#NEXUS BEGIN TAXA; DIMENSIONS NTAX=61; TAXLABELS Dimorphodon_macronyx Campylognathoides_liasicus Rhamphorhynchus_muensteri Anurognathus_ammoni Gnathosaurus_macrurus Gnathosaurus_subulatus Plataleorhynchus_streptophorodon Huanhepterus_quingyangensis Elanodactylus_prolatus Kepodactylus_insperatus Mesadactylus_ornithosphysos Beipiaopterus_chenianus Boreopterus_cuiae Feilongus_youngi Eosipterus_yangi Pterodaustro_guinazui Ctenochasma_elegans Ctenochasma_porocristata Gegepterus_changi Cycnorhamphus_suevicus Gallodactylus_canjuersensis '"Pterodactylus" longicollum' Pterodactylus_kochi Pterodactylus_antiquus Germanodactylus_cristatus Germanodactylus_rhamphastinus Haopterus_gracilis Anhanguera_piscator Anhanguera_santanae Anhanguera_blittersdorffi Liaoningopterus_gui Tropeognathus_mesembrinus Lonchodectes_compressirostris Liaoxipterus_brachyognathus Nurhachius_ignaciobritoi Istiodactylus_latidens Istiodactylus_sinensis Pteranodon_longiceps Pteranodon_sternbergi Nyctosaurus_gracilis Nyctosaurus_bonneri Tupuxuara_longicristatus Tupuxuara_leonardii Thalassodromeus_sethi Tapejara_imperator Tapejara_navigans Tapejara_wellnhoferi Huaxiapterus_jii Sinopterus_dongi Sinopterus_gui Eopteranodon_lii Noripterus_complicidens '"Phobetor" parvus' Dsungaripterus_weii Eoazhdarcho_liaoxiensis Jidapterus_edentus Chaoyangopterus_zhangi Bakonydraco_galaczi Azhdarcho_lancicollis Zhejiangopterus_linhaiensis Quetzalcoatlus_sp.; END; BEGIN CHARACTERS; DIMENSIONS NCHAR=111; FORMAT SYMBOLS= " 0 1 2 3 4 5 6" MISSING=? GAP=- ; CHARSTATELABELS 1 'Skull length exclusive of cranial crests:' / 'between_three_and_five_times_the_height_(3.00_<_skull_length/height_<_5.00)' 'more_than_five_times_the_height_(skull_length/height_�_5.00)' 'less_than_three_times_the_height_(skull_length/height_�_3.00)', 2 'Dorsal margin of skull shape:' / nearly_straight convex concave, 3 'Rostrum shape:' / compressed_laterally 'compressed_antero-posteriorly' 'depressed_dorso-ventrally', 4 'Rostrum length anterior to external nares:' / less_than_50%_of_the_skull_length_exclusive_of_sagittal_crests extremely_reduced at_least_50%_of_the_skull_length_exclusive_of_sagittal_crests, 5 'Anterior end of premaxillae/maxillae shape:' / straight upturned downturned, 6 'Anterior tip of the rostrum with a dorsally reflected palatal surface:' / absent present, 7 'Nasoantorbital fenestra length:' / less_than_40%_of_the_skull_length at_least_40%_of_the_skull_length, 8 'Posterior margin of the nasoantorbital fenestra shape:' / nearly_a_straight_angle rounded_and_concave_outline, 9 'Orbit shape:' / subcircular small_and_circular piriform at_least_one_and_a_half_times_higher_than_wide, 10 'Orbit position:' / 'middle_of_the_skull_with_the_ventral_margin_of_the_orbit_below_the_middle_of_the_antorbital/nasoantorbital_fenestra_and_the_dorsal_margin_of_the_orbit_above_the_dorsal_margin_of_the_antorbital/nasoantorbital_fenestra' 'high_in_the_skull_with_the_ventral_margin_of_the_orbit_the_same_level_or_above_the_middle_of_the_antorbital/nasoantorbital_fenestra' 'low_in_the_skull_with_the_entire_orbit_lower_than_the_dorsal_margin_of_the_antorbital/nasoantorbital_fenestra', 11 'Angle between lacrimal and postorbital and lacrimal processes of the jugal:' / distinct_angle infilled, 12 'Posteriorly-directed process on the jugal process of the lacrimal:' / absent present, 13 'Anteriorly-directed process on the postorbital process of the jugal:' / absent present, 14 'Suborbital vacuity:' / absent present, 15 'Premaxillary sagittal crest:' / absent present, 16 'Premaxillary sagittal crest shape:' / striated_with_a_nearly_straight_dorsal_margin 'round_and_blade-shaped' 'tall_anteriorly,_decreasing_in_height_posteriorly' 'low_anteriorly,_increasing_in_height_posteriorly' 'low_with_a_humped_antero-dorsal_margin' low_ridge, 17 'Elongate dorsal spine extending above premaxillary crest:' / absent present, 18 'Premaxillary sagittal crest position:' / entire_crest_anterior_to_the_nasoantorbital_fenestra crest_starting_anterior_to_and_extending_posterior_to_the_anterior_margin_of_the_nasoantorbital_fenestra crest_starting_at_the_anterior_margin_of_the_nasoantorbital_fenestra_and_extending_backwards crest_starting_near_the_anterior_margin_of_the_skull_and_extending_over_the_occipital_region crest_starting_at_the_posterior_half_of_the_nasoantorbital_fenestra, 19 'Lateral expansion of the anterior tip of jaws:' / absent present, 20 'Posterior ventral expansion of the maxilla:' / absent present, 21 'Nasal process:' / present absent, 22 'Nasal process position:' / placed_laterally placed_medially, 23 'Free nasal process length:' / long short, 24 'Nasal process orientation:' / 'directed_antero-ventrally' directed_ventrally 'directed_postero-ventrally', 25 'Foramen on nasal process:' / absent present, 26 'Bony frontal crest:' / absent present, 27 'Bony frontal crest shape:' / low_and_blunt low_and_elongated high_and_expanded, 28 'Lacrimal process of jugal thickness:' / broad thin, 29 'Lacrimal process of the jugal inclination:' / inclined_anteriorly subvertical strongly_inclined_posteriorly, 30 'Bony parietal crest:' / absent present, 31 'Bony parietal crest shape:' / blunt 'laterally_compressed_and_posteriorly_expanded,_with_a_rounded_posterior_margin' constituting_the_base_of_the_posterior_portion_of_the_cranial_crest, 32 'Posterior region of skull rounded with squamosal displaced ventrally:' / absent present, 33 'Squamosal position:' / above_the_base_of_the_lacrimal_process_of_the_jugal below_or_level_with_the_base_of_the_lacrimal_process_of_the_jugal, 34 'Orientation of quadrate relative to the ventral margin of skull:' / subvertical inclined_about_120�_posteriorly inclined_about_150�_posteriorly, 35 'Position of articulation between skull and mandible:' / posterior_to_the_middle_of_the_orbit anterior_to_the_middle_of_the_orbit under_the_middle_of_the_orbit, 36 'Helical jaw joint:' / absent present, 37 'Supraoccipital shape:' / does_not_extend_backwards extends_backwards, 38 'Distal ends of paroccipital processes:' / not_expanded expanded, 39 'Palate with ridge and mandible with sulcus:' / absent present, 40 'Palatal ridge length:' / tapering_anteriorly confined_posteriorly, 41 'Palatal ridge shape:' / discrete strong, 42 'Interpterygoid opening size:' / larger_than_subtemporal_fenestra smaller_than_subtemporal_fenestra, 43 'Mandibular symphysis position:' / at_same_level_as_rami located_well_below_rami on_a_symphyseal_shelf_dorsal_to_ventral_margin, 44 'Bony mandibular symphysis length:' / less_than_30%_of_the_mandible_length at_least_30%_of_the_mandible_length, 45 'Anterior tip of dentary shape:' / straight projected_anteriorly downturned upturned, 46 'Dentary bony sagittal crest:' / absent present, 47 'Teeth:' / present absent, 48 'Tip of rostrum:' / teeth_present teeth_absent, 49 Tip_of_mandible / teeth_present teeth_absent, 50 'Posterior margin of the maxillary toothrow:' / 'extends_posterior_to_the_anterior_margin_of_naris/nasoantorbital_fenestra' 'at_or_anterior_to_the_anterior_margin_of_the_naris/nasoantorbital_fenestra', 51 'Toothrow length:' / more_than_or_equal_to_one_third_of_the_skull_length less_than_one_third_of_the_skull_length, 52 'All teeth set sub-horizontally in jaws:' / absent present, 53 'Distance between successive teeth:' / most_less_than_the_diameters_of_the_teeth most_more_than_the_diameters_of_the_teeth most_constant_and_subequal_to_the_diameters_of_the_teeth, 54 'Position of largest maxillary teeth:' / middle_of_maxilla located_mesially located_distally, 55 'Variation in the size of anterior teeth with 5th and 6th smaller than 4th and 7th:' / absent present, 56 'Tooth shape:' / 'recurved_and_labio-lingually_compressed' slender sharp_and_conical broad_with_oval_base 'lancet-shaped_and_labio-lingually_compressed' 'spike-shaped_with_wide,_sub-circular_base', 57 'Tooth height:' / elongate_teeth_absent elongate_teeth_present, 58 'Strongly recurved teeth present:' / absent present, 59 'Inclination of teeth:' / all_teeth_upright anterior_teeth_procumbent all_teeth_procumbent, 60 'Tooth number:' / less_than_100 at_least_100, 61 'Notarium:' / absent present, 62 'Atlas and axis:' / unfused fused, 63 'Postexapophyses on cervical vertebrae:' / absent present, 64 'Lateral pneumatic foramen on centrum of cervical vertebrae:' / absent present, 65 'Mid-cervical vertebrae length:' / 'short_(length/mid-width_�_2.5)' 'elongate_(2.5_<_length/mid-width_<_5)' 'extremely_elongated_(length/mid-width_�_5)', 66 'Neural spines of mid-cervical vertebrae shape:' / 'blade-shaped' 'spike-shaped' ridge, 67 'Neural spines of mid-cervical vertebrae height:' / tall low extremely_reduced_or_absent, 68 'Number of posterior-cervical vertebrae:' / one two, 69 'Neural spines of posterior-cervical vertebrae height:' / tall low, 70 'Length of the scapula relative to coracoid length:' / 'scapula_at_least_one_and_a_half_the_coracoid_length_(sca/cor_�_1.50)' 'scapula_between_one_and_one_and_a_half_times_the_coracoid_length_(1.00_�_sca/cor_<_1.50)' 'scapula_at_least_80%_but_less_than_the_coracoid_length_(0.8_<_sca/cor_<_1.00)' 'scapula_at_most_80%_of_the_coracoid_length_(sca/cor_�_0.8)', 71 'Proximal surface of scapula shape:' / elongated 'sub-oval', 72 'Scapula shape:' / elongated stout_with_constricted_shaft, 73 'Scapula rotated outwards:' / absent present, 74 'Scapula articulates with notarium:' / absent present, 75 'Sternum shape:' / 'semi-circular' rectangular triangular, 76 'Coracoid contact surface with sternum:' / 'articulation_surface_flattened,_lacking_posterior_expansion' 'articulation_surface_oval,_with_posterior_expansion', 77 'Coracoidal flange:' / absent present, 78 'Broad tubercle on ventroposterior margin of coracoid:' / absent present, 79 'Cristospine constricted behind coracoid facets:' / absent present, 80 'Cristopine shape:' / shallow_and_elongated deep_and_short, 81 'Length of humerus relative to metacarpal IV length:' / 'humerus_at_least_two_and_a_half_times_the_metacarpal_IV_length_(hu/McIV_�_2.50)' 'humerus_between_one_and_a_half_and_two_and_a_half_times_the_metacarpal_IV_length_(1.50_<_hu/McIV_<_2.50)' 'humerus_between_55%_and_up_to_one_and_a_half_times_the_metacarpal_IV_length_(0.55_<_hu/McIV_�_1.50)' 'humerus_at_most_55%_of_the_metacarpal_IV_length_(hu/McIV_�_0.55)', 82 'Length of humerus relative to femur length:' / 'humerus_at_most_82%_of_the_femur_length_(hu/fe_�_0.82)' 'humerus_between_82%__and_140%_of_the_femur_length_(1.40_>_hu/fe_>_0.82)' 'humerus_at_least_140%_of_the_femur_length_(hu/fe_�_1.40)', 83 'Length of the ulna relative to the femur length:' / 'ulna_at_most_160%_the_femur_length_(ul/fe_�_1.60)' 'ulna_more_160%_the_femur_length_(ul/fe_>_1.60)', 84 'Ventral surface of proximal end of humerus:' / pneumatic_foramen_absent pneumatic_foramen_present, 85 'Dorsal surface of proximal end of humerus:' / pneumatic_foramen_absent pneumatic_foramen_present, 86 'Position of deltopectoral crest on humerus:' / proximally_placed positioned_farther_down_on_humerus, 87 'Deltopectoral crest of humerus shape:' / broad_with_almost_straight_anterior_margin 'subrectangular,_extending_down_the_humerus_shaft_for_greater_than_or_equal_to_30%_of_the_humerus_length' distally_expanded_and_rounded warped elongate_and_rounded elongate_with_straight_proximal_and_distal_margins 'enlarged_and_hatchet-shaped', 88 'Medial (ulnar) crest of humerus:' / reduced distinct, 89 'Medial (ulnar) crest of humerus shape:' / directed_posteriorly massive_with_a_developed_proximal_ridge, 90 'Humerus shaft shape:' / straight bowed, 91 'Midshaft of humerus tapered:' / absent present, 92 'Distal end of humerus:' / 'oval_or_D-shaped' subtriangular, 93 'Distal end of humerus:' / pneumatic_foramen_absent pneumatic_foramen_present, 94 'Length of ulna relative to humerus length:' / 'ulna_less_than_one_and_a_half_times_humerus_length_(ul/hu_<_1.50)' 'ulna_at_least_one_and_a_half_times_humerus_length_(ul/hu_�_1.50)', 95 'Length of ulna relative to tibia length:' / 'ulna_at_most_the_tibia_length_(ul/ti_�_1.00)' 'ulna_longer_than_the_tibia_length_(ul/ti_>_1.00)', 96 'Diameter of the radius relative to the ulna diameter:' / radius_more_than_half_of_the_ulna_diameter radius_at_most_half_of_the_ulna_diameter, 97 'Distal syncarpal shape:' / fused_in_a_rectangular_unit fused_in_a_triangular_unit, 98 'Medial carpal shape:' / longer_than_wide wider_than_long, 99 'Pteroid length:' / shorter_than_half_the_length_of_the_ulna longer_than_half_the_length_of_the_ulna, 100 'Metacarpals I-III:' / articulate_with_carpus 'metacarpal_III_articulates_with_carpus,_metacarpals_I_and_II_reduced' do_not_articulate_with_carpus, 101 'Length of first wing phalanx relative to metacarpal IV length:' / 'first_phalanx_at_least_twice_the_metacarpal_IV_length_(f1dIV/McIV_�_2.00)' 'first_phalanx_up_to_twice_the_metacarpal_IV_length_(1.00_<_f1dIV/McIV_<_2.00)' 'first_phalanx_at_most_the_metacarpal_IV_length_(f1dIV/McIV_�_1.00)', 102 'Length of first wing phalanx relative to tibia length:' / 'first_phalanx_less_than_one_and_a_third_the_of_the_tibia_length_(f1dIV/ti_<_1.33)' 'first_phalanx_at_least_one_and_a_third_the_tibia_length_(f1dIV/ti_�_1.33)', 103 'Length of the second wing phalanx relative to the first phalanx length:' / 'second_phalanx_at_least_the_first_phalanx_length_(f2dIV/f1dIV_�_1.00)' 'second_phalanx_between_70%_and_the_first_phalanx_length_(_0.70_<_f2dIV/f1dIV_<1.00)' 'second_phalanx_at_most_70%_of_the_first_phalanx_length_(f2dIV/f1dIV_�_0.70)', 104 'Length of second wing phalanx relative to the ulna length:' / 'second_phalanx_less_than_the_ulna_length_(f2dIV/ul_<_1.00)' 'second_phalanx_at_least_the_ulna_length_(f2dIV/ul_�_1.00)', 105 'Length of the third wing phalanx relative to the first phalanx length:' / 'third_phalanx_at_least_the_first_phalanx_length_(f3dIV/f1dIV_�_1.00)' 'third_phalanx_less_than_the_first_phalanx_length_(f3dIV/f1dIV_<_1.00)', 106 'Cross-section of second and third wing phalanges:' / round_to_subtriangular 'inverted_T-shaped_cross_section', 107 'Ischiopubic plate:' / unexpanded present absent, 108 'Strongly bowed femur:' / absent present, 109 'Angle of femur caput to shaft:' / at_most_145� greater_than_145�, 110 'Length of the femur relative to metacarpal IV length:' / 'femur_at_least_twice_the_metacarpal_IV_length_(fe/mcIV_�_2.00)' 'femur_between_one_and_a_quarter_and_twice_the_metacarpal_IV_length_(1.25_<_fe/mcIV_<_2.00)' 'femur_between_one_half_and_up_to_one_and_a_quarter_the_metacarpal_IV_length_(0.50_<_fe/mcIV_�_1.25)' 'femur_at_most_one_half_the_metacarpal_IV_length_(fe/mcIV_�_0.50)', 111 'Length of metatarsal III relative to tibia length:' / 'metatarsal_III_between_one_third_and_one_half_of_the_tibia_length__(mtIII/ti_>_0.33)' 'metatarsal_III_at_most_one_third_the_tibia_length__(mtIII/ti_�_0.33)' ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110] [ . . . . . . . . . . . ] Dimorphodon_macronyx 010000-03000000---0000-000-000-0?00?00????0-00000000100000000?00000??1000-?00???11000000-1000000000000000?00010 Campylognathoides_liasicus 000000-00000000---0000-000-000-00000000--00-20001000100000000000000??100?-10000011100010-??00010?00001011?10010 Rhamphorhynchus_muensteri 000000-00000000---0000-000-000-00120000--00110011000110110200000000001000-10000011100020-1000110??0001111010010 Anurognathus_ammoni 011100-0??000?0---0000-100-01?????????????0-00000000110200000???0????1???-??????011??000-1000010??000100???0?00 Gnathosaurus_macrurus ??????????????????1???????????????????100?0?000?0??11?01??11??11222???????????????????????????????????????????? Gnathosaurus_subulatus 10220000000000100110000000-010-1122?0010000?0000?10111011121??????????????????????????????????????????????????? Plataleorhynchus_streptophorodon ?02?00????????????1???????????????????0--???0?000??10101??11??????????????????????????????????????????????????? Huanhepterus_quingyangensis ?02?00????????10011???????????????????????0?000001?0110111110???22201??0??0??????10??04??1?0?1?????0???????01?? Elanodactylus_prolatus ????????????????????????????????????????????????????????????0?1?122011000-?000??2???1040-100?0?0?0101?0100????? Kepodactylus_insperatus ??????????????????????????????????????????????????????????????01121??????????????1?01040-1000????????????0?0??? Mesadactylus_ornithosphysos ??????????????????????????????????????????????????????????????01121??????????????{12}?00040-10?????????1?0????012? Beipiaopterus_chenianus ????????????????????????????????????????????????????????????0???10101100?-?010??2210?04??100?000??0?01201??0?20 Boreopterus_cuiae 1002000?000??00---00?????0-??0-??22?0?????0100000000110110110?00100?????????????210???????0??010??0?11111??0?21 Feilongus_youngi 12020000?00000100000000000-01111?2?0000--?0100000110110111{01}0??????????????????????????????????????????????????? Eosipterus_yangi ??????????????????????????????????????????????????????????????????????0???????????1????????0??00???0101110?0?20 Pterodaustro_guinazui 120210000000000---00000200-000-1122?000--?013000010001011001??1110101?000?0000012110004???00?0100???101110?0?20 Ctenochasma_elegans 120200000000000---00{01}00200-010-1122?00????0100000101210110210?00101011000-0000??21000040-100?000??0?10111?10120 Ctenochasma_porocristata ?20?00?????00?100100??????????????????0--???0?00010121?11021??????????????????????????????????????????????????? Gegepterus_changi 120210000001001000?0000200-000-1?22?0?0--?0?300??10011?11021?011101???????0?10?????????????????????{01}1?1??0?0??? Cycnorhamphus_suevicus 120000000000000---00001?00-?1111122??00--?010000?110???111?0?0001010?1000-00000121000040-100?000???010111010121 Gallodactylus_canjuersensis 120000000000000---00001000-?111?1?2??0????0?00???????????????????????100???000???????????????????????0111?101?1 '"Pterodactylus" longicollum' 120000000000000---00000?00-010-1122?0?0--??1000001001102001000001010?1000-??00??20000040-000?000???010{12}11?10121 Pterodactylus_kochi 100000000000000---00000100-010-1122?000--?0100000100110200000000101011000-?000?021000040-10000{01}00?0010101010120 Pterodactylus_antiquus 100000000000000---00000100-010-1122?000--?0100000100110200000000101011000-00000021000040-100?000000010101010120 Germanodactylus_cristatus 00000000000000100200000100-010-1022?00????0100011000100200000?0?1??011000-?000??2100004??100?000????10111??0121 Germanodactylus_rhamphastinus 00000000000000100200000000-010-1022?0?????0?00000000100200000?0?101???000-?0????210??????100?000????10101?10121 Haopterus_gracilis 0000000?00?00?0---0001110??????0?1??0??????100000000110200000?0?0??0?200?-0000002???0040-000?010????1?1110????? Anhanguera_piscator 10000101201100110010010011001100011100???1010100000011150110011101010311?-01001121101031001111111??1??1??020121 Anhanguera_santanae 1000010120110011?0100100110011000111001001010?0000001115?1100111010103111-?100??211010310011???110?1111??02???1 Anhanguera_blittersdorffi 100001012011001100101----10011000111001001010100000011150110??????????????????????????????????????????????????? Liaoningopterus_gui 100?01?12{02}11001100?0???????01??00111??100?0?0?000?0011050110??110?????????????????????????????????????????????? Tropeognathus_mesembrinus 100001012011001100101----10011000111001011010100000011050110??????????????????????????????????????????????????? Lonchodectes_compressirostris ?00?0?????????0---0???????????????????100???000000?01105??{01}??????????????????????????????01???????????????????? Liaoxipterus_brachyognathus ??2?0?????????????0???????????????????????000?000?100?0400?0??????????????????????????????????????????????????? Nurhachius_ignaciobritoi 10000110?101?00---00???????11????11???????01000?0?10110400101?11010??2?1??0?0???2100?031001??111??011111???0?21 Istiodactylus_latidens 102001101101110---001----??120??0210??0--?0000000110010400001?1?0?01?31111?00011?110?031001111?11?1???11??201?? Istiodactylus_sinensis 102001101101110---001----0-120-002100?????000000001001040000?1???10??3?1????00??2???003??01??11??0?111111020?2? Pteranodon_longiceps 120210012010000---000111012011200111000--111001-------------111101010210111000103101003100110000101211111010131 Pteranodon_sternbergi 12021?012010000---00?????1201120011?0?????110?1-------------?????????2??????????310??????0???00?????111110?013? Nyctosaurus_gracilis 12020001?000000---001----12010-00121000--121001-------------11100001020010000000310101610000?1101?1211111010031 Nyctosaurus_bonneri 1?0?0001?0???00---00?????????0-0??2?00????21001-------------1?1?000??????0??????31???1610000?1?????2211?1?1??3? Tupuxuara_longicristatus ??0??0?0??0???130?0???????????????????111?????1-------------???????????????????????????????????????2??????????? Tupuxuara_leonardii 000000102200001303001----1111120011111111101001-------------1111000??100?1?001?1200100511??0?0000???1021?0?0021 Thalassodromeus_sethi 00000010220000130300011101111120011111111?01001-------------??????????????????????????????????????????????????? Tapejara_imperator 2100201?2200001211001----1111120011?1???????2?1-------------??????????????????????????????????????????????????? Tapejara_navigans 2100201022000012110001010??11??0011?????????2?1-------------??????????????????????????????????????????????????? Tapejara_wellnhoferi 210020102200001201001----11111200110110--101211-------------0?11000??100?-?001??200110511??0?0000?1?10???0?002? Huaxiapterus_jii 01002010??0???1101001----??11????2????0--?01211-------------0???0????????-1001??2001?0511000?000?111101110?0021 Sinopterus_dongi 01002010220???110100010101111120?21?1?????01211-------------??1?000??1?01???0?012101?0511000?1000102101110?0?21 Sinopterus_gui 01002010??0???110100???????11????21???????01211-------------??1?000??1?01??00?0??10??0511000?100??0??0????10??1 Eopteranodon_lii 01002010??0???110-00???????11??????1??0--?012?1-------------??1?100??{12}?0??0?1???210??051?000?100??1?1?11101002? Noripterus_complicidens ??0??0????????????????????????????????????0?000????01?030000??11000?????????0???3000105110000?000??0101??011021 '"Phobetor" parvus' 000000001100011001011----1101120011?11????010001100012030000??????????????????????????5?????????01????????????? Dsungaripterus_weii 000010001100011001011----11011200111110--101300110001203000011110001000011100?01300??05110?0?00001?0101110?1021 Eoazhdarcho_liaoxiensis ??0???????????????0???????????????????0--?11001-------------???0121??1?0????1???210??05??00??00???12101110?0?2? Jidapterus_edentus 020000????????0---00??????????????????0???11001-------------??10121??1?0??10???1210??????00??000??1110211??0?21 Chaoyangopterus_zhangi 020?00????????0---00??????????????????????11001-------------??10121??100?????001300?00???000?000??1{12}10211010021 Bakonydraco_galaczi ??0???????????????0????????????????1??0--?01011-------------??10221???????????????????????????????????????????? Azhdarcho_lancicollis ??0?0?????????????0????????????????1??0--?????1-------------11102221?????1?????????1015??0?0?????????????1????? Zhejiangopterus_linhaiensis 100000002210000---001----0-010-0111?1?????11001-------------11??222?0100??1?10?1300??15??000?{01}00??122020?110021 Quetzalcoatlus_sp. 100?00002210001404001----??01????111??0--?01001-------------1110222101001100100130?101511000100001?2202011?00?? ; END; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=unord PolyTcount=MAXSTEPS ; TYPESET * UNTITLED = unord: 1-64 66-111, ord: 65; END; BEGIN TREES; TRANSLATE 1 Dimorphodon_macronyx, 2 Campylognathoides_liasicus, 3 Rhamphorhynchus_muensteri, 4 Anurognathus_ammoni, 5 Gnathosaurus_macrurus, 6 Gnathosaurus_subulatus, 7 Plataleorhynchus_streptophorodon, 8 Huanhepterus_quingyangensis, 9 Elanodactylus_prolatus, 10 Kepodactylus_insperatus, 11 Mesadactylus_ornithosphysos, 12 Beipiaopterus_chenianus, 13 Boreopterus_cuiae, 14 Feilongus_youngi, 15 Eosipterus_yangi, 16 Pterodaustro_guinazui, 17 Ctenochasma_elegans, 18 Ctenochasma_porocristata, 19 Gegepterus_changi, 20 Cycnorhamphus_suevicus, 21 Gallodactylus_canjuersensis, 22 '"Pterodactylus" longicollum', 23 Pterodactylus_kochi, 24 Pterodactylus_antiquus, 25 Germanodactylus_cristatus, 26 Germanodactylus_rhamphastinus, 27 Haopterus_gracilis, 28 Anhanguera_piscator, 29 Anhanguera_santanae, 30 Anhanguera_blittersdorffi, 31 Liaoningopterus_gui, 32 Tropeognathus_mesembrinus, 33 Lonchodectes_compressirostris, 34 Liaoxipterus_brachyognathus, 35 Nurhachius_ignaciobritoi, 36 Istiodactylus_latidens, 37 Istiodactylus_sinensis, 38 Pteranodon_longiceps, 39 Pteranodon_sternbergi, 40 Nyctosaurus_gracilis, 41 Nyctosaurus_bonneri, 42 Tupuxuara_longicristatus, 43 Tupuxuara_leonardii, 44 Thalassodromeus_sethi, 45 Tapejara_imperator, 46 Tapejara_navigans, 47 Tapejara_wellnhoferi, 48 Huaxiapterus_jii, 49 Sinopterus_dongi, 50 Sinopterus_gui, 51 Eopteranodon_lii, 52 Noripterus_complicidens, 53 '"Phobetor" parvus', 54 Dsungaripterus_weii, 55 Eoazhdarcho_liaoxiensis, 56 Jidapterus_edentus, 57 Chaoyangopterus_zhangi, 58 Bakonydraco_galaczi, 59 Azhdarcho_lancicollis, 60 Zhejiangopterus_linhaiensis, 61 Quetzalcoatlus_sp. ; 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