NEXUS
[File created by TreeBASE:  7/26/07  21:07:04]
[Matrix accession#: M2112]

BEGIN TAXA;
	TITLE M2112;
	DIMENSIONS  NTAX=69;
	TAXLABELS
		Begonia_symsanguinea	Hillebrandia_sandwicensis	Datisca_glomerata	Datisca_cannabina	Begonia_masoniana	Begonia_malabarica	Begonia_sp._Pritzelia	Begonia_lobata	Begonia_angularis	Begonia_violifolia	Begonia_thomeana	Begonia_sutherlandii	Begonia_subscutata	Begonia_staudtii	Begonia_socotrana	Begonia_sericoneura	Begonia_scutifolia	Begonia_scapigera	Begonia_samhaensis	Begonia_salaziensis	Begonia_roxburghii	Begonia_quadrialata	Begonia_prismatocarpa	Begonia_potamophila	Begonia_poculifera	Begonia_palmata	Begonia_olbia	Begonia_obliqua	Begonia_nossibea	Begonia_molleri	Begonia_meyerijohannis	Begonia_mannii	Begonia_mananjebensis	Begonia_madecassa	Begonia_luxurians	Begonia_loranthoides	Begonia_longipetiolata	Begonia_letouzeyi	Begonia_kisuluana	Begonia_johnstonii	Begonia_iucunda	Begonia_integerrima	Begonia_incarnata	Begonia_horticola	Begonia_holtonis	Begonia_grandis	Begonia_goegoensis	Begonia_geranioides	Begonia_gabonensis	Begonia_francoisii	Begonia_floccifera	Begonia_engleri	Begonia_edmondoi	Begonia_duncanthomasii	Begonia_dregei_partita	Begonia_dregei_ssp_homonyma	Begonia_dregei	Begonia_dipetala	Begonia_dewildei	Begonia_crassirostris	Begonia_convolvulacea	Begonia_cinnabarina	Begonia_capillipes	Begonia_bogneri	Begonia_balansana	Begonia_aspleniifolia	Begonia_annobonensis	Begonia_ankaranensis	Begonia_aequata
	;
END;

BEGIN CHARACTERS;
	TITLE M2112;
	DIMENSIONS NCHAR=1541;
	FORMAT DATATYPE = DNA MISSING = ? GAP = -  ;
	MATRIX
		Begonia_symsanguinea              TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGCG-AACAAGTTTTT----CCAATG---TCCCGCTG----CGCGC----GGGAA-TGGCGCGGG-GCGCGTCGG-AGTTGCAGCT--CGC-GCTT---------GCTCGAGCCCA--GCC--TCGCA-CGCCTCGTGCCCCC----------------TAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TCTAGAA-CTT---AGAATGTTGA---TGCCCT-GCCCCAAATTTTGG-GACGT----AGGGG--TCT-TCTGATCTT-----GATCACA--TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCT-------GCACCCAAAAAAAAAA------------CCAAG-T--CAA---TCCCTC--------GATC--GAAMYMA--GACTC---GTGCT-GCTTC--GTTGTG------GTTTT----GTT-GGCCTTT----------GGGTGGTC----GGGGAGGCGACGTTGGCCTCCCGTGCGGGT----ARCY-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGTCAT--GCCCTCGACGAATTAC--TTTGCGCTTGCGAGACGCGTGCTTGG-GTA-CTCC-TCGTC-------GACCC--------TAAATGCGTCGACTC-GTT---------GGACR-CACC------AGCGAGAAGGCGTTAT-----CGAAGC--GACCCCAGGTCAGG--------------------GTCGGGTTGTTTGGGAATGCAGCCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-ACGGCGTGCTTCGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGC-------
		Hillebrandia_sandwicensis         GATCATTGTCGATTCCTGC--AAAA---GCAG-AAAAACTCGCG-AACTCGGTTA--AAAACAAC----AC---TGTGGC-CCGTCGGCAAAAAAGT-CGGGGCGCGTCCGTGC--CGAGCACTT------------------TGCTTGCCCGGCC------GCGGC-GCCCCTCGCAAC--------------TC----AACGAACCCC--GGCGCAAGTCGCGCCAAGGAAACTC-TTAA-CTTTTCTTAA---GGC----ACGTT-GCCCC-GTTTTCGG-GACGAAC----GCTG-GTCTTGCGAAAGTGCTAGTATTATCACGCTCA--ACGACTCTCGACAACGGATATCTTGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACACATCG-TTGCCCCCC--------CGAAAAAAGATTT----------------------AAGGA--AAAATAGTTCGAATCCCGA---CTCCGACCGGGT-CGGCGGGATGCG---------CGCTGTTG----------------------------------------GAGGTGCTGAAGTTGGCCTCCCGTGCGTCG------TT-CGAC--GGTTGGCCTAAATTAGAGATCTCGGCGT-CGCGCGGTGCGATCGACGGTGGTTGCGAT--GCCCTCGGCGAAAAACAAGTTGCACTCGCGCGTCGCATCTCCGA-GAAGCTCCC-CGAT-------GACCC--------TCT-TGTTGTGCCGCGTCGCTTCCGGCTCCG-------------GACGACGCGCGCCTGCACCATCGAAGCGCGACCCCAGGTCAGGCGGGATCACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Datisca_glomerata                 GATCATTGTCGATGCCTT--AAAA---------AACAACCCGTG-AACGTGTTTT------CAATC---TCGGGTGTGCG-CGGGCTC-GAAAGAGCCCTCGCGCTCCCGCGCGGGGAGGCGGGCGTGCTAGCACCTATCCCTTTACGGGCGCGGTCTGGCTCGTTG-CGTTCCTTCTTG--------------CAAACTAACGAACCTCC-GGCGCGGACTGCGCCAAGGAA--CAA-TAA-CA----ATAGCGCGGC----CCCTC-GCCCC-GATACCGG-TGCGC----GTGGGG-TTGACGTGACATG----TGATAT----TCATA--ACGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATTG-TTGCCCCC--------ACGAACACAACC--------------------------------ATCTCTTT--------GAGGTGACTG--------------TATC--GT-------------------------------------------------------GG-GAGCAGACATTGGTCTCCCGTGCGCAT----TGCC-GTGT--GGTTGGCCTAAAA-CACGACCTCGGCTT-CATTCGCCGTGACAAACGGTGGTTGTCAA-AGC-CTCGGCGTCCTGT------CGCGTGCGTT-TGATGAAACATCGAGG-TC--TAATA-------GACCC--------TTC-TGCGTCGACCCT------------------------------CGA-----CGCTAT-----CGACGC--GACCCCAGGTCAGGCGGGACTACCCG------------------------------SCCTAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-CGGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAAAAGAGAGTCAAAGAGTGCTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGTGTT-GAGCCGGTCCGCCGATCGACTCGGGACGTGGACCGACGCGGATCGCGATGGCGGCCCAAGCCCGGGCCTTTGTTACGCTCGTGGAGACGTCGTCGTCGCGATCGTGGCTTGCAGCGCGCGCCTCATGGCGTGCTTCGGCAACTGCGCGCTCCTGGCGTCGGCCTGTGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCGAGTAAACCCGTAAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GTGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGAGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCG------
		Datisca_cannabina                 GATCATTGTCGATGCCTT---AAA---------AACAACCCGTG-AACGTGTTTT------CAAAA---TCGAGGGGG-T-GGGTCTC-GAGAGGGCCCTCACGTTCTTGTGCAGTGAGGCGGGCGTGCTGGCACCTGTCCCTTTACGGGCGTGGTCTGGCTCGTTC-CGTTCCTTCCTG--------------CAAACTAACGAACCTCC-GGCGCGGACTGCGCCAAGGAA--CAA-TAA-CA----ATAGCGCGGC----CCCTC-GCCCC-GATAACGG-TGTGC----GGGGG--CTGACGTGACCTG----TGATAT----TCATA--ACGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGAGGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTYTTTGAACGCAAGTTGCGCCCGAAGCCTTTTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATTG-TTGCCCCC--------ACGAACACAACC--------------------------------ATCTCTTT--------GAGGTGACTG--------------CATC--GT-------------------------------------------------------GG-GGGCAGATATTGGTCTCCCGTGCGCAT----TGTT-GTGT--GGTTGGCCTAAAA-CAAGACCTCGGCTT-CATTCGCCGCGACAAACGGTGGTTGTCAA-AGC-CTCAGCGTCCTGT------CACGTGTGTC-TGATGAAAAACCGAGG-TC--GAATA-------GACCC--------TTC-TGTGTCGACCTT------------------------------CGA-----CACTAT-----CGACGT--GACCCCAGGTCAGGCGGGACTACCCG-----TAGATCGGGTTGTTTGGGAATGCAGCCCCAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-CAGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAAAAGAGAGTCAAAGAGTGCTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGTGTT-GAGCCGGTCCGCCGATCGACTCGGGACGTGGACCGACGCGGATCGTGATGGCGGCCCAAGCCCGAGCCTTTGTTACGCTCGTGGAGACGTCGTCGTCGCGATCGTGGCTTGCAGCGCGCGCCTCATGGCGTGCTTCGGCATCTGCGCGCTCCTGGCGTCGGCCTGTGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCGAGTAAACCCGTAAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GTGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGAGAACATGCCTGTCCGGACC-GAAAGATGGTGAACTATGCCTGAGCG------
		Begonia_masoniana                 --------------------AGAGA---TC-G-AAC-----GCG-AAC-TAGTTT-GCCTGCCTTCG--TCGCGCCG----CGCGM----GGGAA-CGGTGCGGG-GCGCGTCWG-AGCTGCCAAC--CGC-GCTT---------GCGCGTTTTGAT-GCC--TCGCK-TGCCTCGCGCCCCC----------------TAACRAACCCC--GACGCGAGTTGCGTCAAGGAA--TCTTGAA-CTT---AGAACGCGGC---TCCCCT-GCCCC-GATTTCGG-GACCC----AGGGG--TTCCTGCGAACT------GAACA----CATCAA-DCGACTCTCGACTACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCWCGTCTGCCTGGGCGTCWCGCATCG-TWGCCCCCCG-------CACCACTTAWWAGAAWAW----------GAA-AC--CTA---TGCCTCGC-----TGCTC--GAARTG---GACAC---TGGCTCGCGCCTCCTACTTC-AAATCAGGGTTTTGTT--GGCCTWTT-----GGGTGGTGGTC----GGGGAGGCCACTTTGGCCTCCCGTGCGGGT----AACC-TCGC--GGTTGGTCTAAATTCGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCAGCATGCCCTYGACGAATTAC--GTTGCGCCCGCGAGAAGCGCGATCGG-GAA-CTCC-TGGTY-------GACCC--------TAA-TGCGCCGACTY-GTT---------GGACARCACC------AGCGAGAGGACGTTAT-----CGAAGY--GACCCCAG-TCWGGTKYATGTBCACT-----------------------------------------------ATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCTCGGTCGGATGTGTAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGAACC-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_malabarica                --ATTGT-CGATTCCTG-C-ARAAT---GCAG-AACAACCCGCG-AAC-TAGTTTTG----CCAACG--TCCCGCCG----CGAGC----GGGAA-CGGCGCGGG-GCGGGTCGG-WACTAAGGGC--CGC-GCTC---------GCGCGCGCCCTGTGCCC-CGGCG-AGCCTCGCGCCCCCC---------------TAACGAACCCC--GGCGCAAGTCGCGCCAAGGAA--TCC-GAA-CTC---AGAACGCGGC---TCCCCT-GCCCC-GGCTTCGG-GCTGC----GGGGG--TCC-TGCGAACTC-----GATCATT--ATACTAAACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGCCCCTCC------CCACCCACCCACCCACCCA---------CAAA------------------------------------------------------------GGGTGTT------------------------------------GGTCGGTC----GGGGAGGCGACGTTGGCCTCCCGTGCGGGC----AACC-GCGC--GGTTGGCCTAAATTTGAGTCCCCGGCGT-CTTGCGGCGCGATCGACGGTGGTAGCCATATGCCCTCGACGACTTAC--GTTGCGCCTGCTCGACGCGCGCTCGG-GAG-CTCC-TCGAC-------GACCCTC------TAA-TGCGTCGACTC-GCT---------GGGCAATGCC------AGCGAGCGGACGTCAT-----CGAAGC--GACCCCAGGTCAGGAGGGACTACCCG---------------------------------AAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCCCGGTCGGATGTGGAACGGCGCT-CAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTCGTTGTGCTCGCGGAGACGTCGTCGCCCCGATCGTGGCTGGCAGCGCGCGCCTCAGGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTS-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGGA----
		Begonia_sp._Pritzelia             -----GT-CGATTCCTG-C-AGAAT---GCAR-AACAACCCGCG-AAC-TAGTTTGG----CCAGCK--TCTCGTTT----CGCAC----GGGA--CGGCGCGGG-GCGTGCTGG-TGCTGTGGCC---GC-CTTT---------GTGCGCGCCCT--GCC--TGGT-ATGCCTTGCGCTCCCC--------------TTAACAAACCCC--GGCGCGAGTAGCGTCAAGGAA--TCTCGAA-CTC---AGAACGCACC---TCCCTC-GCCCC-GATTTTGG-GATGC----AGGAG--TGT-CGCGAACTT-----GATCATA--CATGAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------ACACCAAAAATAA----------------GAGGAGGTCAA---TGTCTC--------GATC--GAAGCA---GACTC---GCGCT-GCATC--GTCGCGG-----CTT-----GGCTC-GCCCTT----------GGGCGGTG----GTGGTGGCGACATTGGCTTCCCGTGGGGGC----GACC-TCGC--GGTTGGCCTAAATTTGAGTCCTTGGCGTTCTTGCGGCGCGATCGACGGTGGTTTCCAT--GCCCTCGACGAATCAC--ATTGCGCTTGCGAGATGC-TGCTGTT-GAA-CTCT-TGGAC-------GACCCTC---------------------------------------------------------AAATGTTAT-----CGAAGC--GACCCCAG-TCAGGCGGGATTACCCG-----------------------------------ATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGAG-TAGCCGGTCTGCCGATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCA-TTGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGACCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCATAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTC-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGC-------
		Begonia_lobata                    -----GT-CGATTCCGG-C-CGAAT---GCAA-AACAACCCGCG-AAC-TAGTTTGG----CCAATG--TCCCGTTT----CGCAC----GGGA--CGGCGCGGG-GCGTGTTGG-TGCTGTGGCC---GC-CTCT---------GTGCGCGCCCT--GCC--TGGC-AGGCCTCGCGCTCCCC--------------TTAACAAACCCC--GGCGCGAGTAGCGTCAAGGAA--TCTCGAA-CTT---AGAACGCACC---TCCCTC-GCCCC-AATTTTGG-GACGC----AGGAG--TGT-AGCGAACTT-----GATCATA--CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------ACACCAAAAATAA----------------GAGGAGGTCAA---TGTCTC--------GATC--GAAGCA---GACTC---GCGCT-GCATC--GTTGCGA-----CTT-----GGCTC-GCCCTT----------TGGCGGTG----GTGGCGGCGACATTGGCTTCCCGTGGGGGC----GACC-TCGC--GGTTGGCCTAAATTTGAGTCCTTGGCGTTATTGCGGCGCGATCGACGGTGGTTTCCAT--GCCCTCGACGAATCAC--ATTGCGCTTGCGAGATGCGTGCTATT-GAA-CTCT-TGGAC-------GACCCTCT------AAT----GTTCC------------------------------------TCAAATGTTAT-----CGAAGC--GACCCCAGGTCAGGYGGGAT-ACCC----------------------------------AAATCGGGC--TAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGAG-TAGCCGGTCTGCCGATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-TTGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCATAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGC-------
		Begonia_angularis                 TCATTGT-CGATTCCTG-C-AGAAT---GCAG-AACAACCCGCG-AAC-TAGTTTGG----CCAATG--TCCCGTTT----CGCAC----GGGA--CGGCGCGAG-GCGTGTTGG-TGCTGTGGCC---GC-CTCT---------GTGCGCGCCCT--GCC--TGGC-AGGCCTCGCGCTCCCC--------------CTAACAAACCCC--GGCGCGAGTAGCGTCAAGGAA--TCTCGAA-CTT---AGAACGCGCC---TCTCTC-GCCCC-GATTTTGG-GACGC----AGGAG--TGT-AGCGAACTT-----GATCATA--CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------ACACCAAAAATAA----------------GAGG------------------------------------------------------------------------------------------------------------TG----GTGGCGGCGACATTGGCTTCCCGTGGGGGC----GACC-TCGC--GGTTGGCCTAAATTTGAGTCCTTGGCGTTATTGCGGCGCGATCGACGGTGGTTTCCAT--GCCCTCGACGAATCAC--ATTGCGCTTGCGAGATGCGTGCTATT-GAA-CTCT-TGGAC-------GACCCTCA------AAT----GTTCC------------------------------------TCAAATGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG--------------------------------CAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGAG-TAGCCGGTCTGCCGATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-ATGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_violifolia                --------CTTTCTATG-CGGAGAT---GCAG-A-C--CCCGCG-AATTAGTTTG-----CCAACG---TCCCGCTT----CGCGC----GGGAA-CGGCGCGTT-GCGCGTCGG-GGCAGCGGCT--CGC-CTTT---------GCGCGCGCCCT--GTC--TCGC-GCGCCTCGCGCCCCC----------------TAACGAACCCC--GGCGCAAGTCGCGTCAAGGAA--TCTCAAAACTC---ATAACGCGGC---TCCCTT-GCCCC-GTTTTCGG-GATGC----AGGGG--TCC-AGCGAAATTT----GATCA----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCTGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGTCCCCC-------GCACCAAAAAAAGAAA--------------------CCAA----CCCTC--------GGTC--GAAGTA---GACTC---GTGCT-GCTTC--GTCAGGG-----CTT-----GGGT-GGCCTTC----------GTTTGGTC----GGGGTGGCGACATTGGCCTCCCGTGGGGGC----AACC-TCGC--GGTTGGCCTAAATTCGAGTCCTCGGCGC-CTTGCTGCGCGATCGACGGTGGTTTCTAT--GCCCTCGACGAATTAC--GTTGCGCTTGCGAGGCGCGCGCTCGG-GAA-CTCC-TCGAC-------GACCC--------TAA-CGCGTCGGCTG-GTTGG---------GCAACACC------GGCGAGCAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGTGGGACTAC----------GATCGGGTTGTTTGGGAATGCAGCCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCCCGATCGTGGCTGGCAGCGCGCGCCA-ACGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTTAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTC-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCG------
		Begonia_thomeana                  TC-TTGT-CGATTCCTG-M--GAGA---GCAG-AACAACCCGCG-AACTCGT-CTG----CCATCT---TCCC---------GTGC----GGGAA-CGGCGGG---GCGCACCGG-AGCCGCGGCC--CGGGAGC-----------------------------------------------------CTCGCGCACC-TAACGAACCC---GGCGCGAGTCGCGCCAAGGAA--TGTTTAA-CTG---AGARCGCGGT---TCCTCT-GCCCC-GGCCTCGG-GAAGC----AGGGG--CGC-TGCGAACTT-----GCTCG----TCACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TGGCCCCCC-------GCACCGAAAAGAMC----------------GAGT---CCGA--TCTCCCCGCGACGCAAGCAGGCCCGTTCTGCC-A-C-TC--------GGGGGGGGGTTCY----------GGT--GGCCTTT----------GGCCGGAC----GGGGCGGTAACGCTGGCCTCCCGYGCGAGC----GATC-GCGC--GGTTGGCCCAAACTCGAGCGTTCGGTGT-CGCGCGGTGCGATCGGCGGTGGTTGCCGT--ACCCTCGGCGAATTAC--GTCGCACTCGCGGG-CGCG--ATCCC-GAGAATCC-YTCAC-------GACC---------TCC-TGCGCTGC-TC-GTCGGCT----CMTTCG-GAAC------GACGGG-AGGCGAYGT-----CGAAGC--GACCCCAG-TCAGGTGG-AT-ACCC----------------------------------AAATCGGGCGGTAAATTCYGTCCAAGGCTAAATA-CTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGGCCCGGTCGGATGTGGAACGGCGG--TAGCCGGTCTGCCAATCGACTCGGGGCGCGGACCGACGCGGATTGCGGCGGCGGCCCAAGCCCGAGCCTTCGTCGTGCTCGTGGAGACGTCGTCGCCGCGATCGTGGCTGGCAGCGCGCGCCC-TCGGCGTGCTTCGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGCGTGCGAGTCAACGGGCCAGCAAACCCGTGAGGCGCAAGGAAGCTGASTGGCGGGATCCCCTC-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGG-TTCGAGTGCGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGGA----
		Begonia_sutherlandii              TCATTGT-CGATTCTTG-C-AAAAT---GCAG-AACAACCCGCG-AACTAGTTTG-CCCA-CAACGATGTCCTGCCA----CGCGC-----AGAA--GGCGTTG--GGGCGTCTA-AATTGTGGCC--CGT-GCTT-------GCGCGTGCGCCAT--GCCC-TCGA-ATGCCCCGCGCCCTCT--------------ATAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TATCGAACTTGTAAAGAACGCGGY---TACCCT-GCCCC-AGTTTTGG-GACGT----AGGGTG-TCC-TGCGAACTT-----GATCG----CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCMTCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCGTTYTGGC-------------GCCTGGGCGTYACRCATCG-TARCCCCCC-------GCACCCCAAAAAAAA----------------------------TCCCTC----GATCGATC--GAAGCTA--GACTC---GTGCT-GCTTC--GTCGTAGGG-----------GGCT-GGTT-------------GGCTGGTC----GGGGCGGAGATGTTGGCCTCCCGTGTGGGC----AACC-TCGC--GGTTGGCCTAAAATT-AGTCCTCGGCGT-CTTGYGGCGCGATCGACGGTGGTTGTCAT--GGCCTHGACGAATTAC--ATTGCGCACGCGAGACTTGCTGTCTG-GAA-CTC--TAGATGACGAATGACCC--------TMA-YGCGTCGGCTT-GTCGG---------ACAACACTC-GTMAGGCGAGCGGACGTTAT-----CGAAGC--GACCCCAG-TCA---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_subscutata                ------TGTCGATTCCTGC-CMWWW---GCAG-AACAACCCGCG-AACCCGTCGTG-----CAACGA--ACCC--------CGCAA---GGGGAG-CGGCGAGGA--GTGTTGGGG------------------------------------------------CGC-ACGCCCCGCATTC-------CGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAA-CTG---AGAACGCGGC---TCCTCT-GCCCC-GCCTGCGG-GATGCATGCAGGGG--CGC-TGCGAGCTT-----GAGAATC--AAACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCGCCAAAAAA-------TGC----------------TCGA--TCTC-TTC----------------------------------------TT-GGGGTCG------------GGCT-GGACTTT----------GGTCGGAC----GAGGGGGCGATGTTGGCCTCCCGTGCGAGC----GATC-GCGC--GGTTGGCCTAAAGTCGAGTCCTCGGCGT-CGCGCAGTGCGACCG-CGGTGGTTTTTAT-AGCCCTCGGCGAAATAC--GTTGCACGTGCGAGACGCGGTATCGG-GAA-CTCC-CACGC-------GACCC--------TGC-TGCGCTGCCCT-GCC-GTT-----CTCC--GAAC------GACGGGGAGGCGCTAT-----CGAAGC--GACCCCAGGTCAGG-GGGATTACCC--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_staudtii                  GATCATTGTCGATTCCT-----------GCAG-AACAACCCGCG-AATT-GGTAAA-GAACAATCTAC-TTCCCCAGTCG-TAACT---AGGGAA-GGGCGGGGC-GTGCTGGTG-C--CGCGGTG--CGC-GCTGC--------GCGCTYGGCCG--GCT--CGGC-ACTCCTCGCTG----------------ACCGTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--CATCTAA-CCG---AGAGCGCGGA---TCCTCT-GCCCC-GGACCCCGGGAAGC----AGGGT--TCT-CGCGCGCTT-----GCAAA----CGACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGTCCCCCCC-----GTACCGAAGAAGCCAA---------------------GCTA--GGGCTCCCGAAGC---AGAGC---------------------------TT-GCG-TCGGTA---------GCTC-GGATTTGCACCTC----GTTTGGA-----GGGGCGTCAA-GTTGGCCTCCCGCGCGACC----AGCT-GCGC--GGTTGGTCTAAGTTAGAGTGCTCGGCATTCGCGTGGTGCGATCGACG-TGGT-TCCTA--GCCCTCGGCGA---------------------------------------------------------------------------------------------------TTTGCCC-------GCGAGAAGGCGCTTT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_socotrana                 TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGCG-AACCGGTT-A-AGCTTCGAGCG--TCCTGCCT----CGTGC----GGGAA-CTGTGCGGG-GCGCGTCGG-ATCCGCGGAC--AGC-GCCC--------TGCGCGC-------GCC----GC-GCGGCCCGCGCCCCC----------------TAACGAACCCC--GGCGCGATCAGCGTCAAGGAA--CACGTAA-CTG---AGAACGCGGCT--TCCCCT-GCCCC-GATTTCGG-GACGC----GGGGGG-GCA-TGCGAAGTT-----GATCACG--TATCGAAACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCGCATGCCACCCC------GCACCCCGAAA------------CCAACAAGGACCGCTGGC--TCCCCC--------GGAC--GAAGCA---GGCTC---GTGCC-GCTTC--GCAGGGGCGGG-CTG-----CGCC-GGCCTTGGCGGG-----AGGGGGTC----GGGGAGGCGAAGTTGGCCTCCCGCGCGGGC----AGCC-TCGC--GGTTGGCCTAAATTTGAGTCCTCGGCGTCTTCGCAGCGCGATCGACGGTGGTAGCCAT--GCCCTCGACGAATCGC--GTCGCGCATGCGGGACGCGCTCGCGG-GAA-CTCC-TCCAG-------GACCC--------TAA-TGCGCCTGCTC-GCCGG---------ACGAGACC------GGCGAGGAGACGTCGTAT---CGAAGC--GACCCCAGGTCAGGCGGGACTACCCG-----------------------------GCCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGGC-GAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTCGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-ACGGCGTGCTTCGGCATCGGCGCGCTCCTGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGCGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTYTGAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCG------
		Begonia_sericoneura               ----TGT-CGAT-CCTG-A-GCRRW---GCAM-AACAACCCGCG-AACTAGTTTT-----CCAACG---TCCCGCTT----CGTGC----GGTAA-TGGC--AGG-GCGCGTCGG-AGCTGCGGCC--TGC-TTTT---------GTGCTGGCACT--CCC--TCGC-ACGCCTCTCGCCCCC----------------TAACGAACCCC--GACGCAAGTCGCGTCAAGGAA--TCTCAAA-CTC---ATAACGCGGC---TCCCTT-GCCCC-GATTTTGG-GACGC----AAGGG--TCC-TGCGAACTT-----GATCAAA--TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTTTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCCAAAAAAAAAA--------------------CCAA----CTCTC--------GGTC--GAAGTA---GACTC---GTGTT-GCTTT--GTCTGGG-----CTC-----AGTT-GGCGTTT----------AGGTGGTT----GGGGTGGCAACATTGGCCTCCCGTGTGGGC----AACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCAGCGT-CTTGCGGCGCGATCGACGGTGGTTGTCAT--GCCTTC-ACGAATTAC--GTTGCGCTTGCGAGGCGCACGTT-GG-GAA-CTCC-TCGAC-------GACCC--------TAA-CGCGTCAGCTA-ATCGG---------ACAAGACC------GGCGAGCAGACGTTAT-----CGAAGC--GACCCCAGGT-----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_scutifolia                ----------------------------------------CGCGTAA-TGGT----------ATCTGC-TTTCCCAGCCA-GCCCCT--GGGGAA-AGGCGGGGC-GTGCTGGTG-A--CGGGTTC--CGC-GCCCC--------GCGCGCGGCCG--GCA--TAGC-GCGCCTCGCAG----------------ACCGTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--GATGTAA-CCG---AGAGTGCGGA---TCTCCT-GCCCC-GGCCCTGG-GAAGC----AGGGGG-TCT-AGCGCGCTT-----GCTCA----CGAAGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCAAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCACCA-TAGGCCCCC-------GCACCGAAAAGGCCAA---------------------GTTC--GAGCTCCCCAAGCGG-CGGGCAG------TTAG---TCTGC-------TT-GCG-TCGGTG---------GTTT-GGATTTGCATCTC----GTTCGGA-----GGGGCGGCAAGGTTGGCCTCCCGCGCGGGC----AATG-GCGT--GGTTGGTCTAAAGTAGAGTGCTCGGCAT-CGCGCGGTGCGATCGACGGTGGTTTCCTT--ACCCTCGGCGAATTAC--GTCGCACGCGCGGGATGCG-GCGCGGAGAA-CTCC-ATTCG-------GACCC--------TTCTTTTATGA-TGCCTTTCT-----CGCGG-TCCCCC-------GCG---AGGCGCTGC-----CGAAGC--GACCC----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_scapigera                 ------TGTCGATTCCTM-----------CAG-AACGACCCGCG-AACC-GGTTAT-CAACCACACGA-TTCCCCAGCCA-GCGCT---GGGGAA-AGGCGGGGC-ACGCTGGTG-C--CGCGTTG--CGC-GCTCC--------GCGCGCGGCCG--TCC--CCGC-GCGCCTCGCCG----------------ACCGCAACGAACCCC--GACGCGAGTCGCGTCAAGGAA--CATGAAA-CTG---AGAGCGCGGA---TCCCAT-GCCCC-GGCTTCGG-GAAGC----AGGGA--TCT-CGCGCGCTT-----GCTCA----CGACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGTCCCCC-------TCACCGAAGAGGCCAA---------------------ATTC--GAGCTCCTGAAGC---AGTGCAG------TC-----TCTGCGC-----TT-GCG-TCGGGC---------GCTT-GGATTTGCAGCTC----GTTTGGA-----GGGGCGTCAAGGTTGGCCTCCCGCG-AAGC----AACC-TCGC--GGTTGGTCTAAAGTAGAGTGGTCGGCAT-CGCGCGGTGCGATCGACGGTGGTTTTGTA--GCCCTCGGCGAATTAC--GTCGCATGCGCGGGATGCG-GTGCGGAGAA-CTCC-ATACG-------GACCC--------TGC--TGATGATTGCC-GCCT-----CGCGGGTTCCCT-------GCGAGAAGGCGCTG------CGAAGC--GACCCCAGGTCAGG-GGGATTACCCG---------------------------------AAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCCCGGTCGGATGTGGAACGGCGC--GAGCCGGTCTGCCGATCGACTCGGGGCGTGGACCGACGCGGATCGTGGCGGCGGCCCAAGCCCGAGCCTTCGTTGAGCTCGCGGAGACGTCGTCGCGGCGATCGTGGAAGGCAGCGCGCGCCT-TCGGCGTGCTTAGGCATCAGCRCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGCGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGG-TTCGAGTGCGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGC-------
		Begonia_samhaensis                ---TTGT-CGATTCCTG-M-RRAAT---GCAG-AACAACCCGCR-AACTGGTTTATAAATTCTASCG--TCCTGCCT----CGTGC----GGGAA-CTGTGCGTG-GTGCGTCAG-ATCCCCGGAC--AGC-GCCC--------TGCGGGCGATGCGCGGY--TCGC-SCGGCCCACGCCCCC----------------TAACGAACCCC--GACGCGATCAGCGTCAAGGAA--CAATTAA-CTC---AGAACATGGCT--GCCCAT-GCCCC-GATTTCGG-GACAC----AAGGG--TCC-TGTGAAYTT-----GGTCATT--TATCGAAACGACTCTCGACAATGGATATCTTGGCTCTCGCATCGACGAAGAACGAAGTGAAATGCGATACTTGGTGTGAATTGCATAATCCCGTGAACCATTGAGTTTTTGAACGCAAGTTGCGTCCGAAGCCTTCTGGCTGAGGGCACGTCTGCCTGGGCGTCACGCACCGTTTGCCCCCCTTCCCSTGCACCCTAAAA------------CCAACAAGGATCAGTCGG--TCCCTC--------GAAC--CAAGCA---GACTC---GCGCT-ACTTT--GTAGGGGTGGG-CTG-----TGTC-GGCCTTGGCGGGCCGGCAGGCGGTCAGGCTGGGGGGCGAAGTTGGCCTCCCATGCGGGC----AGCC-TCCC--GGTTGGCCTAAATTTGAGTTCTCGGYATCCTTGCAGCGCGATCAACGGTGGTAGCCAT--GCCCTCGACTGATCAC--GTTGCGCATGCGGGACGCGC--GCGG-GAA-CTCC-TCCAG-------GACCC--------TAA-TGCGCCTACTC-GCCGG---------ACGAGACC------GGTGAGGAGACGTCATAT---CGAAGC--GACCCCGGGTCGGGCGGGACCACCCA--------------------------------------------------------------------------------------------------------GAAA-ATGAAAAGGACTTTGAGAARAGAGTCAARRAGTACTTGAAATTGTCGGGAGGGAAGCGSATGGGRGCAGACKKKKCGTC-CKGTC-SATSTGTAWYM-CRYY-GATC-WGTCTGCCWRTSSACTC-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_salaziensis               ----------------------------GCAG-AACAACCCGTG-AACTATTTAA-----CCATCG---TCCC--------CGCGC----GGGAA-TTGCGARGC--ATGTTGG--AGCTACGGCACAA---GCTT---------GCGCGTGCCCT--GCC--TTGC-ATGCCTCGCGA----------------ACCATAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TCTTTAA-CTG---AGAACGCTGC---TCTCCTAGCCC--GACTTTGG-GATGT----AGGGG--TGC-TGCGAACTT-----GTTTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGC-------------CACCCACACCAAAGCCTTGCACGCTCAATATCTCGGTCGA--AATAGACTTGTGCTATTTCTCTC----------------------------GGGGTTG------------GGCT-TGCATT----------GGGTTCGAT----GAGGTGGTAATGTTGGCCTCCCGCGCAAGC----AAGT-GCAC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-TGCGCAGTGCGATCGGCGGTGGTTGCCAT--GCCCTCGGCGAATTAC--ATTGCACTTGCGGGATGCATGTTCGG-GAA-CTCC-TTTGC-------GACCC--------TAT-TGCGCTGCCTC-GCTTGTT---CCYTCCGAGCGC------VATGAGAAGACGWWAT-----CGAAGC--GACCCCAG-TCAGGYGGGAYCACCC-------------------------------CCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGCGTC-TAGCCGGTCTGCCAATCGACTCAGGGCGTGGACCGACGCGGATTGTAGTGGTGGCCCAAGCCTGAGCCTTTGTTGTGCTTGTGGAGACGTCATCGCCGCGATCGTGGCTGGCAGCGCGCGCCT-CTGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GCGGGT-TGCACCGCCGACCGACCTTGATCTTTTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_roxburghii                --TCATTGTCGAGTCCTGC-ARAAT---GCAC-AACAACCCGCG-AACTAGTTT-G----CCAATG---TCCCGCCT----TGTGC----GGGAG-CAGTGCGGA-GCACGTTGG-AGCTGTGGAC--AGC-GCTT---------TTGCGCCCCTT--GCC--CTGCG-TGCCCCGCGCCCCC----------------TAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TCTAAAA-CTT---AGAATGCGGT---TCCCTT-GCCCC-GATTTCGG-GATGC----AGGGG--TCC-TGCAAACTT-----GATCG----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGTCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCCAAAAATAAAAAAA---------CCAACAA--CAA---TCCATC--------GATC--GAAGCA---GACCG---GCGCT-GGTTC--GTTGGGA-----GTTTTGGTTGTT-GGCCTTTT---------GGGCGATC----GGGGAGGCGACGTTGGCCTCCCGTACGGGC----AACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGCAGCGCGATCGACGGTGGTTGCCAT--GCCTTCGACGTGTTAC--ATTGTGCATGCAAGGCGCGCCGTGGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGACTG-GTC---------GGACAACAAC------GGCGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGYGGGATTACCCG------GAGTCGGGTTGTTTGGGAATGCAGCCCAAATCGGGCGGTAAATTTCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATTGTGGCTGGCAGCGCGCGCCT-ACGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTTAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTG-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGC-------
		Begonia_quadrialata               -----------------------------------------------------------------TGC-TTCCCCARCCA-GC--AT--GGGGAA-AGGCGGGGC-GTGCTGGTG-A--CGGTTAC--CGC-GCCCC--------GCGCGCGGCCG--GCC--CAGC-GCACCTCGCGG----------------ACCGTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--GATGTAA-CCG---AGAGTGCGGA---TCCCCT-GCCCC-GGCCCCGG-GAAGC----AGGGGG-TCT-AGCGCGCTT-----GCTCA----CGACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCACCG-TAGGCCCCC-------GGACCGAAGAGGCCAA---------------------GTTC--GAGCTCCCAAAGCGG-CGGGCAG------TTTC---TCAGC-------TT-GCG-TCGGCTGCG------GTTG-GGATTTGCATCTC----GTTCGGA-----GGGGCGGCAAGGTTGGCCTCCCGCGCGGGC----AATG-GCGC--GGTTGGTCTAAAGTAGAGTGCTCGGCAT-CGCGCGGTGCGATCGACG-TGGTTTCCTT--ACCGTCGGCGAATTAC--GTCGCACGCGCGGGATGCG-GCGCGGAGAA-CTCC-ATTCG-------GACCC--------TTCTTTTACGA-TGCCTTTCT-----CGCGGGTTCCCC-------GCGTG-AGGCGCTGC-----CGAAAC--GACCCCAGGTC----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_prismatocarpa             ---TCTTGTCGATGACTGVSMAAAK---ACAG-AACGACCCGCG-AACTGGTTAAA--AACCATCTGC-TTCCCCAGCCA-GAAAAT--GGGGAA-AGGTGGGGC-GTGCTGGTG-A--CGGGTTA--CGC-GCCCC--------GCGCGCGGCCG--GTC--TATC-ATGCCTCGCGG----------------ACCGTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--GATGTAA-CTG---AGAGCGCGGA---TTCCCT-GCCCC-GGCCCCGG-GAAGC----AGGGGG-TCT-AGCGCGCTT-----GCTCA----CRATGC-ACAACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCACCG-TAGGCCCCCC-----AGCACCGAAGAGGCAAA---------------------GGTC--GAGCTCCYGAAGCGG-CGGGCAA-----TTTTG---TCTGC-------TT-GCG-CGGGCG---------GCTG-GGATTTGCATCTC----GTTTGGA-----GAGGCGGCAAGGTTGGCCTCCCGCGCGGGC----AACA-GCGC--GGTTGGTCTAAAGTAGAGTGCTTGGCAT-CGCGCGGTGTGATCGACGGTGGTTTCCTT--ACCCTCGGCGAATAAC--ATCGCACGCGCGGGATGCT-GAGCGGAGAA-CTCC-ATTGG-------GACCC--------TTCC-TTGTGA-TGCCTTTCT-----CGCGGTTCCCCT-------GCGTGGAGGCGTTGC-----CGAAGC--GACCCCAG-TCAGGHGG-AT-ACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_potamophila               --------TCGATGACTGCA--------GCAG-AACAA-CCGCG-AACTGGTTAAA--AACCATCTGC-TTCCCCAGCCA-GCAAAT--GGGGAA-AGGCGGGGC-GCGCTGGTG-A--CGTTKAC--CGC-GCCCC--------GCGCGCGGCCG--GCC--CAGC-GCACCTCGCGG----------------ACCGTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--GATGTAA-TCG---AGAGTGCGGA---TCCCAT-GCCCC-GGCCCCGG-GAAGC----AGGGGG-TCT-AGCGCGCTT-----GCTCA----CGACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCACCG-TAGGCCCCC-------GGACCGAAGAGGCCAA---------------------GTTC--GAGCTCCCAAAGCGT-CGGGCAG------TTTC---TCTGC-------TT-GCG-TCGGCGGCG------GTTG-GGATTTGCATCTC----GTTCGGA-----GGGGCGGCAAGGTTGGCCTCCCGCGCGGGC----AGTG-GCGC--GGTTGGTCTAAAGTAGAGTGCTCGGCAT-CGCGCGGTGCGATCGACG-TGGTTTCCTT--ACCCTCGGCGAATTAC--GTCGCACGCGCGGGATGCG-GCGCGGAGAA-CTCC-ATTCG-------GACCC--------TTCTTTTACGA-TGCCTTTCT-----CGCGGTTTCCCC-------GCGTGRAGGYGCTGC-----CGAAGC--GACCCCAGGTCMGG-GG-ATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_poculifera                ---TCTTGTTGATTCCTGC-ARAAA---GCAG-AACAACCCGCG-AACTTGTCGTG-----CAACGA--ACCC--------CGCAA----GGGAA-TGGCGGGG-GCGTGCTGGGG----------------------------CGCTA---------------GCGCGCTCCTCGCGCTC-------CCCGCGCACCTTAACGAACCCCC-GGCGCGAGTTGCGCCAAGGAA--TCTTTAACTTG---AGAATGTGGC---TCCTCT-GCCCCCGGTTGCGG-GATGC----AGGGG--CGC-TGCGAGCTT-----GAAAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TGGCCCCCCACGCTCC-CGCCAAAAAATC-----------------GAGT---TCAA--TCTCCTAC-----------------------------------------T-GGGGTTT------------GGCC-GGACTTT--------GGTTGGGGAC----GAGGGGGCAATGTTGGCCTCCCGTGCGAGC----GATT-GCGC--GGTTGGCCTAAAGTTGAGCCCTCGGCGT-CGCGCAGTGCGATCGACGGTGGTTTTTA---GCCCTCGGCGAGATAC--ATTGCACGTGCGAGATGCGGTATCGG-GAG-CTCC-CACGY-------GACCC--------TGC-TGCGCTGCCTC-GTCGGTT----CCTCGGAGAAG------GACGAGGAGGCGCTAT-----CGAAGC--GACCCCAGGTCAGGYGGGATTACCCG-------------------------------CCARATCGGGCGGTAAATCCTGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAAGAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGTGCCTTGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCAGGGCGCGGACCGACGCGGATTGCGGCGGCGGCCCAAGCCTGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGGCGCGATCGTGGCTGGCAGCGCGCGCCAAGAGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGTTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTA-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGAGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_palmata                   -CATTGT-CGATTCCTG-C-AGAAT---GCAG-AACAACCCGTG-AACTAGTTT-G----CCAACG---TCCCGCCC----CGTGC----GGGAA-CGGCGCAGA-GCACGTTGG-AACTGTGGAA--AGC-GCTT---------TTGCGCGACCT--AAG--AGACA-TGCTCTGCGCCCCC----------------TAACGAACCCC--GGCGCAAGTCGCGTCAAGGAA--TCTGAAA-CTT---AGAACGCGGA---TCCCCT-GCCCT-GATTTCGG-GATGC----AGGGG--TGA-CGTGGACTT-----GATCA----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCCAAACCAAAAAAA----------CCAATAC--CAA---TCCATC--------GATC--GAAGCA---CACTC---GTGCT-GCTTC--GTTGGTG-----GTTTTGGTGGCT-GGTCTTT-----------GGGGGTC----GGGGAGGCGACGTTGGCCTCCCGTATGGGC----AACC-TCCC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--ATTGTGCTTGCGAGGCGCGCCGTGGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGACTC-GTC---------GGACAACACC------GACGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGTGGGATTACCCG-------------------------------CCAAATCGGGCGGTAA-TTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGTGCG-TAACCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCT-ACGGCGTGCTTCGGCATCAGCGTGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTTAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTG-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_olbia                     TCATTGT-CGAC-YCTG-C-AAAAT---GCAC-AACAACCCGTG-AACTAGTATG-----CCAACG---TCTCGCTT----CGCGC----GGGTA-WGGCGCGGG-GTGCGTCKG-AGCTGCGGCT--CGC-TTTT---------GCGCGGGCCCA--GCC--TCKC-GCGCCATGCGCCCAC----------------TAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TCTCAAA-CTT-ATAGAACGCACT---TCCCT--GCCCC-AAATTTGG-GATGC----AAGGG--TCC-TGCGAACTT-----GATCA----TACCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCAAAGGAAAAAA------------CCATGA--YCGA---TCCCTC---------TTC--GAAGTA---GACTG---GTGCT-GCTTT--GTTGGGG-----CTT-----GGCT-GGCCTTT----------GGGTGGTC----GGGGTGGCAAAATTGGCCTCCCGTG-GAGC----AACC-TCGC--GGTTGGTTTAAATTCGAGTCCTCGGCGT-CTTGGGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--ATTGCGCTTACGAGACGCGTGCTGGG-GAA-CTCC-TGGAC-------GACCC--------TAA-TGCGTCGGCTA-GCCGG---------ACAACGCC------GGCGAGCAGGCGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGAT-ACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_obliqua                   TCATTGC-CGATTCCTG-C-AAAAT---GCAG-AACAACCAGCG-AACTAGTT-GC----CGAAT----TCCCGCAT----TGCGC----GGGA-------------CGCGTGGG-AGGTCGTGGT--CAC-TTTT---------GTGCGCCCACG--GCC---GGC-ACGCCTCTCGCCCCC----------------TAACGAACCCC--GGCGCGAGCTGCGTCAAGGAA--TGTGTAA-CTT---AGAACGCTCA----CCCTT-GTCCG-GATTTTCG-GATGC----AGGGT--TAGTTGCGAACTG-----GATCAAA--TATTGA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-GAGCCCCCC-------GCACCAAAAGAAAAAA------------CCAA---GTCAGT--TCCGTG--------CGTC--GAAGTA---GACTT---GTGCT-GCTGC--TTTG--------CTC---GGGGCA-GA-TGGGA--------GAA-TGGTC----GGGGCGTAGAGAATGGCCTCCCGTGCGAA-----ATCCATCGC--GGTTGGTCTAAAACTGAGTCCTCGGCGT-CTTGCGGCGCAATCGRCGGTGGTTGACAT--GCCTTCGACGAAT-AC--ATTGCGCTGGGAAGGCGCGGTTCCGG-GAA-CTC--TGGAG-------GACCCC--------AG-TGCGTCGGCGA-GTC---------GGACAGGAAC------GATGAGCAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG------------------------------CCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGAT-GAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCCCGATCGTGGCTGGCAGCGTGCGCCC-TCGGCGTGCTCAGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGAGTGGCGGGATCCCCTC-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_nossibea                  ------T-CGATTCCTGMC--GAAA---GCAG-AACAACCCGTG-AACTAGTTAA-----CCATCG---TCCC--------CATGT----GGGAA-TTGCGAGGC--ATGTTCG--AGGTACGGCATA----GCTT---------CCGCGTGCCTT--GCC--TTGC-ATGCCTCGGGA----------------ACCATAACGAACCCC--GGCGCTAGTTGCGTCAAGGAC--TCTTTAA-CTG---AGAACGCTGC---TCTCCTAGCCC--GACTTCGG-GATGC----AGGGG--TGC-TGCGAACTT-----GCTTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACACATCG-TAGC-------------CACCCACACCAAAGCTTTGAACGCTCAATATCTCTGTTTA--AATAGACCTATGCTATTTTTCTA----------------------------GGGGTTG------------GGCT-GGCAAT----------GGGTTCGAT----GAGGTGGTCATGTTGACCTCCCTTGCAAGC----AAGT-GCAT--GGTTGGTCTAAATTTGAGTCCTCAGCGT-TGCGCAGTGCGATCGACGGTGGTTGCCAT--GCCCTCGGCGAATTAC--ATTGCACTTGCGGGATGCATGTTTGA-GAA-CTCC-TTTGC-------GACCC--------AAT-TGTGCTGCCTT-GTTGGTT----CCTTCGAGAAC------AATGAGAATACGTTAT-----CGAAGC--GACCCCAGGTCAGGYGGGATCACCC------------------------------------ATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGCGTC-TAGCCGGTCTGCCAATCGACTCAGGGCGTGGACCGACGCGGATTGTAGTGGTGGCCCAAGCCTGAGCCTTTGTTGTGCTTGTGGAGACGTCATCACCGCGATCGTGGCTGGCAGCGCGCGCCT-ATGGCGTGCTTCGGCATCAGCGTGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GTGGGT-TGCACCGCCGACCGACCTTGATCTTTTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAA-ATGGTGAACTATGCCTGA---------
		Begonia_molleri                   -----TTGTCGATTCCTGCM--AA----GCAG-AACAACCCGCG-AACCCGTTGTG-----CAACGA--ACCC--------CGCAA---GGGGAG-CGGCGAGGA--GTGTTGGG-------------------------------------------------CGC-ACGCCCCGCATTC-------CGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TCTTTAA-TTG---AGAACGCGGT---GCCTCT-ACCCCCGTTTGCGG-GATGC----AGGGG--CAC-TGTGAGCTT-----GCCAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-CAGCCCCCC-------GCGCCAAAAAAA-------GC----------------TCAA--TCTC-TTCTTCCGG----------------------------------TT-GGGGTTG------------GGCC-GGACTTT----------GGTCGGAC----GAGGGGGCAATGTTGGCCTCCCGTGCGAGC----GATC-GCGC--GGTTGGCCTAAAGTCGAGTCCTCGGCGT-CGCGCAGTGCGACCGACGGTGGTTTTTG---GCCCTCGGCGAAATAC--GTTGCACGTGCGAGACGCGGTATCGG-GAA-CTCC-CACGC-------GACCC--------TGC-TGCGCTGCCCC-GTC------------GGAGAAC------GACGAGGGGGCGCTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATCACCCG---------------------------------AAATCGGGCGGTAAATTCC-TCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGTGT--TAGCCGGTCTGCCAATCGACTCAGGGCGCGGACCGACGCGGATTGCGGCGGCGGCCCAAGCCTGAGCCTTTGTTGTGCTTGCGGAGACGTCGTCGGCGCGATCGTGGCTGGCAGCGCGCGCCAGTAGGCGTGCTTCGGCATCGGCGCGCTCCGGGCGTCGGCCTGCGGGTTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTA-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGAGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGA---------
		Begonia_meyerijohannis            --TCATTGTCGATTCCTGM-GAAA----GCAG-AACAACCCGTG-AACTTGTTGA------CACCG---TCCC-------ACGC------GGGAA-CGGCGGGGCGCGTCTGGG---------------------------CTGCGCACACKCTCKCG---------TGCGCCCGGCCTCGCACGCCTCGC-CG--CTTTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TCTTTAA-CTG---AGAACGCGGC---TCCCCT-GCCCC-GGCTCCGG-GATGC----ACGGG--TGC-TGCGAACTT-----ACTTACTA-T-ACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACACATCG-TATCCCCCCC------GCACCAAACAAG----------------------------------------------------------------------------------------------------------CAAGTTT----------GGCCAGAC----GGGGCGGTAATGTTGGCCTCCCGTGCGA-C---TGCTC-GCAC--GGTTGGCCTAAAGTCGAGTCCTCGGCAT-CGCGCGGTGCGATCGACGGTGGTTGCCAT--ACCCTCGGCGAATTAC--ATTGCACTTGCGAGTTGCGCGTTGGG-GAA-CTCT-GCCAA-------GACCC--------TGC-TGCGTCGTCTC-GTCGGTT----CCTCGGGGATC------GACGGGGRGACGTCAT-----CGAAGC--GACCCCAGGTCAGGYGGGAT-ACC-------------------------------GCCCAAATCGGGCSSTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGCGTC-GAGCCGGTCTGCCAATCGACTCAGGGCGTGGACCGACGCGGATTGCGGTGGTGGCCCAAGCCTGAGCCTTTGTTGTGCTTGTGGAGACGTCGTCACCGCGATCGTGGCTGGCAGCGCGCGCCT-TCGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCTAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_mannii                    -----GT-CGATTCYCG-CWAGAA----GCAG-AACAACCCGCG-AACTCGTTGTG-----CAACGA--ACCC--------CGCAA---GGGGAG-TGGCGAGGA--GTGTTGGGG------------------------------------------------CGC-AGGCCCTGCATTT-------CGCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TATATAA-CTG---AGAACGCGGT---TCCTCT-GCCTC-GCCTGCGG-GATGC----AGGGG--CGC-TGTGAGCTT-----GCGAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCAAAAAAA------TGC----------------TCGA--TCTC-TTC----------------------------------------TT-GGGGTTG------------GGTT-GGACTTT----------GGTCGGAC----GAGGGGGCGATGTTGGCCTCCCGTGCGAGC----GATT-GCGC--GGTTGGCCTAAAGTTGAGTCCTCGGCGT-CGCGCAGTGCGACCGACGGTGGTTTTTA---GCCCTCGGCGAAATAC--GTTGCACGTGCGAGATGCGGTATCGG-GAA-CTCCCTTTGC-------GACCC--------TGC-TGCGCTGCCCC-GTC-GTT-----CTAC--GAAC------GATGGGGAGGCGCTAT-----CGAAGC--GACCCCAGGTCAGGCGGGA--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_mananjebensis             --------CGATTCCTGMC--GAAW---GCAG-AACAACCCGTG-AACTAGTTAA-----CCATCG---TCCC--------CATGC----GGGAA-TTGCGAGGC--ATGTTCG--AGCTACGACACA----GCAT---------TCGCGTGCCTT--GCC--TTGC-ATGCCTCGTAA----------------ACCATAACGAACCCC--GGCGCGAGTTGCGTCAAGGAC--TCTTTAA-CTG---AGAACGCCGC---TCTCCTAGCCC--GACTTCGG-GATGC----AGGGT--TGC-TGCGAACTT-----GATTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGC-------------CACCCACACCAAAGC-----ACGCTCAATCTCTCGGTTGA--AATAGACCAATGCTATTTTTCTA----------------------------GGGGTTG------------GGCT-TGCAAT----------GGGTTCGAT----GAGGTGGTAATGTTGGCCTCCCGTGCAAGC----AAGT-GCAC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-TGCGCAGTGCGATCGACGGTGGTTGCCAT--GCCCTCGGCGAATTAC--ATTGCACTTGCGGGATGCATGTTCGA-GAA-CTCC-CTTGC-------GACCC--------AAT-TGTGCTGCCTC-GTTGGTT----CCTTCGAGAAC------AATGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGG-GGG-TCACCC--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_madecassa                 TCATTGT-CGATTCCTG-C-AAAAA---GCAG-AACAACCTGCG-AACTAGTTAA-----CCATTG---TTCC--------CACGC----GGGAACTTGCGAGGC--ATGTAAT--TGCT-TGGCACAA---GCTT---------GCGAGTGCCTT--GCC--TTGC-ATGCCTTGCGA----------------ACCATAACGAACCCC--GACGCGAGTTGCGTCAAGGAA--TCTTTAA-CTG---AGAACGCCGC---TCTCTTAGCCC--GACTTCGG-GATGC----AGGG---TGT-TGCGAACTT-----GCTTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACACATCG-TAGC-------------CACCCACACCAAAGCTT-GCACGCTCAATCTCTA-----------------------------------------------------------GGGGTTG------------GGTT-GGCATT----------GGGTTCAAT----GAGGTGGTAATGTTGGCCTCCCGTGCGAGC----AAGT-GCAT--GGTTGGTCTAAATTTGAGTCCTCGGCGT-TGCGCAGTGCGATCGGCGGTGGTTGTCAT--GCCCTCGGCGAATTAC--ATTGCACTTGCGGGATGCATGTTCGG-GAA-CTCC-TTTGT-------GACCC--------TAT-TGCGTTGCCTC-GTTGGTT----CCTCGAAGAAC------AACGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATCACCCG---------------------------------AAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGCGTC-TAGCCGGTCTGCCAATCGACTCAGGGCGTGGACCGACGCGGATTGCAGTGGTGGCCCAAGCCTGAGCCTTCGTTGTGCTTGTGGAGACGTCATCACCGCGATCGTGGCTGGCAGCGCGCGCCT-CTGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACT----------------------------------------------------------------------------------------------------------------------
		Begonia_luxurians                 TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGCG-AACTAGATTGG----CCAATG---TCCCGTTT----CGCAC----GGGA--CGGCGCGGG-GCGTGCTGG-TGCTGTGGCC---GC-CTCT---------GTGCGCGCCCT--GCC--TAGC-AGGCCTCGCGCTCCCC--------------TTAACAAACCCC--GGCGCGAGTAGCGTCAAGGAA--TCTCGAA-CTT---AGAACGCGCC---TCCCTC-GCCCC-AATTTCGG-GGTGC----AGGAG--TGT-CGCGAACTT-----GATCATA--CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------ACACCAAAAATAA----------------GAAGGGGTCAA---CGTCTC--------GATC--GAAGCA---GACTC---GCGCT-GCACC--GTCGCGA-----CTT-----GGCTCGCCCTT-----------GGGCGGTG----GTGGTGGCGACATTGGCTTCCCGTGGGGGC----GAC--TCGC--GGTTGGCCTAAATTTGAGTCCTTGGCGTTCTTGCGGCGCGATCGACGGTGGTTTCCAT--GCCCTCGACGAATCAC--ATTGCGCTTGCGAGATGCGTGCTACT-GAA-CTCT-TGGAC-------GACCCTC---------------------------------------------------------AAATGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------CCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGAG-TAGCCGGTCTGCCGATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGACTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-TTGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCATAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_loranthoides              ------TGTCGATTCCTGC-ARAAA---GCAG-AACAACCCGCG-AACTTGTCGTG-----CAACGA--ACCC--------CGCAA----GGGAA-TGGCGGGGGCGTGATGGGG---------------------------------------CG--CTC--GCGC-GCTCCTTGCGCTC-------CCCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TCTTTAA-CTG---AGAACGCGGC---TCCCCT-GCCCCCGGTTGCGG-GATGC----AGGGG--TGT-TGTGAGCTT-----GAAAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TGGCCCCCC-------GCGCCAAAAAAAAAAAC-T-----------GAGT---TCAA--TCTC-TTC----------------------------------------TT-GGGGTTG------------GGCC-AGACTTT----------GGTCGGAC----GAGGGGGCAATGTTGGCCTCCCGTGCGAGC----GATC-GCGC--GGTTGGCCTAAAGTTGAGTCCTCGGCGT-CGCGCAGTGCGATCGACGGTGGTTTTTT---GCCCTCGGCGAGATAC--ATTGCACGTGCGAGACGCGGTATCGAGAGCGCTCC-CACGC-------GACCC--------TGC-TGCGTTGCCTC-GTTGATT----CCTCGGAGAAG------GACGAGGAGGCGCTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACC---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_longipetiolata            ------TGTCGATTCCTGCCGAAA----GCAG-AACAACCCGCG-AACTCGTAGTG-----CAACGA--ACCC--------CGCAA---GGGGAT--GGCGAGGC--GTGATGGGG----------------------------CGCTC---------------GCCCGCTCCTCACGCTC-------CCCGCCCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAARATCTTYAA-TTG---AGAACACGGC---TCCTCT-GCWCCCGGCTGCGG-GATGC---WAGGGG--CGT-TGCGAGCTT-----GAGAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACSTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTKGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TGGCCCCCC-------GCGCCAACAAAAAAAACC-----------CGAGT---CCAA--TCCC-TTA-TTATTA-------------------------------TTTT-GGGGTTG------------SACC-GGACTTTT---------GGTTGGAC----GAGGGGGCAATGTTGGCCTCCCGTGCGAGC----GATC-GCGC--GGTTGGCCTAAAGTTGAGTCCTCGGCGT-CGCGCGGTGCGATCGACGGTGGCCTGTAT-AGCCCTCGGCGAGATAC--ATTGCACGTGCGAGACGCGGTATCGG-GAG-CTCC-CACGC-------GACCC--------TAC-TGCGCTGCCTC-GACCCTT----CTTGAGGGAAGGG--AAAGAGAAGGCGCGCTAT-----CGAAGC--GACCCCAGGTCAGG-GGGAT-ACCC--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_letouzeyi                 -------GTMGATGCTA------------CAG-AACGACC-GCG-AACT-GGTTAT-CAACCACACGA-TTCCCCAGCCA-GCGCT---GGGGAA-AGGCGGGGC-ACGCTGGTC-T--CGCATTG--CGC-GCTCT--------GCGGG---------------------CCTCGCCG----------------ACCGCAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--CATGAAA-CTG---AGAGCGCGGA---TCCCCT-GCCCC-GGCTCCGG-GYAGC----AGGGA--TCT-CGCGCGCTT-----GCTCA----CGACGC-ACGACTCTCGACGACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGTCCCCC-------TCACCGAAGAGGCCAA---------------------ATTC--GAGCACCCAGAGC---AGTGCAG------TC-----TCTGCAC-----TT-GCA-TCGGGC---------GCTT-GGATATGGAGCTC----GTTTGGA-----GAGGCGTCAAGGTTGGCCTCCCGCG-GAGC----AACC-TCGC--GGTTGGTCTAAAGTAGAGTGCTCGGCAT-CGCGCAGTGCGATCGACGGTGGTTTCGTA--GCCCTCGGCGAATTAC--GTCGCAC-----AGATGCG-GCGCGGAGAA-CTCC-ATACA-------GACCC--------TGC--TGATGATSGCC-GGCT-----CGTGGGTTCCCC-------GCGAGAAGGCACTGC-----CGAAGC--GACCYCAG-TCAGG-GG-ATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_kisuluana                 ----TGT-CGATTCCTG-M-CGAAW---GCAG-AACAACCCGCG-AACCYGTTGTG-----CAACGA--ACCC--------CGCAA---GGGGAG-CGGCGAGGA--GTGTTGGGG------------------------------------------------CGC-ATGCCCCGCATTT-------CGCGCACACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAA-CTG---AGAACGCGGC---TCCTCT-GCCCC-GCCTGCGG-GATGC----AGGGG--TGC-TGCGAGCTT-----GAGAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCGCCAAAAAA-------TGC----------------TCGA--TCTC-TTC----------------------------------------TT-GGGGTTG------------GGCT-GGACTTT----------GGTCGGAC----GAGGGGGCGATGTTGGCCTCCCGTGAGAGC----GATC-GCGC--GGTTGGCCTAAAGTCGAGTCCTCGGCGT-CGCGCAGTGCGACCGACGGTGGTTTTTA---GCCCTCGGCGAAATAC--GTTGCACGTGCGCGACGCGGTATCGG-GAA-CTCC-CACGC-------GACCC--------TGC-TGCGCTGCCCT-GTC-GTT-----CTCC--GAAC------GACGGGGAGGCGCTAT-----CGAAGC--GACCCCAGGTCAGGYGGGA-TACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_johnstonii                GATCATTTTCG-TTCCAGC-AAAAA---GCAG-AACAACCCACG-AACTGATTAAA-----CAACG---TCTCTCGC-----TCGG----GGGAACGGAACGGTGCATGCTAA---GATAGAGGCA----T-GCAC-------GTGCACGCACCCT--CCC--TGGC-GAGCCCCGCG----------------GACCCTAACAAAGCCT--GGCGAGAGTCGCGTCAAGGAA--TCTTGAA-CTT---ATAACGCAAT---TCCTTT-TCCCT-GGCTTCGG-GACGC----GGGGG--CTA-TGCAAACTT-----GATCAAGTATCAAAC-ACGACTCTCGACAATGAATATCTCGGCTCTCGGATAGATTAAGAACGTAGTGAAATGCGGTACTTGGTGTGAATTTCATAATCCAGTGAACCATCGAGTTTTTGAATGCAAGTTGCGCCCGAAGCCTTCTGGCTAAGGGCACGTCTGCYTGCGCGTCACGCATCG-TAGCCCCCC---------ACCAAAAACCACTCGC----------------------------------------------------------------------------------------------------GGGCTTTT---------GCTGMTT-----GAGATGGCGACGTTGGCCTCTCGTGAGGGG----AACC-TCAC--AGTTTGTCTAAATTAGAGTCCTCAACTT-CACACAGCTCGATCGACGGTGGTTGCAAT--TCCCTCAGTAAATCAC--ATTGCGCTCGCGAGACGTGCCTACAAGGAA-ATCG-TTCGC-------GACCC--------TAC-TGCGTAGAGTCGCTGCTAC----CTCTAGGTGCGGTSSMGTC------GGCGTCA------TCAAGC--GACCCYTG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_iucunda                   ------TGTCGATTCCTGMCVAAA----GCAG-AACAACCCGCG-AACTCGTT----AATAATCGCG--TTCCCCACGCA-----------GGAA-CGGCGGGTCGTGCCGGGGC-TGCGGATCGCC-------------GCTGTGCGCTCCACGCC------TGGC-ACGCCTCGCAC----------------ACCCTAACGAACACC--GGCGCAGGTCGCGTCAAGGAG--CATTTAA-TTG---AGAACGCAAT---TCCTCTTGCCCT-ACCACGGG-GAT-----AAGGGG---TT-TGCGAACCC-----TCTAA----AAACAC-ACAACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTCTTGGCTGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCAAACAACAA-----------------------GT-C--ACGCTCCACATGTCAGTAGGCTAGT----TCTA---CTTGCG------TT-GGAGTT-------------GGCT-GGCATATT---------GCTTGGT-----GGGGTGGTAACGTTGGCCTCCCGCGCGAC-----GAAATGCAC--GGTTGGCCTAAAAATGAGTACTCG-CAT-CGCGCAGTGCGATCGGCGGTGGTTTCCAT--GCCCTCGGCGTATTTC--ATTGCACCTGCCAGATGCATGTTTTGAGTA-GTCC-----G-------GACCC-------TTGC-GCGCGTGCTGCTCGATTT-----CATT-GAGAAC------GAGGAGAAG-CGCTAT-----CGAAGC--GACCCCAGGTCAGGCGGGACTACCCG--ACTTAGATCGGGTTGTTTGGGAATGCAGCCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTCGGTCGGATGTGGAACGGCGTC--AGCCGGTCTGCCAATCGACTCGGGGCGTGGTCCGACGCGGATTGMGATGGCGGCCCAAGCCCGAGCCTTTGTTGWGCTMGTGGAGAWGTCGTCGGCGCGATWGTGGCTGGCAGCGCGCGCCC-CTGGCGTGCTTCGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGCGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACMGCCGACCGACCTTGATBTTCTGAGAAGGGTTCGAGTG-----------------------------------------------------
		Begonia_integerrima               ---TTGT-CGATGCCGG-C-AGAAT---GCAG-AACAACCCGCG-AACTAGTTT-G----CCAACG---TCCCGCTG----CGCGC----GGGAA-CGGCGCCTG-GCGCGTCGG-AGCTGCGGCC--CGC-GTTC---------GCGCGCGCCCT--GCC--TCGC-GCGCCTCGCGCCCCCC---------------TAACGAACCCC--GGCGCGAGTCGCGCCAAGGAA--TCTCAAA-CTTTAAAGAACGCGGCGGCTCCCTT-GCCCC-GATTTTGG-GATGC---AAGGGG--TCC-TGCGGACTT--GACGATCGTCG-TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACAAAAG----------------------TCATCSG-----TCCCTC--------GATC--GAAGCA---GACCC---GTGCT-GCTTC--GTCGGGG-----CTC-----GGCC-GGCGTCT----------TGGTGGCA----GGGGCGGCGACATTGGCTTCCCGTGGGGGC----AACC-TCGC--GGTTGGTCTAAATTCGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATCAC--GTTGCGCTTGCGAGGCGCGCGCTCGG-GAA-CTCA-TGCAC-------GACCC--------TAAAAGCGTCTGGCA-GCC---------GGACAACACC------GGCGAGCAGGC-TTAT-----GGAAGG--GACCCCAG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_incarnata                 --------CGATTCCTG-C-ARAAT---GCAG-AACAACCCGCG-AACACGTTT-G----CCAACT---TCCTGCTT----TGCGC----GGGAA-TGGTGCGGG-GCGCGTCGG-AGTTGTGGCT--TGC-TTTT---------GCGCGCTCACT--GCC--TCGC-ACGCCTCGTGCCCCC----------------TAACGAACCCC--GGCGCGAGTCGCGCCAAGGAA--GCTTATA-CTT---AGAACGCGGC---TCCCTT-GCCCC-GATTATGG-GATGC----AGGGG--TCC-TGTGAACTT-----TATCA----TATTGA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCAAAAAAAAA--------------CCAACAC--CAA---ACTATC--------GATC--GGAGTA---GTCTC---GTGCT-GCTTC--GTCGGGG-----CTT-----GGCT-GGCCTTT----------GGGTGGTC----GGGGTGGCAACATTGGCCTCCCGTTGGGGC----AACC-TCGC--GGTTGGTCTAAATCTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCTTCGACGAATTAC--ATTGCGCTGGCGAGACGCGCGCTCGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGGCTA-GCCGG---------ACAACACT------GGCGAGCAGACGTTAT-----AGAAGC--GACCCCAGGTCAGGYGGGAT-ACCC-------------------------------CCCAAATCGGGC--TAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-ACGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGGAAACCCGTGAGGCGCAAGGAARCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_horticola                 ---TTGT-CGATTCCTG-C-MGAAA---GCAG-AACAACCCGCG-AACCCGTTGTG-----CAACGA--ACCC--------CGCAA--GGGGGAT-CGGCAAGGA--TTGTTGGGG------------------------------------------------CGC-AAGCCCCGCATTY-------CGCGCGCACCTTAACAAACCCCCCGGCGCGACTCGCGTCAAAGAA--TATTTAA-CTG---AGAACGCGGC---TCCTCT-GCCCC-GCCTGCGG-GATGC----AGGGG--CGC-TGCGAGCTT-----GAGAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCGCCAAAAAA-------TGC----------------TCGA--TCTC-TTC----------------------------------------TT-GGGGTCG------------GGGA-GGACTTT----------GGTCGGAC----GAGGGGGCGATGTTGGCCTCCCGTGCGAGC----GATC-GCGC--GGTTGGCCTAAAGTCGAGTYCTCGGCGT-GGCGCAGTGCGACCGACGGTGGTTTTTAT-AGCCYTCGGCGAAATAC--GTTGCACGCGCGAGACGCGGTATCGG-GAA-CTCC-CACGC-------GACCC--------TGC-TGCGCTGCCCT-GYC-GTT-----CTCC--GAAC------GACGGGGAGGCGCTAT-----CGAAGC--GACCCCAGGTCAGGHGGGAT-ACCC--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_holtonis                  TCATTGC-CGATTCCTG-C-AAAAT---GCAG-AACAACCAGCG-AACTAGTTTGT----RGAAT----TCCCGCAC----TGCGC----GGGA--CGGTGCGGG-GCGTGACGG-ARGTGGTGGC--CGC-TTTT---------GCGCG-GCCCA--GCC--TCGC-ACGCCTCTCGCCCCC----------------TAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TGTGTAA-CGT---AGAACGCTTG----CCCTC-GTCCC-GATCTTGG-GATGC----AGGTG--TAGTTGCGAACTT-----GATGAAAA-TATCGA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTGCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGCCCCC--------GCACCAAAAGAAAAA-------------CCAC---GTCAGT--GGCGTG--------CAAG--GAAGTA---GACTT---GTGCT-GCGTC--TTTG--------CTC----GGGCT-GGTTGGGA--------GAGGTGGTC----GGGGCGCAGACATTGGCCTCCCGTGCGA------AATCGTCGT--GGTTGGTCTAAAACGGAGTCCTCGGCGT-CTCGCGGCGCGATCGACGGTGGTTGACAT--GCCCTCGACGAAT-AC--GTTGCGCTCGCAAGATGCGGTTTGGA-GAA-CTC--TTGAC-------GACCC--------TAG-TGCGTCGGCGA-GTC---------GGAGAAGAAC------GACGAGCAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGAT-ACCCG------------------------------SCTAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATAATTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGAT-GAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTGGCCTCGATCGTGGCTGGCAGCGCGCGCCG-TTGGCGTGCTCAGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGAGTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGA---------
		Begonia_grandis                   --TCATTGTCGATTCCTGC-ARAAT---GCAG-AACAACCCGCG-AACTAGTTT-G----CCAACG---TCCCGCCT----CGTGC----GGGAA-CGGCGTTGG-GCGCGTCGG-AACTGCGGCC--TGC-GCTT---------GCGCGCGCCCT--GCC--TCGCG-CGCCTCGTGCCCCC----------------TAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TCTGTAA-CTT---AGAACGTGGC---TCCCCT-GCCCC-GATTTTGG-GACGC----AGGGG--TCC-TGCGAACTT-----GATCA----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCCAAAAAAACA-------------CCAGGAT--CAA---GACCTC--------GATC--GAAGAAGTAGACTC---GTGCT-GCTC---GACGTTGTG---GTTTT----GCT-GGCCTTT----------GGGCGGTC----GGGGAGGCGACGTTGGCCTCCCGTGAGGGC----GACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--ATTGTGCCTGCGAGACGTGCGCTCGA-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGACTC-GTC---------GGGCAACACC------RGCGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGC-GGATTACCCG--------------------------------CAAATCGGGCGGTAAATTTCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAATAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCT-ACGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTTMGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAAC-------
		Begonia_goegoensis                TCATTGT-CGTTTCCTG-CAAAAAT---GCAG-AACAACCCGTG-AACTAGTT-TG----CCAACG---TCCCGCTG----CGTGC----GGGAA-CGGCGTGGA-GCGCGTCGG-AGTTGCGGCT--CGC--ATT--------CGTGTGCGCCCT--GCC--TTGCA-CGCCTCCCGCCCCC----------------TAACGAAACCC--GGCGCGAGTTGCGTCAAGGAA--TCTGGAA-CTT---AGAATGCAGC---TCCCCT-GCCCC-GATTTCGG-GACG-----AGGGA--TCT-TGCAAACTT-----GATCG----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACC-AAACAAAATAGAAA--------CCAGCATMGACY---TCTC-C--------GACC--GAGGCAA--GATTC---GTTCTCGCTTC--GTTGGGGGG---GTCTT----GTT-GGCCTTT----------GGMTGGTTG---GGGGAGGMGACGTTGGCCTCCCGTGCGGGC---AAACY-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGYTGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAA--GTTGTGCTGGCGAGATGCGGGGTGGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGACTC-GTC---------GGACGATAAT---GCCGGCGAGTAGACGTCAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_geranioides               TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGTG-AACTAGTTTG-----CCAATG---TCCTGCCC----CGCGC----GGGAT-TGGCGCGGG-GCGCTTCGG-AGCTGTGGCT--CGT-GCTT---------GCGCGCGCCCT--GCC--TCGA-ACGCCTCGCGTCCCC----------------TAACGAACCCC--GGCGCTAGACGCGTCAAGGAA--TCTCGAA-CTT---AGAACGCGGC---TCCCCT-GTCCC-GATTTTGG-GACGC----AGGGGTGTCC-TGCGAACTT-----GAACA----CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC----ACCGCACCCAAAAAGAAA-----------CCAA------GATCT--TCCCTC----GATC-ATCGGG-------------GGTAG--GTT--------------------------GGCT-GGCCTTT----------GGGTGCTC----GGGGCGGCAAAGTTGGCCTCCCGTGTGGGC----AACC-TCGC--GGATGGTCTAAATTTGAGTCCTCAGCGT-CTCGCGGCGCGATCGACGGTGGTTTCCAT--GCCCTCGACGAATTAC--ATTGCGCCTGCGAGATGTGCGCTCGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGACTC-GTCGG---------AAAACGCC------GGCGAGTGGATGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG----------------------------AGCCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGTT-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-ATGGCGTGCTTCGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGAGTGGCGGGATCCCCTC-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCG------
		Begonia_gabonensis                -----TTGTCGATTCCTGCMMAWA----GCAG-AACAACCCGCG-AACCCGTTGTG-----CAACGA--ACCC--------CGCAA---GGGGAG-CGGCGAGGA--GTGTTGGGG------------------------------------------------CGC-AAGCCCCGCATTT-------CGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAA-CTG---AGAACGCGGC---TCCTCT-GCCCC-GCCTGCGG-GATGC----AGGGG--CGC-TGCGAGCTT-----GAGAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCGCCAAAAAA-------TGC----------------TCGA--TCTC-TCC----------------------------------------TT-GGGGTCG------------GGCT-GGACTTT----------GGTGGGAC----GAGGGGGCGATGTTGGCCTCCCGTGCGAGC----GATT-GCGC--GGTTGGCCTAAAGTCGAGTCCTCGGCGT-CGCGCAGTGCGACCGACGGTGGTTTTTATTAGCCCTCGGCGAAACAC--GTTGCACGTGCGAGACGCGGTATCGA-GAA-CTCC-CACGC-------GACCC--------TGC-TGCGCTGCCCG-GTC-GTT-----CTCC-GGAAC------GACGGGGAGGCGCCAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCC--------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_francoisii                -----TY-CGATTCCTGAC--GAAA---GCAG-AACAACCCGTG-AACTAGTTAA-----CCATCG---TCCC--------CATGT----GGGAA-TTGCGAGGC--ATGTTCA--AGGTACGGCATA----GCTT---------TCGCGTGCCTT--GCC--TTGC-ATGCCTCGCGA----------------ACCATAACGAACCCC--GGCGCTAGTTGCGTCAAGGAC--TCTTTAA-CTG---AGAACGCTGC---TCTCCTAGCCC--GACTTCGG-GATGC----AGGGG--TGC-TGCGAACTT-----GCTTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGC-------------CACCCACACCAAAGCTTTGAACGCTCAATCTCTCGGTTTA--AATAGACCTATGCTATTTTTCTA----------------------------GGGGTTG------------GGCT-GGCAAT----------GGGTTCGAT----GAGGTGGTCATGTTGACCTCCCTTGCAAGC----AAGT-GCAT--GGTTGGTCTAAATTTGAGTCCTCAGCGT-TGCGCAGTGCGATCGACGGTGGTTGCCAT--GCCCTCGGCGAATTAC--ATTGCACTTGCGGGATGCATGTTTGA-GAA-CTCC-TTTGC-------TACCC--------AAT-TGTGCTGCCTT-GTTGGTT----CCTTCGAGAAC------AATGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGTGGGATCACMCG---------------------------------ACATCGGGCGGTAAATTCCGTCCAARGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGCGTA-TAGCCGGTCTGCCAATCGACTCAGGGCGTGGACCGACGCGGATTGTAGTGGTGGCCCAAGCCTGAGCCTTTGTTGTGCTTGTGGAGACGTCATCACCGCGATCGTGGCTGGCAGCGCGCGCCT-CTGGCGTGCTTCGGCATCAGCGTGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GTGGGT-TGCACCGCCGACCGACCTTGATCTTTTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGC-------------
		Begonia_floccifera                -----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------TGCCTGGGTGTCACGCATCG-CGGCCCCCC-------CCATCCATCCATCCATCCG--------------------------------------------------------------------CCAG-CCTCCCG-T-------------GCTGGGCCCT-----------GGGTGTTT----GGGGAGGCGACGTTGGCCTCCCGTGCGCGC----AGCC-TCGC--GGTTGGCCTAAATTTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATCGC--GTCGCGCCCGCGAGACGCGCGGCGGG-GAA-CTCC-TTCAC-------GACCC--------TCA-CGCGTCGACCC-CCC---------GGACGAC-TCGT-CCCGGCGGGAAGACGTCAT-----CGGAGC--GACCCCAGGTCAGGCGGGATTACCCG---------------------------------AAATCGGGCGGTAAATTCCGTCCAAG-CTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCTTTGGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGCTGTGCTCGCGGAGACGTCGTCGCCCCGATCGTGGCTGGCAGCGCGCGCCTCACGGCGTGCTTCGGCATCAGCGCGCTCCRGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGCGAGGCGCAAGGAARCTGACTGGCGGGATCCCCTB-G-GGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGA---------
		Begonia_engleri                   GATCATTGTCGATTCCTGC-AAATA---GCAG-AAAAACCCGCG-AATTGGTTAAA-----CATCG---TCCCTCAC-----TTGG----GGGAACGGCGCGGTGAGCACTAA---GGCCGAGGCA--CGT-GCAC---------GCACAAGCTCT--GCC--TGGC-GAGCACCGCGTG----------------CCCTAACAAATCCT--AGCGCGAGTCACGTCAAAGAA--TCTTGAA-CTT---AGAACGCGCA---TCATCT-GCCCT-GGTTTCAG-GATGC----AGGGG--ACC-TGCAAACTT-----GATCAAATATCAGAC-ACGACTCTCRGAAATGGATATCTCGGCTCTCATATCGATGAAGAATGTAGTGAAATGGGATACTTGGTGTGAATTGCAGAATCCCGTAAACCATCAAGTTTTTGAACGCAAGTTACGCCTTAAGCCTTCTGGCCKAGGGCACGTCTACCTGGGCATCACGAATTG-TAGCCCCCC--------CACCAAAAACCCT----TGC-------------------------------------------------------------------------------------------------GGGCTTT----------GGCTGGCC----GAGGTGGCGATGTTGGCCTCCCGTGAGGGG----AACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCAGCAT-CGCGCAACACGATCGATGGTGGTTGGAAT--GCCCTCGGTGGATCAC--GTTTCCCTCACGAGACGCGCCTACGAGGAA-CTCT-TTYGC-------AACCC-----------------------------------------------------------------------------------------------------TACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_edmondoi                  -CATTGT-CGATGCCTG-C-AGAAT---GCAG-AACAACCCGCG-AACTAGTCTG-----CCACCG---TCCCGCTT----CGCGC----GGGAA-AGGCGCGGG-GCGCGTTGG-TGCATCGGCC---GC-CTCT---------GCGCGCGCCCT--GCC--TCGC-GCGGCCCGCGCCCCC----------------TAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TATCAAA-TTC---AGAACGCGGT---TCCCCT-GCCCC-GATTTCGG-GATGC----AGGGG--TCC-TGCGAACTC-----GATCGTCA-TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-CAGCCCCCC-------GCACCCAACAAGAAAGA-------------------CCAA----GTCTC--------GATC--GAAGCG---GACTC---GTGCT-GCTTC--GTCGGGG-----CTC-----GGCC-GGCCCAT----------GGGCGGTC----GGGGTGGAGACATTGGCCTCCCGTGGGGGC----AACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTTCAAT--GCCCTCGACGAACTAC--GTTGCGCTCGCGAGGCGCGCGCTCGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGCCGGCTG-GTCGG---------CTAACGCC------GGCGAGCAGACGCTAT-----CGAAGC--GACCCCAGGTC-----------------------------------------------CAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCCCGGTCGGATGTGGAACGGCGTC-GAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGCGGCGGCGGCCCAAGCCTGGGCCTTTGTTGTGCTCGTGGAGACGTCGTCGCCGCGATCGTGGCTGGCAGCRCGCGCCC-CTGGCGTGCTTCGGCATCGGCGCGCTCCTGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAACTGACTGGCSGGATCCCCTT-G-GGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGCGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCG------
		Begonia_duncanthomasii            TGCGGAAGGATCATTGTCGATTCCT---ACAG-AACGACCCGCG-AACT-GGTTAT-CAACCACACGA-TTCCCCAGCTA-GCGCT---GGGGAA-AGGCGGGGC-ACGCTGGTC-C--CGCATTG--CGC-GCTCC--------GCGCG---------------------CCTCGCCG----------------ACCGCAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--CATAAAA-CTG---AGAGCGCGGA---TCCCCT-GCCCC-GGCTTCGG-GAAGC----AGGGA--TCT-CGCGCGCTT-----GCTCA----CGACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCAGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGTCCCCC-------TCACCGAAGAGGCCAA---------------------ATTC--GAGCACCCAGAGC---AGTGCAG------TC-----TCTGCAC-----TT-GCG-TCGGGC---------GCTT-GGATTTGCAGCTC----GTTTGGA-----GGGGCGTCAACGTTGGCCTCCCGCG-GAGC----AACC-TCGC--GGTTGGTCTAAAGTAGAGTGCTCGGCAT-CGCGCGGTGCGATCGACGGTGGTTTCGTA--GCCCTCGGCGAATTAC--GTCGCAC-----AGATGCG-GCGCGGAGAA-CTCC-ATACG-------GACCC--------TGC--TGATGATTGCC-GGCT-----CGCGGGTTCCCC-------GCGAGAAGGCACTGC-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_dregei_partita            TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGCG-AACTAGTTTG-----CCAAAG---TCCCGCCC----CGCGC----GGGAA-CGGCGCGGG-GCGCTTCGG-AGCCGTGGCT--CGT-GCTT---------GCGCGCGCCCT--GCC--TGGA-ACGCCTCGCGCCCC----------------TTAACGAACCCC--GACGCCAGTCGCGTCAAGGAA--TGTATAA-CTC---AGAACGCGGC---TCCCCT-GCCCC-GATTTCGG-GACGC----AGGGGTGTCT-TGCGAACTT-----GATAA----CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTYTGCCTGGGCGTCACGCATGG-TAGCCCCCC-------GCACCACAAAAAAAA------------CAAGGCCGAGATCG--TTCCTC----GGTC-ATC----------------------------------------------------GGCT-GGCCTTT----------GGGTGGTC----GGGGCGGCAATGTTGGCCTCCCGTGCGGGC----AACC-GCGC--GGATGGCCTAAATTTGAGTCCTAGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--GTTGCGCCTGCGAGACGCGCGCTTGG-GAA-CTCC-TCGAA-------GACCC--------TAA-TGCGTCGGCTC-GTCGG---------AAAAAGCC------GGCGAGTGGGCGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_dregei_ssp_homonyma       TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGCG-AAYTAGTTTG-----CCAAAG---TCCCGCCC----CGTGC----GGGAA-CGGCGCGGG-GCGCTTCGG-AGCCGTGGCT--CGT-GCTT---------GCGCGTGCCCT--GCC--TGGA-ACGCCTCGCGCCCC----------------TTAACGAACCCC--GACGCCAGTCGCGTCAAGGAA--TGTATAA-CTC---AGAACGCGGC---TCCCCT-GCCCC-GATTTCGG-GACGC----ATGGGTGTCT-TGCGAACTT-----GATAA----CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCYTTHTGGCGGAGGGCACGTYTGCCTGGGCGTCACGCATGG-TAGCCCCCC-------GCACCACAAAAAAA-------------CAAGGCCGAGATCG--TTCCTC----GATC-ATC----------------------------------------------------GGCT-GGCCTTT----------GGGTGGTC----GGGGCGGCAATGTTGGCCTCCCGTGCGGGC----AACC-ACGC--GGATGGCCTAAATTTGAGTCCTAGGCGT-CTTGCGGCGCGATCGACGGTGGTTGTCAT--GCCCTCGACGAATTAC--GTTGCGCCTGCGAGACGCGCGCTTGG-GAA-CTCC-TCGAA-------GACCC--------TAA-TGCGTCGGCTC-GTCGG---------AAAAAGCC------GGCGAGCGGGCGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-----------------------------------------------------------GCTAAATA-TTGGCGAGAGACCGATASCRRACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAKAGTCAAAKAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGKTKCGTCCYGGTCGGATGTGGAAYGKCKCGY-------------------------------------------------------------------------------------------------------------------------------------------ATCAGCGTGCTCCAGGCRTBGGSCTGYGGGCTCCCCATTCGGYCCGTCWTGAAACAYGGACCAAGGAGTCTGACATGTGTGCGAGTYAACGGGCCAGAAAAMYCGTGAGGCGCAAGGAAGCTGACTGGCGGGATSCCCTY-GMGGGT-TGCACCGCCGAC-GACCTTGATCKTCTGAGAAGGGWTCGAGTGWGAGCAYGCCTGTCGG-ACCCGAAAG---------------------------
		Begonia_dregei                    TCATTGT-CGATTCCTG-C-AAAAT---GCAG-AACAACCCGCG-AATTAGTTTG-----CCAAAG---TCCCGCCC----CGTGC----GGGAA-CGGCGCGGG-GCGCTTCGG-AGCCGTGGCT--CGT-GCTT---------GCGCGTGCCCT--GCC--TGGA-ACGCCTCGCGCCCC----------------TTAACGAACCCC--GACGCCAGTCGCGTCAAGGAA--TGTATAA-CTC---AGAACGCGGC---TCCCCT-GCCCC-GATTTCGG-GACGC----ATGGGT-GTCTTGCGAACTT-----GATAA----CATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATAG-TAGCCCCCC-------GCACCACAAAAAAA-------------CAAGGCCGAGATCG--TTCCTC----GATC-ATC----------------------------------------------------GGCT-GGCCTTT----------GGGTGGTC----GGGGCGGCAATGTTGGCCTCCCGTGCGGGC----AACC-ACGC--GGATGGCCTAAATTTGAGTCCTAGGCGT-CTTGCGGCGCGATCGACGGTGGTTGTCAT--GCCCTCGACGAATTAC--GTTGCGCCTGCGAGACGCGCGCTTGG-GAA-CTCC-TCGAA-------GACCC--------TAA-TGCGTCGGCTC-GTCGG---------AAAAAGCC------GGCGAGCGGGCGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_dipetala                  TCATTGT-CGATTCTTG-C-AAAAT---GCAG-AACAACCCGCG-AACTAGTTTTG----CCAACT---TCCCGCCGCCG-CGCGCGCGCGGGAA-CGGCAGGGG-GCGCGTCGG-AGCTGTGGTCCGCGCTGCTTTGCTT----GCGCGCGCCCT--GCC--TCGCG-CGCCTTGTGCCCCC---------------TTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TCTGGAA-CTT---AGAACGTGGC---TCCCCT-GCCCC-GGTTTCGG-GACGC----GGGGG--TCC-TGCGGACTTTT---GATCATAT-AATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------TCACCCAAAAAACACAACC---------CAAAGAT--CAA---TCCCTC--------GAACGAAAAGCA---GACTC---GTGCT-GCCTC--GTTTGGG----GTTTTGTGTTGCTGGGCCTTT----------GGGTGGTTT---GGGGAGGCGACGTTGGCCTCCCGTGCGGGC----AACC-TCGC--CGTTGGTCTAAATTTGAGTCCTCGGCGTCTTTGCGGGGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--GTCGCGCCTGCGGGACGCGCGCTCGG-GAG-CTCC-TCGAC-------GACCC--------TCA-CGCGTCTATTC-GTC---------GGACATTATAT--GCCGGCGGGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGYGGGATTACCCG----------------------------CACCCAAAYTGGGCGGTAAATTCCATCCAAGGCTAAATA-TTGGCGAGAGRCCKATAGCAAACAAGTTYCGCGAGAGAAAGATGAAAAGGACTTTGAGAAATGAGTCAAAGAGTCACTRGAATTTTCGKAGAGAGAAGCGTGATTGGGGCTCGA-----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_dewildei                  GATCATTGTCGATTCCTGCGAAAAAAGAGCAG-AAAAACCCGCG-AACT-GGTTTT-TAAGCACGCGC-TTTCCCAGCCC-GCGGT---GGGGAA-AGGCGGGGC-GTGCTGCG--A--CGAGTTC--CGC-GCCCA--------GCGCGCGGCCG--TTC--GAGC-GCGCCTCGCGC----------------ACCGTCACGAACCCC--GGCGCGAGTCGCGTCAAGGAAA-GATGTAA-CCG---AGAGCGCGTA---TCCCCT-GCCCC-GGACCCGG-GAAGC----AGGGG--TGT-AGCGCGCTT-----GCTAA----CGACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGAAGCGCCCGAAGCCTTCTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGGCCCCC-------GCACCGAAGAGGCCAA---------------------GTTC--CAGCTCC-AGAGGGGGCGGGCG------GTTTG---TCCGC-------TT-GCG-TCGTCG---------GCTG-GGATTTGCGTCTC----GTTCGGA-----GAGGCGGCAAGGTTGGCCTCCCGCGCGAGC----AAGA-GCGC--GGTTGGTCTAAAGTAGAGTGATCGGCAT-CGCGCGGTGCGATCGACGGTGGTTTCCAT--TCCCTCGGCGAATCAC--GTCGCACGCGCGGGATGCG-GCGCGGAGAACACTCCATGGG-------CACCC--------TGC--TGATGA-CGCC-TTCC-----CGCGGGTTCCCC-------CCGWGGAGGCGCTGC-----CGAAGC--GACCCCAGGTCAGGYGGG----------------------------------------CAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTCGGTCGGATGTGGAACGGCGG--GAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATCGCGGCGGCGGCCCAAGCCCGAGCCTTGGTTGAGCTCGCGGAGACGTCGTCGCGGCGATCGTGGGCGGCAGCGCGCGCCA-GCGGCGTGCTCAGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGCGTGCGAGTCAGCGGGCCAGCAAACCCGCGAGGCGCAAGGAAGCTGARTGGCGGGATCCCCRA-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGCGAGC------------------------------------------------
		Begonia_crassirostris             ----------------------------GCA--AWCAACCCGCG-AA-TAGTTT-G----CCAACG---TCCCRCCC----CGTGC----GGGAA-CAGYGCGGA-GCACGTCAG-AGTTGTGGAY--AGC-GCTT---------TCGCGCCCCYT--GCC--CTGCA-TGCCCYGCGC--TTC---------------TAACGAACCCC--GGCGCAAGTCGCGTCAAGGAA--TCTGAAA-CTT---AGAACGCGGC---TCCCCT-GCCCC-GATTTYGG-AACGC----AGGGG--TTC-TGCGAACTT-----GATCA----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCCAAACAAAAAAAA----------CCA------------TCCATC--------TATCC-GAAGCT---GACTC---GTGTC-GACTC--GTT-GGG-----GTTTTGGTGGCT-GGCCTTT----------GGGCAGTC----GGGGAGGCGACGTTGGCCTCCCGTACGGGC----AACC-TTGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTCGCAGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--ATTGTGCCTGCGAGGTGCGTCGTGGG-GAA-CTCC-TTCAC-------GACCC--------TAA-TGCGTTGACTC-GTC---------GGACGACAAC------GGCGAGAAGATGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG------------------------------CCCAAATCGGGCGGTSCATTTCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGTGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGATCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCT-ACGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTTAGGCGCAAGGAAGCTGACTGGCGGGATCCCGTG-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_convolvulacea             ---TTGT-CGATTCCTG-C-MRAAT---GCAG-AACAACCCGCG-AACTAGTTTTG----CCAATG---TCTCGTCT----CGCAC----GTGA--CAGCGCGGG-GCGTGTTGG-TGCTGTGGCT--TAC-CTAY---------GTGTGAGCTCT--GCC--TGGC-ATGCCTCGTGCCCCC----------------TAACAAACCCC--GGCGCGAGTAGCGTCAAGGAA--TATCAAA-CTT---AGAACGCGCC---TCCCTT-TTCCC-TATTTTGG-GATTG----AGGGT--CGT-TGCGTACTT-----GATCAAA--TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCAAAGCCTTTTGGCCGAGGGCACGCCTGCCTGGGCGTCACGCATCG-TAGCCCCTCC------ACACAAGTCATAG---------------AGAGGTGGTCAA---GGTCGC--------GATG--AAAGCA---GACTT---GCGCT-GCAAC--GTCGGGA-----CTT-----GGCTTGCTCTT-----------GGGTGATG----GTGGTGGCGACATTGGCTTCCTGTGGGTTC----AATC-TCGC--GGTTGGTCTAAATTTGAGTCTTTGGCGTTCTTGCAGCGCGATCGACGGTGGTTTCCAT--GCCCTCGACGAATCAC--ATTGCGCTTGCGAGTTGCATGCTAGT-GGA-CTGC-TCAAC-------GACCCTCA-----TAA---CGTTA--------------------------------------TGAAACGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-------------------------------CCA-ATCGGGTGGTAWATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCTAG-TAGCCGGTCTGCCGATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGATGGCAGCGCGCGCCA-ACGGCGTGCTTCGGCATCAGCGTGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGT-TGCACCGCCGACCGACCTAGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_cinnabarina               ---TTGT-CGATGCCTG-C-M-ART---GCAG-AACAACCCGCG-AACTAGTTTG-----CCAACT---TCCCGCTT----CGCGC----GGGAA-CGGCGCGGG-GCGCGTCGG-ARCTGTGGCA--CGC-TGTT---------TTGCGCGCCCT--GCC--TCGC-ACGCCTCGTGCCGCC----------------TAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TCTAAAA-CAT---AGAACGCGGC---TCCCTT-GCCCC-GATTTCGG-GATGC----AGGGG--TACCTGCGAACTT-----GATCA----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTTTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCAAAAAAGAAAAAA----------CCAAGA--TCAA---TCCCTC--------GATC--CAAGTA---GACTC---GTGCT-GCTTT--GTTGGGG-----GTT-----GGCT-GGCCTTT----------GGGTGATC----GGGGCGGCGACATTGGCCTCCCGTGGGGGC----AACC-TCGC--GGTTGGTCTAAATTCGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCAAT--GCCCTCGACGAATTAC--GTTGCGCATGCAAGATGCACGCTCGG-GAA-CTCC-TCGAC-------GACCC--------TAA-TGCGTCGGCCA-GTCGG---------ACAACACC------GGCTTGCTAACGTTAT-----CGAAGC--GACCCCAGGTCA---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_capillipes                --TT-GT-CGATTCCTG-C-MMAWA---GCAG-AACAACCCGCG-AATCCGTTGTG-----CAACGA--ACCC--------CGCAA---GGGGAG-CGGCGAGGA--GTGTTGGGG------------------------------------------------CGC-AAGCCCCGCATTT-------CGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAA-CTG---AGAACGCGGC---TCCTCT-GCCCC-GCCTGCGG-GATGC----AGGGGG-CGC-TGCGAGCTT-----GAGAA----TCACAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCGCCAAAAAAA------TGC----------------TCGA--TCTC-TTC----------------------------------------TT-GGGGTGG------------GGCT-GGACTTT----------GGTGGGAC----GAGGGGGCGATGTTGGCCTCCCGTGCGAGC----GATC-GCGC--GGTTGGCCTAAAGTCGAGTCCTCGGCGT-CGCGCAGTGCGACCGGCGGTGGTTTTTA---GCCCTCGGCGAAACAC--GTTGCACGTGCGAGACGCGGTATCGG-GAA-CTCC-CACGC-------GACCC--------TGC-TGCGCTGCCCG-GTC-GTT-----CTCC--GAAC------GACGGGGAGGCGCTAT-----CGAAGC--GACCCCAG-TCAGGYGGGAT-ACC-----CTTAGATCGGGTTGTTTGGGAATGCAGCCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGTG-C-AAGCCGGTCTGCCAATCGACTCAGGGCGCGGACCGACGCGGATTGCGGCGGCGGCCCAAGCCTGAGCCTTTGTTGTGCTTGCGGAGACGTCGTCGGCGCGATCGTGGCTGGCAGCGCGCGCCATTAGGCGTGCTTCGGCATCGGCGCGCTCCGGGCGTCGGCCTGCGGGTTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTA-GCGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGG-TTCGAGTGAGAGCATGCCTGTCGGGACC-GAAAGATGGTGAACTATGCCTGAGCG------
		Begonia_bogneri                   TCATTGT-CGATTCCTG-C-AGAAA---GCAG-AACAACCCGCG-AACTAGTTAA-----CCATCG---TCCC--------CATGT----GGGAA-TTGTGAGGC--GTGTTGG--AGCTGCGACACAA---GCTT---------GCGCGTGCCCT--GCC--TTGC-ATGCCTCGCAA----------------ACCATAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TCTTTAA-CTG---AGAACGCCGC---TCTCCTACCCC--GACTTCGG-GATGY----AGGGG--TGC-TGCGAACTT-----GCTTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACACATCG-TAGC-------------CACCCACACCAAAGCTT-SCAAGCT-------------------------------------------------------------------------------------------------------------------AT----GAGGTGGTAATGTTGACCTCCCGTGAGAGC----AAGT-GCAC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-TGCGCAGTGCGATCGGCGGTGGTTGCCAT--GCCCTCGGCGAATTAC--ATTGCACTTGCGGGACGCATGTTCGG-GAA-CTCC-TTTGC-------GACCC--------TAT-TGTGCTGCCTC-GTTGGTT----CCTYCGAGAAC------CATGAGAAGACGTCAT-----CGAAGC--GACCCCAGGTCAGGYGGGATCACC---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_balansana                 GATCATTGTCGATTCCTGC-AAAAT---GCAG-AACAACCCGCG-AACTAGTTT-G----CCAACG---TCCCGCCT----CGTGC----GGGAA-CGGCGCGGA-GCGTGTCGG-AGCTGTGGAC--AAC-GCTT---------TCGCGCGGCCT--GCC--TCGCA-TGCCCCGCGCCCCT----------------TAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TCTGAAA-CTT---AGAACGCGGCT-CTCCCCT-GCCCC-GTTTTCGG-GATGC----AGGGG--TCC-TGCGAACTT-----GATCA----TATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------GCACCCAAACAAAAAAA-----------GCAACAC--CGA---TCCATC--------GATC--GTAGCA---GACTG---GTGCT-GCTTC--GTTTGGG-----GTTTTGGTTGCT-GTCCTTT----------GGGCGGTC----GGGGAGGCGACGTTGGCCTCCCGTATGGGC----AACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CTTGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--ATTGYGCCTGCGAGGCGKGCCGTGGG-GAA-CTCC-TCGAC-------GACCC--------TAA-YGCGTCGACTS-GTC---------GGACAA-ACC------GGCGAGAA-A-GYYAT-----CGAAGC--GA--------------------------------------------------------CAAATCGRGCG-AWA--TYCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCC-TAGCCGGTCTGCCAATCGACTCGGGGCGTGGACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGGGCCTTTGTTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCT-ACGGCGTGCTTCGGCATCAGCGCGCTCCGGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTTAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTG-GAGGGT-TGCACCGCCGACCGACCTTGATCTTCTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_aspleniifolia             GATCATTGCCGATTCCTGC-AAAAA---GCAG-AACGACCCGCG-AACTCGTTGGA----CCC------TCTTCCCCACT---------GGGGAG-AGGCGGCGG-TTCCGCRGA-A-----------CGC-GCCCT--------GCGCGAGCCCG--GCG--CCGC-ACGCCTCGCGC----------------ACCCTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--TATGTAA-CTG---AGAGCGCGGT---TCCCCC-GCCCC-GGCCCCGG-GAAGC-----GGGG--GTCTCGCGTACTT-----GCTCG----ACACGC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCYTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGGCCCCC-------GCAGCAAAGAGGCCAAGCCCGAGAT-----------------------CCCGAAGCATGCAGGCG-------TC-----TCTGT-------TT-GCG-TCGGGGA--------GCGG--GC-TGGCATCCG----GTTTGGC-----GAGGCGGMMAGGTTGGCCTCCCGCGCMAGC----AGCG-GCGC--GGTTGGTCTAAAGTAGAGTACTCGGCAT-CGCGCGGTGCGATCGGCGGTGGTTCASTC--ACCCTCGGCGGATCGC--GRTGCACTCGCGGGATGCG-ACGCGGAGAA-CTCC-ATGCG-------GACCC--------TGC-TGATGCCTGCT-----------CGCGGAGATCT--------GCGAGATGGCGCTGC-----CGAAGC--GACCCCAGGTCAGGCGRGATTACCCG---------------------------CAGCYCAAATCGGGCGGTAAATTCCGTCYAAGGCTAAATA-YTGGCGAGAGACCGATAGCAAACAAGTACCGYGAGGGAAAGATGAAAAGSACTTTGARAARAGAGTCAAASAGTACTTGAAATTGTCGGGAGGGAAGCGVATGGGGKCCGRCGKKKCKCCTCGGTCSSATSTGKAWCGGCRH--GAGYCGGTCCGCCARTCGACKCG-GSCKYGGWCCGMYGSGSATTGYRRCGGSGGCCCAARCCYRASCCTYCGYYGADCTYGYKRARAC-TCKTCGC-----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
		Begonia_annobonensis              GATCATTGTCGATTCCTGCAAAGA----GCAG-AACAACCCGGG-AACTTATTAAAA----CATCGT--CCCC--------TACAA--GGGGGAA-TGGTGCGGT-GCGTGCCA--A-----GGCT---GT-G-TT---------GTGTGTACGC---------GCCCAGCTTTGGCAAGC-------CCCGCGCGCCCTAACGAACCCC--GGCGCGAGTCGCGTCAAGGAA--ACTTGAA-CTT---AGAACGCATG---TCCTTTTGCCCC-GGCTTCGG-GATGC----AGGGGG--CCTTGTGAACTT-----GATGAAATATCAAAC-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCCCCC-------CCACCAAAAACCCCT---TGCGG------------------------------------------------------------------------------------------------GATTTGGAT-------GGTC--------GAGGTGGCGACATTGATCTCCCGTGTGG--------CY-TCGC--GGTTGGTCTAAATTAGATTCCTCAGCAT-CGCACCACGCGATGGACGGTGGTTGCAAT--GCCCTTGGTGGATCAC--ATTGCGCTTGTGAGATGCGGATACGAGGAA-ATCC-TTTGC-------GACCC--------TAT-TGTGTCAATTC-GTTGCTAA--CCGTTAGGTGCA------GCGGCYTCGACATCAT-----CGAAGC--GACCCCAGGTC------------------------------------------------AAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGYGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTCGGTCGGATGTGGAACGGCGTT-GAGCCGGTCTGCCAATTGACTCGGGGCGTGGACCGACGCGGATTGTGGTGGCGGCCCAAGCCTGGGCCTTTGTTGTGCTCGTGGAGACGTCGTCGCCGCAATTGTGGCTGGCAGCGCGCGCCC-TAGGCGTGCTTCGGCATCAGCGCGCTCCTGGCGTCGGCCTGTGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCTTT-GGGGGT-TGCACCGCCGACCGACCTTGATCTTYTGAGAAGGGTTCGAGTGTGAGCATGCCTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGG-----
		Begonia_ankaranensis              TCATTTT-CGATTCCTG-C-AAAAA---GCAG-AACAACCCGTG-AACTAGTTAA-----CCATCG---TCCC--------CATGC----GGGAA-TTGCGAGGC--ATGTTCG--AGCTACGACACA----GCTT---------TTGCGTGCCTT--GCC--TTGC-ATGCCTCGCGA----------------ACCATAACGAACCCC--GGCGCGAGTTGCGTCAAGGAC--TCTTTAA-CTG---AGAACGCCGC---TCTCCTAGCCC--GACTTTCG-GATGC----AGGAG--TGC-TGCGAACTT-----GCTTAT---CATGA--ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCKGCCTGGGCGTCACGCATCG-TAGC-------------CACCCACACCAAAGCTTTGCACACGCTCAATCTCCCGGTTGAGATAGACCTATGCTATMTTTCTC----------------------------GGGTTTG------------GGCT-GGCAAT----------GGGTTCGAT----GAGGTGGTAGTGTTGGCCTCCCGTGCATGG----AAGT-GGAC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-TGCGCAATGCGATCGACGGTGGTTGCCAT--GCCCTCGGCGAATTAC--ATTGCATTTGCGGGATGCATGTGCGA-GAA-CTCC-TTTGC-------GACCC--------AAT-TGTGCTGCCTC-GTTGGTT----CCTTCGAGAAC------AATGAGAAGACGTTAT-----CGAAGC--GACCCCAGGTCAGGMGGGATCACCCG----------------------------CAGCCAAATCGGGCGGTAAATTCCGTCCAAGGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGCCTTGGTCGGATGTGGAACGGCGTC-TAGCCGGTCTGCCAATCGACTCAGGGCGTGGACCGACGCGGATTGTAGTGGTGGCCCAAGCCTGAGCCTTCGTTGTGCTTGTGGAGACGTCATCACCGCGATCGTGGCTGGCAGCGCGCGCCT-CTGGCGTGCTTCGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGTAAACCCGTGAGGCGCAAGGAAGCTGACTGG--------------------------------------------------------------------------------------------------------------------
		Begonia_aequata                   TCATTGT-CGATTCCTG-C-ARAAT---GCAG-AACAACCTGCG-AACAAGTTTTG----CCAATG---TCTCGCTG----CAYGC----GGGAA-TGGCGCGGG-GCGCGTCAG-AGTTGTAGCC--TGC-GTTT--------GCGCGRAGCCCA--GCC--TCGCG-CGCCTTGTGCCCCC----------------TAACGAACCCC--GGCGCAATTTGCGTCAAGGAA--TCTAAAA-CTT---AGAACKTTGA---TGCCCT-GCCCCAAATTTTGG-GATGC----AGGGG--TCC-TCTGATCTT-----GATCACATATATCAA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCTGAGGGCACGTCTGCCTGGGCGTCACGCATCG-TAGCCTCTC-------GCACCCAAAAAAAAA-------------CCAAGTA--CAA---TTCCTC--------GATC--GAAGTA---GACTC---GTGCT-GCTTT--GTTGTC------GTTTT----GTT-GGCCTTT----------GGGTTGTC----GGGGAGGCGGTGTTGGCCTCCCGTGCGGGT----AACC-TCGC--GGTTGGTCTAAATTTGAGTCCTCGGCGT-CCAGCGGCGCGATCGACGGTGGTTGCCAT--GCCCTCGACGAATTAC--TTTGCGCTTGCGAGACGCGTGCTTGA-GTA-CTCC-TCATC-------GACCC--------TAAATGCGTCGACTC-GTT---------GGACAACACC------AGCGAGAAGGCGTTAT-----CGAAGC--GACCCCAGGTCAGGCGGGATTACCCG-----------------------------------------------ATTCCGTCCAASGCTAAATA-TTGGCGAGAGACCGATAGCAAACAAGTACCGCGAGGGAAAGATGAAAAGGACTTTGAGAAAAGAGTCAAAGAGTACTTGAAATTGTCGGGAGGGAAGCGGATGGGGGCCGGCGGTGCGTCCCGGTCGGATGTGGAACGGCGCG-TAGCCGGTCTGCCAATCGACTCGGGGCGTGAACCGACGCGGATTGGGGCGGCGTCGCAAGCCCGAGCCTTTGCTGTGCTCGCGGAGACGTCGTCGCCTCGATCGTGGCTGGCAGCGCGCGCCC-ATGGCGTGCTTCGGCATCAGCGCGCTCCAGGCGTCGGCCTGCGGGCTCCCCATTCGGCCCGTCTTGAAACACGGACCAAGGAGTCTGACATGTGTGCGAGTCAACGGGCCAGAAAACCCGTGAGGCGCAAGGAAGCTGACTGGCGGGATCCCCTT-GAGGGTTTGCACCGCCGACCGACCTTGATCTTTTGAGAAGGGTTCGAGTGTGAGCATGCTTGTCGGGACCCGAAAGATGGTGAACTATGCCTGAGCGGAAG--
	;
END;

BEGIN CODONS;
CODESET * UNTITLED = Universal: all ; 
 END;

BEGIN ASSUMPTIONS;
CHARSET ITS = 1 - 946 ; 
 CHARSET 26S = 947 - 1541 ; 
 CHARSET exclusion = 1 - 28 49 - 194 529 - 651 869 - 891 934 - 990 1531 - 1541 ; 
 CHARSET exclusion2 = 243 - 349 505 - 516 525 - 663 690 - 708 840 - 868 ; 
 OPTIONS DEFTYPE = unord PolyTcount = MINSTEPS ; 
 EXSET * excluded = 1 - 29 49 - 194 529 - 651 869 - 891 934 - 990 1531 - 1541 ; 
 END;