NEXUS
[File created by TreeBASE:  7/26/07  21:07:04]
[Matrix accession#: M2111]

BEGIN TAXA;
	TITLE M2111;
	DIMENSIONS  NTAX=69;
	TAXLABELS
		Hillebrandia_sandwicensis	Datisca_glomerata	Datisca_cannabina	Begonia_violifolia	Begonia_thomeana	Begonia_symsanguinea	Begonia_sutherlandii	Begonia_subscutata	Begonia_staudtii	Begonia_sp._Pritzelia	Begonia_socotrana	Begonia_sericoneura	Begonia_scutifolia	Begonia_scapigera	Begonia_samhaensis	Begonia_salaziensis	Begonia_roxburghii	Begonia_quadrialata	Begonia_prismatocarpa	Begonia_potamophila	Begonia_poculifera	Begonia_palmata	Begonia_olbia	Begonia_obliqua	Begonia_nossibea	Begonia_molleri	Begonia_meyerijohannis	Begonia_masoniana	Begonia_mannii	Begonia_mananjebensis	Begonia_malabarica	Begonia_madecassa	Begonia_luxurians	Begonia_loranthoides	Begonia_longipetiolata	Begonia_lobata	Begonia_letouzeyi	Begonia_kisuluana	Begonia_johnstonii	Begonia_iucunda	Begonia_integerrima	Begonia_incarnata	Begonia_horticola	Begonia_holtonis	Begonia_grandis	Begonia_goegoensis	Begonia_geranioides	Begonia_gabonensis	Begonia_francoisii	Begonia_floccifera	Begonia_engleri	Begonia_edmondoi	Begonia_duncanthomasii	Begonia_dregei_ssp_homonyma	Begonia_dregei_partita	Begonia_dregei	Begonia_dipetala	Begonia_dewildei	Begonia_crassirostris	Begonia_convolvulacea	Begonia_cinnabarina	Begonia_capillipes	Begonia_bogneri	Begonia_balansana	Begonia_aspleniifolia	Begonia_annobonensis	Begonia_ankaranensis	Begonia_angularis	Begonia_aequata
	;
END;

BEGIN CHARACTERS;
	TITLE M2111;
	DIMENSIONS NCHAR=45;
	FORMAT SYMBOLS = " 0 1 2 3 4 5 6 " MISSING = ? GAP = - ;
	CHARSTATELABELS
		1 Tubers_derived_from_stem / absent present, 2 Tubers_derived_from_root / absent present, 3 Bulbils_derived_from_buds / absent present, 4 Tuberciles / absent present, 5 Caudex / absent present, 6 Leaf_shape / simple compound, 7 Peltateness / not_peltate peltate, 8 Trichome_ring / absent 'present - hairs' 'present - ridge', 9 Stipule_edge / entire fringed, 10 Fuzzy_hairs / absent present, 11 Stellate_hairs / absent present, 12 Lifestyle / perennial monocarpic, 13 Inflorescence_position / axile terminal 'pseudo-terminal', 14 Sexual_separation / dioecious monoecious, 15 Inflorescence_type / monochasial_cyme dichasial_cyme raceme, 16 Dichasia / symmetric asymmetric, 17 'Flowers - Inflorescence' / less_than_70 over_100, 18 'Inflorescence - sexual separati' / staminate_and_pistillate_interspersed pistillate_basal separate_inflorescences, 19 Flower_size / male_and_female_similar female_distinctly_larger, 20 'Flower colour - base' / white_or_pink yellow red orange, 21 'Flower colour - pattern' / all_one_colour with_red_veins_or_patches tepals_different, 22 Scent / imperceptible strong, 23 Perianth_tube / absent present, 24 'Male - tepal no.' / 2_tepals 4_tepals 0_tepals 10_tepals, 25 Male_tepal_fusion / free partly_fused, 26 Androecium / anthers_face_all_ways anthers_face_2_ways anthers_face_one_way anthers_face_inwards, 27 Stamen_no. / less_than_10 10_or_more, 28 Stamen_colour / yellow orange red, 29 Stamen_fusion / free fused_in_centre fused_at_one_side on_a_column, 30 Female_tepal_no. / 2_tepals 3_tepals 4_tepals 5_tepals 6_tepals 0_tepals 10_tepals, 31 Female_tepal_fusion / free 2_tepals_partly_fused all_tepals_partly_fused, 32 Style_no. / 2_styles 3_styles 4_styles '(5-) 6 (-7) styles' 5_styles, 33 Style_colour / yellow greenish white pink red, 34 Style_fusion / free fused, 35 Ovary_position / inferior 'semi-inferior', 36 Locule_number / 1_locular 2_locular 3_locular 4_locular '5- 6 -7 locular', 37 Placentation / parietal septal axile, 38 'Placentation - dividedness' / '1-fid' '2-fid', 39 Wing_no. / absent 2_wings 3_wings 4_wings ca._6, 40 Wing_symmetry / equal_to_subequal one_larger, 41 'Fruit dry-fleshy' / dry fleshy, 42 Fruit_orientation / upright pendant_to_nodding recurved, 43 Beaked_fruit / absent present, 44 Dehiscence / not_between_styles between_styles, 	45 Bracteole_subtending_ovary / absent 2_bracteoles 3_bracteoles
	;
	MATRIX
		Hillebrandia_sandwicensis         000000010?0001100000??03?0??06?4?01?010???01?
		Datisca_glomerata                 00000100?000?02???0??002?0?005?1?000000?0?010
		Datisca_cannabina                 00000100?000102???0??002?00005?1?000000?0?010
		Begonia_violifolia                0000000000000110001000000?1030010101212101000
		Begonia_thomeana                  00000000100001100001??00021020010102202101000
		Begonia_symsanguinea              000000000000012001020000101213213102212001000
		Begonia_sutherlandii              100?000010000110000{13}0001001003010102202001000
		Begonia_subscutata                00000000001001??0{02}001001030032010102010?1?00?
		Begonia_staudtii                  000000100000010?0001100002103002010320300000?
		Begonia_sp._Pritzelia             000000000000?11???????0???????????0????????0?
		Begonia_socotrana                 001000100000111000000001011004010?0220210100?
		Begonia_sericoneura               000000000100011000000000001002010002212101001
		Begonia_scutifolia                000000101000010?0001100002102002010320300?000
		Begonia_scapigera                 000000101000010?00011000001020020103200?0000?
		Begonia_samhaensis                0?1000100000111000000001001004010002202001001
		Begonia_salaziensis               00000000001001100000??010?101201{02}00???0?1?000
		Begonia_roxburghii                00000000000000?00200010100100?????0???????00?
		Begonia_quadrialata               000000101000010?0001100002103002010320300000?
		Begonia_prismatocarpa             000000001000010?0001100002103002010320{03}01{01}00?
		Begonia_potamophila               00000010100001??0001{01}?000210?00{12}?10{23}2?{23}00000?
		Begonia_poculifera                00000000001001?002001010021020020103010?1?00?
		Begonia_palmata                   000000000000011000000{01}01001033000101212102000
		Begonia_olbia                     000000010000?110???0??0???????????02212?0100?
		Begonia_obliqua                   000000000100011000000001001003010102212101001
		Begonia_nossibea                  00000000?0?001100??0??00?0???2???0022?200?00?
		Begonia_molleri                   000000000010011002000001001032020103010?1?00?
		Begonia_meyerijohannis            0000000000000{01}100200?10100100{02}040005{01}00?1?001
		Begonia_masoniana                 000000000000011000?0000100101101010001210100?
		Begonia_mannii                    000000000010011002001001001{02}12024003110?1?001
		Begonia_mananjebensis             00000000???001100??0??00?0????????022?200?00?
		Begonia_malabarica                000000000000011000000001001011010002212101000
		Begonia_madecassa                 00000000000001100??0??0??????????1022?2101000
		Begonia_luxurians                 000001?10100011010000001001003010002202001001
		Begonia_loranthoides              00000000001001100200?00101102202?1030?0?1?00?
		Begonia_longipetiolata            00000000001001?00200100102102202{04}1030?0?1?000
		Begonia_lobata                    000000000100?11???????0???????????0????????0?
		Begonia_letouzeyi                 000000101000010?0001{01}0000210300201032?301{01}000
		Begonia_kisuluana                 00000000?01001100200?001021?320{12}{04}10{23}0?0?1?00?
		Begonia_johnstonii                000000010000011000000001001013010002212101001
		Begonia_iucunda                   1001000?10?001??0{02}01?00002101001010220210100?
		Begonia_integerrima               000000000000011????0??0100???301?00221210?00?
		Begonia_incarnata                 0000000000000110000000010?1?{01}3011102212101000
		Begonia_horticola                 00000000001001100200?001001?320{12}?10{23}0?0?1?00?
		Begonia_holtonis                  00000000000021101??0000100100?01??022{01}210100?
		Begonia_grandis                   1001000{01}000001100000010100103301010221200?000
		Begonia_goegoensis                000000100000011000002001001032000101202101000
		Begonia_geranioides               0100?00????001100000??01???????1??0220200100?
		Begonia_gabonensis                000000000010011000000001020032010102010?1?000
		Begonia_francoisii                000000000000?1100??0??00??????????022?200?00?
		Begonia_floccifera                000000000000011000000000001032?1?102202??100?
		Begonia_engleri                   00000001000001100000000100101401010221210100{12}
		Begonia_edmondoi                  0?000000???0?11?00?0??010?10?3010?022?2001001
		Begonia_duncanthomasii            000000101000010?000110000210200201032?30?000?
		Begonia_dregei_ssp_homonyma       0?00100000000110000000000?10{01}3010?022?200100?
		Begonia_dregei_partita            0000?00000000110000000000?10?3010?022?200100?
		Begonia_dregei                    0000?0000000011000000000001013010002202001000
		Begonia_dipetala                  000000000000011000000000021010010{01}02202001000
		Begonia_dewildei                  000000101000010?00000000020020020103203000000
		Begonia_crassirostris             00000000000001?000000001001004011002212011000
		Begonia_convolvulacea             000000000000011010000001001003010002202101001
		Begonia_cinnabarina               01000000110001??0??3??0?0?????????02212?0100?
		Begonia_capillipes                000000000010011002000001000?32004101010?1?000
		Begonia_bogneri                   00000000000001??000000010000{23}4010102{02}1{25}111000
		Begonia_balansana                 0000000?1000011000000?0100100203010421401?00?
		Begonia_aspleniifolia             000000001000010?00?010000?00300101022020?1000
		Begonia_annobonensis              00000001000101?00000000100000{34}0100022{01}2101001
		Begonia_ankaranensis              00000000?0?0?1100??0??0000????????0?2?2???00?
		Begonia_angularis                 000000000000011010000001001003010102202101001
		Begonia_aequata                   000000000000012?0??0?00??????001010221200?000
	;
END;

BEGIN ASSUMPTIONS;
CHARSET morphology = 1 - 45 ; 
 CHARSET poor_morphology = 21 25 27 45 ; 
 CHARSET exclusion = 45 ; 
 OPTIONS DEFTYPE = unord PolyTcount = MINSTEPS ; 
 EXSET * excluded = 16 45 ; 
 END;