAACTAAAATTGATGAGAAGCAATCT--ATAGATGGCATTGTTTTG---AGAGAGAAAACGACAACTAAAATGTTGAGCAGAACTACATGTCATATCCTTGCAAAAGACTCAAGGGTGAAG Aquilegia_chrysantha -AGACTCTCCCTTGAAATCGCTCGCAAGTTCTTTGATCTGCAAGACATGCTTGGTTTTGATAAGGCAAGTAA---AACTGTTGAATGGTTGCTGGGGAAGTCAGAAGAAGCAATTAATGAGCTCCATAAA----GGGTGCACTGGTGAAACCAAAAGCACAGTA---TCTTCTACTTCTGAATGTGATGTAGTGTCAGAAATTGAT---GAGGGGCAG-AAAGGC-TTAACTAAAATTGATGAGAAGCAATCT--ATAGATGGCATTGTTTTG---AGAGAGAAAACGACAACTAAAATGTTGAGCAGAACTACATGTCATATCCTTGCAAAAGACTCAAGGGTGAAG Aquilegia_ecalcarata -------------------------------------------------CTTGGTTTTGATAAGGCAAGTAA---AACTGTTGAATGGTTGCTGGGGAAGTCAGAAGAAGCAATTAATGAGCTCCGTAAA----GGGTGCACTGGTGAAACCAAAAGCACAGTA---TCTTCTACTTCTGAATGTGATGTAGTGTCAGAAATTGAT---GAGGGGAAG-AAAGGC-TTAACTAAAATTGATGAGAAGCAATCT--ATAGATGGCATTGTTTTG---AGAGAGAAAACGACAACTAAAATGTTGAGCAGAACTACATGTCATATCCTTGCAAAAGACTCAAGGGTGAAG Callianthemum_anemonoides GAGACTTTCAATTGAAATCGCCCGCAAGTTCTTTGATCTTCAAGACATTCTTGGTTTCGATCAGGCTAGTAA---AACTGTTGGGTGGTTGCTGATGAAGTCGGAGGAGGCAATCAAGGGGCTCACTAAA----GGGTTCACTGCTGACACCAAGAGCAATGTA---TGTTCTACTTCTGAATGTGGTTATGTGTCGGAAATTGAT---GAGGGGCGT-AAA----------------CGCGCAAGA--------GAGGAACTTCTTCTAGCT---GAGAATACAATGATGAACAAAACGACATGTAGGACT---------------ACAAGAGATTTGAGGGTGAAG Clematis_apiifolia GAGACTGTCTCTTGAAATTGCTCGCAAGTTCTTTAATCTTCAAGATATGCTTGGTTTTGGCATAGCAAGCAA---GACTGTTGAGTGGTTGTTGAAGCAGTCGGAGGAAGCGATTGATGAGCTCAAACAA----GCTTGTGCTATCGAAACCAAGATTCCTTTTTCTTCTCCTACTCCTGAATGTGGTCAGGTTTCAGGACACAAG------GGTGTG-AAA----------------GGTAAGGAT--------TTGGATGGCAATGTGTCA---AAATGGATGCCGGTGAATAAGGCTCCGTCAGGTAGAGAT------------GCAAGAGATTTGAGGGTAAAG Clematis_terniflora AAGACTGTCTCTTGAAATTGCTCGCAAGTTCTTTAATCTTCAAGATATGCTTGGTTTCGGCATAGCAAGCAA---GACTGTTGAGTGGTTGTTGAAGCAGTCGGAGGCAGCGATTGATGAGCTCAAACAT----GCTTGTACTAGCGAAACCAAGACTGCTTTTTCTTCTTTTACTCCTGAATGTTGCCAGGTTTCAGGGCACAAG------GGTGTG-AAA----------------GGTAAGGAT--------TTGGATGGCAATGTGTCA---AAATGGATACCGGTGAATAAGGCTCCGTCAGGTAGAGAT------------GCAAGAAATTTGAGAGTCCAG Clematis_vitalba GAGACTGTCTCTTGAAATTGCTCGCGAGTTCTTTAATCTTCAAGATATGCTTGGTTTTGGCATAGCAAGCAA---GACTGTTGAGTGGTTGTTGAAGCAGTCGGAGGCAGCGATTGATGAGCTCAAACAA----GCTTGTGCTATTGAAACCAAGACTCCTTTTTCTTTACCAACTCCTGAATGTGGTCAGGTTTCAGGACACAAG------GGTGTG-AAA----------------GGTAAGGAC--------TTGGATGGCAATGTGTCA---AAATGGATGCCGGTGAATAAGGCTCCGTCAGGTAGAGAT------------GCAAGAGATTTGAGGGTACAG Consolida_regalis GAGACTGTCTCTTGAAATCGCTCGAAGATTTTTCGGTCTTCAAGATAAACTCGGTTTCAGCAAGGCTAGTAA---AACTGTTGAGTGGTTGCTGAACAAATCGGAAGAAGCCATCAATGAGATCAATAAAGGA-AGCAGTTCAATTGATAACAAGAGCCCT------TCTTCTGTTTCCGAGTGTGATGTAGTGTCTGAC---------GAAGGGCAA-AAGGGT--TTGTCTCGA--GTTGAGAAC--------ATGAAA---------------------------------------ATGCTTACAACTATGTCAAATCTGCGTGCTAAGGAAGCTAGGGAGGAG Coptis_quinquefolia -------------------------------------------------------------------------------------------------------------------CATGACATTGTGGAA----CGATGTACTGGCGATACCAAGAGTGTA------TCTTGTAGTACTGAATGTGATGTGGTTTCTGAAATTGAT---GAGGGACAG-AACGTC--ATCTTTAAA--GCGAAA-TCT-------TTAGACCGTGCTGTTTCG---AGAGAGAAAACAAAGAACAAATCCTCATGTCGGACTACATTGTATCCTGTTGCAAGAGATTCAAGGGCGAAG Delphinium_cheilanthum GAGACTGTCCCTTGGAATCGCTCGCAGGTTTTTCAATCTTCAAGATAAGCTTGGTTTCGACAAGGCTAGTAA---AACTGTTGAGTGGTTGCTGAATAAATCGGAAGGAGCGATTGATGAGATCAACAAAGGT----------------------------------------GTTTCTGAGTGTGATGTAGTGTCTGAAAATAAG---GAGGGGCAA-AAGGGT--TTCTCTAGA--GTTGAGAAG--------ATGAAA---------------------------------------ACAGCTACACCAACATCATATATGCGTGTTAGGGAAGCAAGGGAGAAG Delphinium_pictum GAGACTTCCCCTAGAAATTGCACGCAAGTTTTTCGATCTTCAAGACATGCTTGGCTTTGATAAGGTTAGTAA---AACTGTTGAGTGGTTGCTGCATAAGTCAGAAGATGCGATAGGGAAGATCAGTAAGGGT-GTTAGTACTGGTGATAACAAGAGCAAA------TCTTCTATGGCCGAAAGTGATGTAGTATCTGAAAAT------GAGGGGCAA-AAACGT--TTCTCTAAA--GTTGAGAAA--------GTGAAA---------------------------------------ACATTCAGGACTATGTCATATCCACTTGATAGGGATTCAAGAAAGAAG Dichocarpum_stoloniferum GAGACTCTCCCTTGGAATCGCTCGCAAGTTCTTTGATCTGCAAGACATGCTTGGTTTTGATAAGGCAAGTAA---AACTGTTGATTGGTTGCTGGGGAAGTCGGAACAAGCAATTACTGAGCTCAGTAAA----GGGTACACTGGTGACGCCAAAAGCACAGTA---TCTTCAACTTCTGAATGTGATGTAGTGTCGGAAATTGAT---GAGGGGCAG-AAACAC--TTAACTAAA--GGGAAA-TCT-------GTAGATGGCATTGTTTTG---AGAGAGAAATCGATAAATAAAATGTTGTGTACAACTACCTGTCATACCCTTGCTAAAGACTCAAGGGTGAAG Ficaria_verna GAGACTTTCCCTAGAAATCGCTCGTAAGTTTTTTGATCTTCAAGATATGCTTGGTTTTGATAAAGCAAGTAA---AACTGTTGAGTGGTTGTTGAGACATTCGGAGGAAGCCATTAATGAGCTTAGTAAG----GGGTGCACAGGCGACACAAAGAGTACTCTG---TCTTCTATTATGGAATGTGAGCTAGGGTCGGAAATTGCC---AGAGGAAAG-AAA----------------CGGAAA-TCC-------CTTGATGCTAATGTATCA---ATTGAGAAAAAGATGAGTAAATCGCCATCAAGTGGTGCT------------ACAAGAGATTCAAGAGTAAAG Isopyrum_thalictroides GAGACTCTCCCTTGAAATTGCTCGCAAGTTCTTTGATCTGCAAGACATGCTTGGGTTCGACAAGGCAAGTAA---AACTGTTGAATGGTTGCTGGGGAAGTCGGAACAAGCAATTAATGAGCTCAGGAAA----GGGTGCACTGGTGACACCAAAAGCACAGTA---TCTTCTACTTCTGATTGTGATGTAGTGTCGGAAATTGAC---GAGGGGAAG-AAAGGC--TTAACTAAA--GGGAAA-TCT-------ATAGATGGCATTGTTTTG---AGAGAGAAAACGACAAATAAAATGTTGTGCAGAACTACATGCCATATCCTTGCAAAAGACTCAAGGGTGAAG Nigella_damascena GAGGCTCTCCCTTGAAATTGCTCGCGAGTTCTTTAATCTTCAAGATTTGCTTGGTTACGACAAAGCAAGCAA---AACTGTTGACTGGTTATTGAGGTTATCTGAAGAAGCCATCAACCAGATTAGTAAA----GCTTGCCCTGCTGAGACCAAGAGTGCTAAA---TCTTCTACTTCTGAATGTAATGTAGTGTCCCATATTAAT---GAGACGAAG-CAAAAA-CTCCCAT-----GGGAAGAAG--------CCTCTAGATGTGGTTGCCTCCAGAGAAAAA---------------------------------------------AATGATTCCAGACTGAAA Ranunculus_repens ----------------------------------------------------------GATAAAGCAAGTAA---AACTGTTGATTGGTTGTTGAGGCAGTCAGAGGAAGCGATTAATGAGCTTACGAAA----GGGTTTGTG------------------------TCTTCTATTTCTGAATGTGAACTAGTGTCAGAAATTAAA---GACATGAAA-CGG----------------AAGATG-TCT-------ATTGATGGAAATGTTGCA---AGCCAGAAAAAGATGAATAAATTGGAAGCAAGTGGAACT------------ACAAGGGATTCCAGGATGAAG Semiaquilegia_adoxoides GAGACTCTCCCTTGAAATCGCTCGCAAGTTCTTTGATCTGCAAGACATGCTTGGTTTTGATAAGGCAAGTAA---AACTGTTGAATGGTTGCTGGGGAAGTCAGAAGAAGCAATTAATGAGCTCCGTAAA----GGGTGCACTGGTGAAACCAAAAGCACAGTA---TCTTCTACTTCTGAATGTGATGTAGTGTCAGAAATTGAT---GAGGGGCAG-AAAGGC--TTAACTAAAATTGATGAGAAGCAATCT-ATAGATGGCATTGTTTTG---AGAGAGAAAACGACAACTAAAATGTTGAGCAGAACTACATGTCATATCCTTGCAAAAGACTCAAGGGTGAAG Thalictrum_aquilegiifolium -------------------------AAGTTCTTTGATTTGCAAGACATGCTTGGTTTCGATAAGGCAAGTAA---AACTATTGAATGGTTGCTGGGGAAGTCGGAAGAAGCAATTAATGAGCTCAGTAAA----GGGTGCACTGGTGACAACAAAAGTGCAGTA---TCTTCTACTTCTGAATGTGATGTAGTGTCGGAAATTGAC---AAGGGGGAG-AAAGGC--TTAACTAAA--GGGAAA-TCT-------GTAGATGGCATTGTTGTT---AGAGAAAAAAGGACAAATCAAATGTTGTGCAAAACTACATGCCTTATTCTTGCAAAAGACTCAAGGGTGAAG Thalictrum_minus -------------------CGCTCGCAGTTCTTTGATCTGCAAGACATGCTTGGTTTCGATAAGGCAAGTAA---AACTATTGAATGGTTGCTGGGGAAGTCGGAAGAAGCAATTAATGAGCTCAGTAAA----GGCTGCACTGGTGACACCAAAAGTACAGTA---TCTTCTACTTCTGAATGTGATGTAGTGTCGGAAATTGAT---GAGGGGCAG-AAAGGC--TTAACTAAA--GGGAAA-TCT-------GTAGATGGCATTGTTGTT---GGAGAGAAAAGGATAAATGAAATGTTGTGCAGAACTACATGTCATATTCTTGCAAAAGACTCAAGGGTGAAG Trollius_chinensis GAGACTGTCCGTTGAAATTTCCCGCAAGTTCTTTGATCTTCAAGATATGCTTGGTTTTGATAAAGCAAGTAA---GACTGTTGAGTGGTTGATGAAGCAGTCAGAGCAAGCTATCGGTGAGCTAAGAAAAGAT-AGCAGCACTGGTGAT---------------------------------------GTAGTTTCAGAAATTGAT---GAGGGGAAA-AAGGGA--TTCTCCTCA--AGGAAA-TCC-------TTAGAC------------------------GTTTCAAAAAAAACGTCGTGCAAAACT---------ATTCTTGCAGGGGATTTGAGGGTAAAG Trollius_europaeus GAGACTGTCCGTTGAAATTTCCCGCAAGTTCTTTGATCTTCAGGATATGCTTGGTTTTGATAGAGCAAGTAA---GACTGTTGAGTGGTTGATGAAGCAGTCGGAGCAAGCTATCGGTGAGCTAAGAAAAGAT-GGCAGCACTGGTGACACCAAGACA------------------------------ACAGTTTCAGAAATTGAC---GATGGGAAA-AAAGGA--TGCTCCTCA--AGGAAA-TCC-------TTGGAC------------------------GTTTCAAAAAAAACGTCGTGCAAAACT---------TTTCTTGCAAGGGATTTGAGGGTAAAG ; END;