NEXUS
[File created by TreeBASE:  1/16/09  15:38:35]
[Matrix accession#: M4143]

BEGIN TAXA;
	TITLE M4143;
	DIMENSIONS  NTAX=71;
	TAXLABELS
		Anthaenantia_lanata	Panicum_repens	Echinochloa_pyramidalis	Urochloa_brizantha	Digitaria_abyssinica	Digitaria_aequiglumis	Digitaria_ammophila	Digitaria_angolensis	Digitaria_arenicola	Digitaria_argillacea	Digitaria_argyrostachya	Digitaria_argyrotricha	Digitaria_atra	Digitaria_badia	Digitaria_balansae	Digitaria_bicornis	Digitaria_bonplandii	Digitaria_botryostachya	Digitaria_brownii	Digitaria_californica	Digitaria_catamarcensis	Digitaria_cayoensis	Digitaria_chaseae	Digitaria_ciliaris	Digitaria_coenicola	Digitaria_cognata	Digitaria_curvinervis	Digitaria_debilis	Digitaria_diagonalis	Digitaria_divaricatissima	Digitaria_eggersi	Digitaria_eriantha	Digitaria_eriostachya	Digitaria_fragilis	Digitaria_fuscescens	Digitaria_gazensis	Digitaria_gerdesii	Digitaria_gymnostachys	Digitaria_heterantha	Digitaria_hitchcockii	Digitaria_horizontalis	Digitaria_insularis	Digitaria_junghuhniana	Digitaria_katangensis	Digitaria_laxa	Digitaria_lecardii	Digitaria_leiantha	Digitaria_maitlandii	Digitaria_mariannensis	Digitaria_melanochila	Digitaria_minutiflora	Digitaria_mollicoma	Digitaria_monodactyla	Digitaria_nitens	Digitaria_nodosa	Digitaria_parviflora	Digitaria_patens	Digitaria_pearsonii	Digitaria_pennata	Digitaria_perrottetti	Digitaria_phaeotrix	Digitaria_pittieri	Digitaria_pubiflora	Digitaria_sacchariflora	Digitaria_sanguinalis	Digitaria_similis	Digitaria_swalleniana	Digitaria_tenuis	Digitaria_ternata	Digitaria_thouaresiana	Digitaria_violascens
	;
END;

BEGIN CHARACTERS;
	TITLE M4143;
	DIMENSIONS NCHAR=113;
	FORMAT SYMBOLS = " 0 1 2 3 " MISSING = ? GAP = -  ;
	CHARSTATELABELS
		1 Inflorescence_primary_branches, 2 Depth_of_epidermal_cell_indentations, 3 Epidermal_cell_width, 4 Spikelet_length_mm, 5 'Spikelet-width mm', 6 Anther_length_mm, 7 'Spacer, uninformative', 8 'Spikelet, hair apex' / not_dilated slightly_dilated 'dilated - claviform hairs', 9 'Spikelet, hair apex' / acute 'apiculate-rounded' rounded, 10 'Spikelet, hair walls' / smooth verrucose, 11 'Spikelet, hair apex' / straight recurved, 12 'Spikelet, hair length' / up_to_2_mm '3-5 mm', 13 'Spikelet, hair pigmentation' / 'silver-white' isabelline brownish, 14 'Spikelet, purple-tinged hairs' / wanting present, 15 'Spikelet, first rachilla internode' / elongated inconspicuous, 16 'Spikelet, second rachilla internode' / elongated inconspicuous, 17 Spikelet_position / Upper_lemma_abaxial Upper_lemma_adaxial, 18 Spikelet_compression / biconvex 'plano-convex' dorsiventrally_flattened, 19 'Lower glume, whether present or not' / wanting present, 20 'Lower glume, shape' / 'truncate, reduced to a rim' 'ovate, acute' obtuse, 21 'Lower glume, margins' / not_clasping clasping_at_the_base 'clasping, with overlapping margins', 22 Lower_glume_texture / membranaceous hyaline, 23 'Lower glume, number of nerves' / 'many-nerved' '3-nerved' without_nerves, 24 'Upper glume, number of lateral veins on each glume half' / 0. 1. 2. 3_or_more, 25 'Upper glume, apex' / acuminate acute obtuse, 26 'Upper glume, relative length' / obsolete shorter_than_the_lower_lemma 'sub-equal to the lower lemma' longer_than_the_lower_lemma, 27 'Upper glume, relative width' / 'wider than the upper floret, the glume covers the upper floret sides' 'narrower than the upper floret (upper floret laterally visible)', 28 Upper_glume_indumentum / wanting present, 29 'Upper glume, distribution of indumentum' / pilose_throughout 'alternate, inner internervial spaces pilose' 'pilose between margins and outer veins, otherwise glabrous' 'alternate, inner internervial spaces glabrous', 30 'Upper glume, scabrousness of nerves' / smooth scabrous, 31 'Upper glume, distribution of nerves' / equidistant 'unequidistant, lateral nerves contiguous', 32 'Upper glume, distal convergence of lateral nerves' / not_convergent convergent, 33 'Lower lemma, number of lateral veins on each lemma half' / 1. 2. 3. 4_or_more, 34 'Lower lemma, apex' / acuminate acute obtuse apiculate, 35 'Lower lemma, texture' / papyraceous membranaceous hyaline cartilaginous, 36 'Lower lemma, indumentum' / wanting present, 37 'Lower lemma, distribution of indumentum' / 'pilose between margins and outer veins, otherwise glabrous' 'alternate, inner internervial space glabrous' 'inner internervial space glabrous, otherwise pilose' all_internervial_spaces_fringed_with_hairs, 38 'Lower lemma, scabrousness of nerves' / smooth scabrous, 39 'Lower glume, pectinate bristles' / lacking present_at_least_in_pedicellate_spikelets, 40 'Lower lemma, vein distribution' / equidistant 'unequidistant, lateral nerves contiguous' 'unequidistant, three central nerves contiguous', 41 'Lower lemma, distal convergence of lateral veins' / not_convergent convergent, 42 'Lower lemma, axillary flower' / lacking reduced_to_a_palea 'male-fertile', 43 'Upper floret, abscission' / not_occurring occurring, 44 'Upper floret, lemma apex' / closed open, 45 'Upper floret, lemma texture' / membranaceous cartilaginous_with_membranaceous_margins crustaceous, 46 'Upper floret, lemma and palea surface' / smooth papillose, 47 'Upper floret, disposition of lemma margins' / inrolled_around_the_palea folded_upon_the_palea, 48 'Upper floret, short, stiff prickles towards lemma apex' / wanting present, 49 'Upper floret, pigmentation at maturity' / stramineous_to_ochraceous brown dark_purple, 50 'Upper floret, relative length' / longer_than_the_lower_lemma 'sub-equal to the lower lemma' shorter_than_the_lower_lemma, 51 'Upper floret, apex' / acute acuminate obtuse, 52 'Upper floret, nerves of the upper lemma' / 3_nerves_distinguishable 5_nerves_distinguishable 7_nerves_distinguishable, 53 'Flower, pigmentation of anthers' / yellow purplish, 54 'Flower, pigmentation of stygmata' / pale purplish, 55 'Caryopsis, outline' / 'elliptical-oblong' obovate orbicular, 56 'Caryopsis, pigmentation' / pale brown, 57 'Reproduction, cleistogamy' / none occurring, 58 'Inflorescence, distribution of LPc along the main axis' / racemes_always_alternate 'proximal racemes (frequently) verticillate' 'distal racemes conjugate, otherwise alternate', 59 'Inflorescence, relative length of 1 order branches' / decreasing_towards_apex about_the_same_length_throughout 'longer at the middle, infl. fusiform', 60 'Inflorescence, length of racemes relative to main axis' / racemes_not_reaching_the_apex the_lowermost_racemes_almost_reaching_the_apex racemes_overtoping_the_apex, 61 'Inflorescence, orientation of LPc' / ascendent_to_spreading rigidly_horizontal, 62 Inflorescence_abscision_as_tumbleweed_sensu_Watson_and_Dallwitz / not_occuring occuring, 63 'Inflorescence, homogenization' / paniculate 'proximally paniculate, homogenized distally' kPc_homogeneous_throughout, 64 'Inflorescence, main florescence (=terminal spikelet) and distal kPc' / wanting present, 65 'Inflorescence, long paraclades' / wanting present, 66 'Inflorescence, phylotaxis' / distichous polystichous, 67 'Inflorescence, second order long paraclades' / 'wanting - occasionally a reduced 2 LPc on the proximal raceme' regularly_present, 68 'Inflorescence, pulvini at the base of racemes' / 'inconspicuous - lacking' 'conspicuous, bulky', 69 'Inflorescence, pulvini at the base of pedicells' / lacking present, 70 'Inflorescence, peduncle indumentum' / glabrous distally_hirsute hirsute_throughout, 71 'Inflorescence, rachis of racemes' / trichetrous narrowly_winged, 72 'Inflorescence, rachis indumentum' / glabrous_to_scabrous hirsute ventrally_pubescente, 73 'Inflorescence, pedicell development' / 'very brief, spikelets subsessile' 'developed, normal' extraordinarily_developed, 74 'Inflorescence, pedicell indumentum' / glabrous 'scabrous to antrorse-echinulate (more-than-scabrous)' puberulous 'hirsute-ciliolate', 75 'Inflorescence, pedicell apex sensu Webster 1983' / truncate cupuliform discoid, 76 'Inflorescence, coronula at the apex of pedicells' / lacking composed_of_short_hairs composed_of_long_cilia, 77 'Inflorescence, the more frequent number of spikelets per node' / one two three_or_more, 78 'Inflorescence, pedicell concrescence' / pedicells_free pedicells_concrescent_with_the_rachis, 79 'Inflorescence, whether sub-sessile or pedunculate racemes' / '(sub)sessile' with_long_peduncles, 80 'Inflorescence, relative length of raceme internodes' / 'short - less than twice the length of the spikelets' 'long -more than twice the length of the spikelets', 81 'Growth form, life cicle' / annual perennial, 82 'Growth form, branching of culms' / unbranched branched, 83 'Growth form, growth direction of culms' / orthotropous_to_geniculate decumbent 'plainly plagiotropous, stoloniferous', 84 'Growth form, internode indumentum' / glabrous distally_hirsute, 85 'Growth form, stypochitum' / wanting present, 86 'Growth form, leaf sequence along tillers' / the_prophyll_is_followed_by_cataphylls the_prophyll_if_followed_by_foliage_leaves, 87 'Leaves, pubescence of cataphylls' / glabrous pubescent, 88 'Growth form, direction of growth in the innovation zone' / orthotropous 'mainly orthotropous, but shortly plagiotropous proximally' plainly_plagiotropous, 89 'Growth form, rhizome internodes' / all_internodes_short 'elongated in the plagiotropous portion, distally shortened' all_elongated, 90 'Growth form, accesory innovation zones along the culm' / lacking present, 91 'Growth form, transition from short to long internodes' / abrupt gradual, 92 'Growth form, culm internodes' / hollow solid, 93 'Leaves, leaf sheath indumentum' / glabrous 'pubescent-pilose', 94 'Leaves, prefoliation' / convolute conduplicate, 95 'Leaves, indumentum of the ligular region' / glabrous 'hirsute-pubescent', 96 'Leaves, ligule outline' / truncate acute obtuse, 97 'Leaves, ligule margin' / entire 'irregular, erose', 98 'Leaves, type of ligule' / entirely_membranous membranous_with_a_ciliate_margin 'ciliate, without a conspicuous membranous portion', 99 'Leaves, ligule development' / 'well developed, conspicuous' reduced_to_an_inconspicuous_ridge, 100 'Leaves, leaf blade outline' / 'narrowly lanceolate, acuminate' lanceolate filiform, 101 'Leaves, leaf blade consistence' / papiraceous coriaceous, 102 'Leaves, mature leaf blade exposition' / plane involute, 103 'Leaf anatomy, midrib' / 'differentiate, distinct' undifferentiate, 104 'Leaf anatomy, adaxial papillae' / lacking present, 105 'Leaf anatomy, adaxial prickles on rib zones' / lacking present, 106 'Leaf anatomy, abaxial prickles on rib zones' / lacking present, 107 'Leaf anatomy, differentation of costal and intercostal adaxial regions' / undifferentiate slightly_noticeable... noteworthy, 108 'Leaf anatomy, differentation of costal and intercostal abaxial regions' / undifferentiate slightly_noticeable... noteworthy, 109 'Leaf anatomy, location of bulliform cells' / only_adaxial both_adaxial_and_abaxial, 110 'Leaf anatomy, colorless parenchyma associated to midrib' / adaxial both_adaxial_and_abaxial none, 111 'Leaf anatomy, macrohairs' / lacking present, 112 'Leaf anatomy, crystals in chlorenchymatous cells' / lacking present, 	113 'Leaf anatomy, abaxial microhairs' / lacking present
	;
	MATRIX
		Anthaenantia_lanata             000000000100????31201(01)0102111?00100010010000(012)011100200001?11000001010?001100011?0000?(01)110?112110110000010000?1?01
		Panicum_repens                  ???????10001220(02)31200?1003110?000021020000(12)0211??00000001?11000001110?01111000002201?0010?11(02)101110210?1001100012
		Echinochloa_pyramidalis         ???????00101120020300?1101000?101020020010121011100000021110000011320100011000001000?(01)100?20000001110000011100001
		Urochloa_brizantha              000000000001(12)20031201?0101100?001120021000101110?001200201001001102200?0011110011010?(01)010?11(01)001011110?0000100002
		Digitaria_abyssinica            ???????111211002(12)2(12)00?0(01)12210?00011001110010001000(01)0100211101000011001000?11001?????01012100100001111001111100001
		Digitaria_aequiglumis           0200000111210012(12)020(01)0011201110001100111102001100111100211100000011001000010001?????0(01)011100000001000101100110001
		Digitaria_ammophila             000000100111210212201000021113001010011110220012001011?211101000011(01)010111(01)0000110000101010000010110(12)1?1000?00001
		Digitaria_angolensis            021000110111001211211000021211002011011100120110000110021110100001(01)102000010001?????00012000100000000000010100001
		Digitaria_arenicola             020000100121110(12)(23)1201101021113000010011110120(01)?0??100012111011000210010111100001??0?0(01)000(01)00000100000000101100001
		Digitaria_argillacea            1100000111210002(12)1(12)11(01)0(01)0211110(01)0010011101100010000100021(01)?01000011102000010001????101010010100001010000011100001
		Digitaria_argyrostachya         2200020111210012111110000111120020100111011001100101100211101001011212100010001????000010100000010100001101100001
		Digitaria_argyrotricha          02100011112(01)0012(12)12010000111110000100111001000000011200201101001010102100011001?????01012000100000010101111100001
		Digitaria_atra                  22000(02)01112(01)0002(12)22010000133(01)?00101001110112???000????0210?0?02??11?110??1001???000??10100000000011211?1?11101001
		Digitaria_badia                 1100020111212012121110000(12)211300(01)0100111011001100?01000211?010000111020001000?0000000(01)010100100101021010011100001
		Digitaria_balansae              220002011121001211111000011112002010011101(01)201100001100211101000011102000?(01)0001????0?0000100000010022000111100001
		Digitaria_bicornis              00000011112110021111100012111101(01)11001110010011001112002(01)1101001011001000011001?????01010000000111100010011001011
		Digitaria_bonplandii            2200020111212012322(01)100002211300211001110110011000(01)01002111010000111120001????????0?11010010010111020000111100001
		Digitaria_botryostachya         02000001112100021111100002111100011001110012000???000002?11000000102010001000001100000010000000001101001111100001
		Digitaria_brownii               00001010112110011120100002011300001001111022001(12)000000021(01)?0000001110100111000(01)1(12)00100010100(01)001?11120?1?00?00(01)?1
		Digitaria_californica           0000(01)0(01)001211002102110000201120010100111102100(01)001010002111000000111010001100001100001011100000101112(01)01000?00001
		Digitaria_catamarcensis         00001100112120021021100002011200101001111021000001010002111000000111010001100001210100002100000101012000010100011
		Digitaria_cayoensis             11000001112100121111100002111100101001110110001???000002111010000112020000100?1????001010100000001011000111101001
		Digitaria_chaseae               22000001112120123121100002111300201001111210000???????02100??0001111020?0000001????001010101011001021001111101001
		Digitaria_ciliaris              0000000111211002111110000211110011100111001001100011(12)002(01)1(01)01001011001000011001???0?0(01)012100100101012000111101001
		Digitaria_coenicola             000000100121110(12)10201000020113000010011110210010001011?211101000011101011110000110000(01)0(01)1100000(01)0(01)(01)(01)2001000?00001
		Digitaria_cognata               0000001011211002112010010211130000100111101000100010101(12)111011000211010111100001000000000100100101102000001101101
		Digitaria_curvinervis           ???????11111000212110?0012110?002110011101000110000?000211?01000011102000010001?????010(01)2000(01)00010010000?10100001
		Digitaria_debilis               020(01)001001211002303013000201110010100111022101100010100211101000011001000011001?????0(01)00(01)000000110100111?11101011
		Digitaria_diagonalis            ???????1112(01)(02)01202110?0?0(01)110?000100011100(01)001000?11000211111000011122000100011?000000000010000001112000111100001
		Digitaria_divaricatissima       000000(01)001212102102110000201110(01)1110011110200010001011121110100001110101111000011000000(01)2100000(01)0110(12)00000(01)?00001
		Digitaria_eggersi               000000100121100211101000021111000010011110110000000000021(01)00000001100100011100011000010110000001000011?1?0??00011
		Digitaria_eriantha              00000001112110021111100102111100001001110010011000112002(01)110100101100100011(02)00(01)100100(01)001100000001112000000100001
		Digitaria_eriostachya           00000011112(01)0002(12)120100(01)0201110010100111021101100?(01)1000211101000011101000?(01)1001????10(01)001100100010012000?1010?011
		Digitaria_fragilis              110000011121000211(12)11000021111000010011101100?10000(01)(01)00211101000011102000010001????1010100000001010110?0111101001
		Digitaria_fuscescens            ???????1112(01)001212200?001(12)110?00210101110100001(02)002?2002(01)1?01001010102100112001?2?11100001001001100(01)1000?11100011
		Digitaria_gazensis              02010011112121123121130002111100001001110212011???01(12)001111110000111010001000001100001011000000011100101010111011
		Digitaria_gerdesii              220002011121000231201000021113000000011111100??000???0021(01)?0?00??111020101000?1?000001010000000011010101111101001
		Digitaria_gymnostachys          0200000011210002020012??02311000001001111012001???10101211101000011101011100001?000011010100000001112001?10100001
		Digitaria_heterantha            000000010121110231211000031112010010011110200110001120?2(01)11010000110010(01)111100???01000010000100000001010110111001
		Digitaria_hitchcockii           0000001001212002112110000111130000100111101000100001000211?000000111010001100001100001010100000101(01)010?10(01)1?00001
		Digitaria_horizontalis          0200000111210002111110000211110000000111101000000011(012)002(01)1101001011001000011001???01010121001000(01)111(12)(01)(01)(01)?(01)(01)(01)00011
		Digitaria_insularis             000011000121(12)00210(12)11000020111001010011110(12)10110010100021110000001110100011000011000010(01)1100000001112(01)0(01)001100011
		Digitaria_junghuhniana          00000001112100021111100002111100111001111012011???01100(12)111(01)1000011101000011001???00010(01)0100100011111?11111111011
		Digitaria_katangensis           ???????1112(01)001212110?0002110?00011001110110000???0100021110100101120200010000??1?0000000110000000022000?0010?001
		Digitaria_laxa                  00000100012120020111120??201101000100111101101010011000211110100011101000110000?200001002100100(01)00112000000?00101
		Digitaria_lecardii              11000001112100121111100002111101101001111112001001010002111010000112120000(01)0001????0010101100000011220?1111101001
		Digitaria_leiantha              ???????11121001212110?0012110?0021100111001001100001100211101000011202000100010100001(01)0001100111110200?0100110001
		Digitaria_maitlandii            22000001112100121211100001211200101001110102011??0010002111010(01)0011202000100011?0000010101012001110211?1111100001
		Digitaria_mariannensis          00000001112(01)?????????????311131(01)011000010010000000112002(01)1101001000000?00011001?????0(01)010100000011122101110101011
		Digitaria_melanochila           22000201112100121111100001111300201001110110001???01000211100001011102100000001????00001010000000002200111110?001
		Digitaria_minutiflora           ???????11120??????010????0120?000100011100(01)000000001000211101000011222000010001???0?0(01)0101102001010220?0111100001
		Digitaria_mollicoma             021000011121001211201000121111000100011100100(01)1??02?2002(01)1?01001010102000111001?2?111(01)00(02)100100001011001111101011
		Digitaria_monodactyla           00000101112100021011100001111101001001110010001???????02100??0001130110?010001??10000(01)0(01)01102011010100?1100110001
		Digitaria_nitens                00000011112100021111100002011300101000111012010???00000211101000023111010100????0000?(01)?000000101100110?010110000?
		Digitaria_nodosa                02000001112100121111100002111300011001110112011??0(01)01002111010000101010001100001100101110100000011101000?11100011
		Digitaria_parviflora            0(02)0000011121110211211000022113000110011111(01)20110000100011110100001(01)10100011000010000000(01)00000000011110001??101001
		Digitaria_patens                0000101011211002102110000201120010100111101200100100100211100000011111001110000110000(01)01000000010110(12)0010(01)(01)?000(01)1
		Digitaria_pearsonii             ???????11121100212110?0002110?00001001110212001???001001111110000110010001?000?120?001010100100010022101111111011
		Digitaria_pennata               020000001121010211201000011113000010011111120000001001121110100011110101111?0001??0?001101000000011000?1000100001
		Digitaria_perrottetti           0200001111210002112010000111120020100111001000100011000211101000011001001011001????001010101100010012000?1010?011
		Digitaria_phaeotrix             1100020111210002111110000211110020100111011000000011(01)0021110100001110200(01)100000(01)000001000110211(01)1(01)020020(01)0010(01)001
		Digitaria_pittieri              00000010012110021111100002111100011001110011000000(01)1100211100000011001000?1(01)00?1??0001012000100(01)01102101000?00001
		Digitaria_pubiflora             02000010112111021120100001111300001001111012000001(01)0001211101100021101011110000110000(01)0(01)01001001010010?0011101001
		Digitaria_sacchariflora         000011000121200211(12)11000020111001010011110110100000100021110000001110100011000011000?0000100000001102000110100001
		Digitaria_sanguinalis           00000001112110021111100002111110111001111012011000112002(01)110100101100100001(01)001????00101(02)100(01)00001101101000100001
		Digitaria_similis               0000110001212002(12)121100002011000001001111021000001010002111000000111010001100001100100001100000001112000001100001
		Digitaria_swalleniana           00001100012110021111100002011000001001111021000000010002111000000111010001100001210001012000000(01)0(01)101101001?00011
		Digitaria_tenuis                000000100121100211211000020111000010011110110000000100021(01)?(01)0000011101000110000110000(01)012100000001102101000100001
		Digitaria_ternata               21000201112(01)0012111110000221130020100111011001100021100211?01011213(12)12000000001????000010100000001021000111111001
		Digitaria_thouaresiana          2200020111210012112010000212130020000111011001100001100211101001011102000000001???0000010000100010021010111110001
		Digitaria_violascens            0(02)100001112(01)0002(12)120100(01)12111100211001110110011000(01)11002(01)1101001011(12)02100002001????00(01)000000100000020000111110001
	;
END;

BEGIN ASSUMPTIONS;
OPTIONS DEFTYPE = ord PolyTcount = MINSTEPS ; 
 END;