NEXUS
[File created by TreeBASE:  7/26/07  21:10:03]
[Matrix accession#: M2528]

BEGIN TAXA;
	TITLE M2528;
	DIMENSIONS  NTAX=27;
	TAXLABELS
		Begonia_baccata	Begonia_ampla	Begonia_poculifera	Begonia_bonushenricus	Begonia_loranthoides	Begonia_longipetiolata	Begonia_subscutata	Begonia_kisuluana	Begonia_komoensis	Begonia_oxyanthera	Begonia_horticola	Begonia_eminii	Begonia_capillipes	Begonia_mannii	Begonia_fusialata	Begonia_preusii	Begonia_polygonoides	Begonia_gabonensis	Begonia_ebolowensis	Begonia_molleri	Begonia_furfuracea	Begonia_subalpestris	Begonia_cavallyensis	Begonia_bogneri	Begonia_betsimisaraka	Begonia_squamulosa	Begonia_eleagnifolia
	;
END;

BEGIN CHARACTERS;
	TITLE M2528;
	DIMENSIONS NCHAR=1230;
	FORMAT DATATYPE = DNA MISSING = ? GAP = -  ;
	MATRIX
		Begonia_baccata              AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCC------------------TCCAAC---------------------------------------------------------------------------------??------------------------------?---------------------------------------------?----CTTAAAGTTGAAAAAAGAATTAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACTTGTCGTGCAACGAACCCCGCAAGGGAA-TGGCGGGGGCGTGCTGGGGCG----------------CTCGCGC-GC-TCCTCGCGC---TCCCCGCGCACCTCAACGAACCCCC-GGCGCGAGTCGCGCCAAGGAA--TTTTTAAC-TG-AGAATGCGGCTCCTCTGCCCTCGATTT-CGG-GATGC----AGGGG-CGCYGCGAGCTTGAAAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTGGCCCCCCACGCTCGCGCCTATAAAAA---C-CGAGTTC-AATCTCCTG--TT-----------------GGGGGTGGGCCGGACTTT-GGTT--GGACGAGGGGGGCAGTGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTTGAGTCCTCGGCGTCGCGCAGTGCGATCGACGGTGGTTTTT---TAGCCCTCGGCGAGACACATTGCACGTGCGAGACGCGGTAT-CGGGAG--CTCCCAC---GCGACC?TGCTGCGCTGAAACACGA-A-AAAAAAAAAKAAACAA
		Begonia_ampla                AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAATAAATAATAGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCC------------------TCCAAC---------------------------------------------------------------------------------??------------------------------?---------------------------------------------?----CTTAAAGTTGAAAAAAGAATTCAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCTGCGAACTTGTCGTGCAACGAACCCCGCAAGGGAA-TGGCGGGGGCGTGCTGGGGCG----------------CTAGCGC-GC-TCCTCGCGTA--TCCC-GCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGCCAAGGAA--TCTTTAACTTG-AGAATGCGGCTCCTCTGCCCCTG-GTTGCGG-GATGT----AGGGG-CGCTGCGAGCTTGAAAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTTTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTGGCCCCCCACGCTCCTGCCAAAAAA-----T-CGAGTTC-AATCTCCTA--CT-----------------GGGGTTGGGCCGGACTTT-GGTCGGGGACGAGGGGGGCAATGTTGGCCTCCCGTGCGATCGGTCGCGCGGTTGGCCTAAAGTTGAGCCCTCGGCGTCGCGCAGTGCGATCGACGGTGGTTTT----TAGCCCTCGGCGAGATACATTGCACGTGCGAGACGCGGTGT-CGGGAG--CTCCCAC---GCGACCCTGCTGCGCTGAAACACCA-A-CAAAAAACAGAACMAA
		Begonia_poculifera           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATAGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCC------------------TCCAAC---------------------------------------------------------------------------------??------------------------------?---------------------------------------------?----CTTAAAGTTGAAAAAAGAATTCAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACTTGTCGTGCAACGAACCCCGCAAGGGAA-TGGCGGGGGCGTGCTGGGGCG----------------CTAGCGC-GC-TCCTCGCGC---TCCCCGCGCACCTTAACGAACCCCC-GGCGCGAGTTGCGCCAAGGAA--TCTTTAACTTG-AGAATGTGGCTCCTCTGCCCCCG-GTTGCGG-GATGC----AGGGG-CGCTGCGAGCTTGAAAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTGGCCCCCCACGCTCCCGCCAAAAAA-----T-CGAGTTC-AATCTCCTA--CT-----------------GGGGTTTGGCCGGACTTT-GGTTGGGGACGAGGGGG-CAATGTTGGCCTCCCGTGCGAGCGATTGCGCGGTTGGCCTAAAGTTGAGCCCTCGGCGTCGCGCAGTGCGATCGACGGTGGTTTT----TAGCCCTCGGCGAGATACATTGCACGTGCGAGATGCGGTAT-CGGGAG--CTCCCAC---GTGACCCTGCTGCGCTGAAACACCA-A-CAMAAAACAKAACAAA
		Begonia_bonushenricus        AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATAGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCC------------------TCCAAC---------------------------------------------------------------------------------??------------------------------?---------------------------------------------?----CTTAAAGTTGAAAAAAGAATTCAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCTGCGAACTTGTCGTGCAACGAACCCCGCAAGGGAA-TGGCGGGGGCGTGCTGGGGCG----------------CTAGCGC-GC-TCCTCGCGC---TCCCCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGCCAAGGAA--TCTTTAACTTG-AGAATGYGGCTCCTCTGCCCCCG-GTTGCGG-GATGC----AGGGG-CGCTGCGAGCTTGAAAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCCCCCGAAGCCTTCTGG?GGAGGGCACGTCTGCCTGGGCGTCACGCATCGTGGCCCCCCACGCTC?AGC?AAAAAA-----T-CGA?TTC-AATCTCCTA--TT-----------------GGGGTTGGCCCGGACTTT-GGTCGGGGA??AGGGGG-CAATGTTGGCCTCCCGTGCGAGCGAT?GCGCGGTTGGCCTAAAGTTGAGCCCTCGGCGTCGCGCAGTGCGATCGACGGTGGTTTT----TAGCCCTCGGCGAGATACATTGCACGTGCGAGATGCGGTAT-CGGGAG--CTCCCAC---G?GACCC?GCTGCGCTGAAACCCCA-A-CAAAAAACAGAACMAA
		Begonia_loranthoides         ???TGATAACTTTCAAATTCAGAGAA?CCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---TATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAA-----------------------------?----CTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACTTGTCGTGCAACGAACCCCGCAAGGGAA-TGGCGGGGGCTTGATGGGGCG----------------CTCGCGC-GC-TCCTTGCGC---TCCCCGCGCACTTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TCTTTAAC-TG-AGAACGCGGCTCCCCTGCCCCCG-GTTGCGG-GATGC----AGGGG-TGTTGTGAGCTTGAAAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCAYGTCTGCCTGGGCGTCACGCATCGTGGCCCCCC--GC----GCCAAAAAAAAAAAC-TGAGTTC-AATCTCTTC--TT-----------------GGGGTTGGGCCAGACTTT-GGTC--GGACGAGGGGG-CAATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTTGAGTCCTCGGCGTCGCGCAGTGCGATCGACGGTGGTTTT----TTGCCCTCGGCGAGATACATTGCACGTGCGAGACGCGGTAT-CGAGAGCGCTCCCAC---GCGACCCTGCTGCGTTGAAACAA-C?CAACAACAAACSAAAAGC
		Begonia_longipetiolata       AAGTGATAACTTTCAAATTCAGAGAAACCCTGGCAATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAAGAAATAATGGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCTTTCCAGCGAAACTCCGGAAAG---GATGAAAAATAAACCTATATACATACGTATACGGACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAAT-TTTTTTTC-AATGATATAGTGAAAATGAACGAATTGTTGTAAATCAATT--CCC--AACTTCAAGTTGAAAAAAGAATAAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-TTGAAATTTATAGTGCAGAACAACCCGCGAACTCGTAGTGCAACGAACCCCGCAAGGGGA-TGGCGAGG-CGTGATGGGGCG----------------CTCGCCC-GC-TCCKCACGC---TCCCCGCCCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAWGGAARATCTTTAA-TTG-AGAACACGGCTCCTCTGCWCCCG-GCTGCGG-GATGC----AGGGG-CGTTGYGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATCTCSGCTCTCGCATCGATGAAGAACRTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACRCAAGTTGCGCCCGAAGCCTTCATGCGGAGGGCACGTCTKCCTGGGCGTCAYGCATCRTGGCCCCCC--GC----GCCAACAAAAAAAACCCGAG-TCCAATCCYTTA--TT---------ATTATTTTGGGGTTGGACCGGACTTTTGGTT--GGACGAGGGGG-CAATGTTGGCTTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTTGAGTCBTCGGCGTCGCGCGGTGCGATCGACGGTGGCCT-GT-ATAGCCYTCGGCGAGATACATTGCACGTGCGAGACGCGGTAT-CGGGAG--CTCCCAC---GCGACC-TACTGCGCTGCAMCAA-CWCAAMAAACCACNAAAGAC
		Begonia_subscutata           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTGGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAGAAGGATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATAAAATGATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTTCAGAACAACCCGCGAACCCGTCGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGTTGGGGCG----------------CACGCCCCGCAT-----------TCCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGCATGCAGGGG-CGCTGCGAGCTTGAGAATCAAACACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAA-----TGC----TC-GATCTCTTC--TT-----------------GGGGTCGGGCTGGACTTT-GGTC--GGACGAGGGGG-CGATGTTGG-CTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCYTCGGCGTCGCGCAGTGCGACCG-CGGTGGTTTTT--ATAGCCCTCGGCGAAATACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCAC---GCGACCCTGCTGCGCTGAMACCCCCTCAACCMCAAACVGGAGAC
		Begonia_kisuluana            AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATT-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTTGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAAT?C----------GACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTKCAGAACAACCCGCGAACCYGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAG-A-GTGTTGGGGCG----------------CATGCCCCGCAT-----------TTCGCGCACACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGC----AGGGG-TGCTGCGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAA-----TGC----TC-GATCTCTTC--TT-----------------GGGGTTGGGCTGGACTTT-GGTC--GGACGAGGGGG-CGATGTTGGCCTCCCGTGAGAGCGATCGCGCGGTTGGCCTAAAGTCGAKTCYTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGCGACGCGGTAT-CGGGAA--CTCCCAC---GCGACCCTGCTGCGCTGAAMCAA-CTCAACAACAAACSGGAGAC
		Begonia_komoensis            AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAAAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGGAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCAAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATTTTTTTTTATAATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCCAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTT?TAGTGCAGAACAACCCGCGAACTCGTTGTGCAACGAACCCCGCAAGGGGAGTGGCGAGGA-GTGTTGGGGCG----------------CACGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TATATAA-TTG-AGAACGCGGCTCCTCTGCCCC-G-CTTGCGG-GATGC----AGGGG-CG?TGCGAGCTTGCGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCAGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--G?----GCCAAAAAA-----TGC----TC-GATCTCTTC--TT-----------------GGGGTTGGGTTGGACTTT-GGTC--GGACGAGGGGG-CGATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCGC---GCGACCCTGCTGCGCTGAAACAA-CGCAACACCAAACSGGAGAC
		Begonia_oxyanthera           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAAAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATTTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACC-GCGAACCCGTTGTGCAACGAACCCCGCAAGGGGAGTGGCGAGGA-GTGTTGGGGCG----------------CACGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TATATAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGC----AGGGG-CGCTGCGAGCTTGCGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAAA----TGC----TC-GATCTCTTC--TT-----------------GGGGTTGGGTTGGACTTT-GGTC--GGACGAGGGGG-CGATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCGC---GCGACCCTGCTGCGCTGAAACAA-CTCAACACCAAACCGGAGAC
		Begonia_horticola            AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAAAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGGAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATTTTTTTTTATAATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTGTACATGTCAATACGGACAAAAA-T--AAATTTATAGTGCASAACAACTYGCGAACCCGTTGTGCAACGAACCCCGCAAGGGGATCGGCAAGRA-TTKTTGGGGCS----------------CAASCCCCSCWT-----------TYCSCSCSCACCTTAACAAACCCCCSGGCGMGWCTCGKSTCARAGAA--TATTTAWC-TGAAGAAYGCGGCTCCTCTGCCCC-G-CCTGCGGAGATGC----AGGGG-CGCTGCGAKCTTGAGAATCA--CACAMGACTCTCGACAACRGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCWWGTTGCGCCCGAAGCCTTYTGGCRGAGGGCAHGTCTGCCTGGGCGTCACRCATYGTAGCCCCCC--GC----GCCAAAAAA-----TGC----TC-GATCTCTTC--TT-----------------GGGGTCGGGGAGGAMTTT-GGTY--GGAMGAGGGGG-CGAWGTTGGCCTTCCGTGCGAGVGATTGSGCGGTTGGCCTAAAGTTGAGTTCTTGGSGTTGSGCAGTGSGACCGAMGGTGGTTTTT--ATRSCCTTSGGSGAAAWACGTTGAACGSGSGAGASGCGGTWT-SGGGAA--TTCCCAC---GCGACCCTGCTGCGCTGAAACAA-CYCAACACCAAACGGGAGAC
		Begonia_eminii               AAGTGATMACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGAAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATTAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATCACGAGTCCCATTCTACATGTCAATACGGACAAAAA-T-GAA-TTTATCC?GCAGAACAACCCGCGAA-TCGTTGTGCAACGAACCCCGCAAGGGGAGTGGCGAGGA-GTGTTGGGGCG----------------CACGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TATATAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGC----AGGGG-CGTTGCGAGCTTGCGAATTA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCATAGCCCCCC--GC----TCCAAAAAA-----TGC----TC-GATCTCTTC--TT-----------------GGGGTTGGGTTGGACTTT-GGTC--GGACGAGGGGG-TGATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGATGCGGTAT-CGGGAA--CTCCC--TTTGCGACCCTGCTGCGCTGAAACAA-CKCAACAACCAACCAAAGAC
		Begonia_capillipes           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTTGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAAT?C----------GACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAA--T-GAAATTTATAGTGCAGAACAACCCGCGAATCCGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGTTGGGGCG----------------CAAGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGC----AGGGGGCGCTGCGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAAA----TGC----TC-GATCTCTTC--TT-----------------GGGGTGGGGCTGGACTTT-GGT---GGACGAGGGGG-CGATGTTGGCYTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCYTCGGCGTCGCGCAGTGCGACCGGCGGTGGTTTT----TAGCCCTCGGCGAAACACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCAC---GCGACCCTGCTGCGCTGAACCAA-CYCAACCCAAAACVGGAGAC
		Begonia_mannii               ???TGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAAAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGGAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATTTTTTTTTATAATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTGCCGTGCAGAACAACCCGCGAACTCGTTGTGCAACGAACCCCGCAAGGGGAGTGGCGAGGA-GTGTTGGGGCG----------------CAGGCCCTGCAT-----------TTCGCGCGCACCTTAACGAACCCCC-GGCGCGAGTCGCGTCAAGGAA--TATATAAC-TG-AGAACGCGGTTCCTCTGCCTC-G-CCTGCGG-GATGC----AGGGG-CGCTGTGARCTTGCGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATKGCAGAA-CCCGTGAACCATCGAGTTTTTGAACGCAAGTKGCGCCCGAAGCCTTCTGGCGGARGGCACKTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----ACCAAAAAAA----TGC----TC-GATCTCTTC--TT-----------------GGGGTTGGGTTGGACTTT-GGTC--GGACGAGGGGG-CGATGTTTGGCTCCCGTGCGAGCGATTGCGCGGTTGGCCTAAAGTTGAGTC-TCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGATGCGGTAT-CGGGAA--TTCCC--TTTGCGACCCTGCTGCGCTGAAACAA-CGCAACACCAAACBGGAGAC
		Begonia_fusialata            AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAAAT-GAAATTTATAGTGCAGAACAACCCGCGAACACGTTGTACAACGAACCCCGCAAGGGGAGTGGCGAGGA-GTGTTGGGGCG----------------CATGCCCCGCAT-----------TTCGCGTGCACCTTAACGAACCCCC-GACGCGAGT?GCGTCAAGGAA--TATATAAC-TG-AGAACGC?GCTCCTCTGGCCC-G-CTTGCGG-GATGT----AGGGG-TGCTGCGAGCTTGCGAATCA--CACACGACTCTCGAC?ACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCC---GC---CGCCAAAAAA-----TGC----TC-GATCTCTTC--TT-----------------GGAGTTGGGATGGACTTT-GGTC--GGACGAGGGGG-TGATGTTGGCCTCCCGTGCGAACGATCGCGCGGTTGGCCTAAAGTCGAGTCCTTGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCTTCGGCGAAATACGTTGCACGTGCGAGACGCTGTATGCGGGAA--CTC?CAC---GCGACCCTGCTG?GCTGAAACAA-CTCAACAAAAAACCACAGCC
		Begonia_preusii              AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTTGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAAT?C----------GACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACCCGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGTCGGGGCG----------------CAAGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTTGCGTCAAGGAA--TATTTAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CTTGCGG-GATGC----AGGGG-CGTTGTGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATTTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAAA----TGC----TC-GATCTCTTC--TT-----------------GGGGTGGGGCTGGACTTC-GGTG--GGACGAAGGGG-CGATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGGCGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCAC---GCGACCCTGCTGCGCTGAAACAA-CYCAACCCAAAACCGGAGAC
		Begonia_polygonoides         ???TGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAAAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGGAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTATAATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACCCGTTGTGCAACGAACCCCGCAAGGGAAGCGGCGAGGA-GTGTTGGGGCG----------------CAAGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGC----AGGGG-CGCTGCGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCTGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAA-----TGC----TC-GATCTCTCC--TT-----------------GGGGTCGGGCTGGACTTT-GGT--GGGACGAGGGGG-CGATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTT--GT--TAGCCCTCGGCGAAACACGTTGCACGTGCGAGATGCGGTAT-CGAGAA--CTCCCAC---GCGACCCTGCCGCGCTGAAMCACACYCAACCMAAAACAGGAGAC
		Begonia_gabonensis           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAACCAAATAATGGTTTCGAGAGCAAGAACAAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAACGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTATAATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACCCGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGTTGGGGCG----------------CAAGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATTTAAC-TG-AGAACGCGGCTCCTCTGCCCC-G-CCTGCGG-GATGC----AGGGG-CGCTGCGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAA-----TGC----TC-GATCTCTCC--TT-----------------GGGGTCGGGCTGGACTTT-GGT--GGGACGAGGGGG-CGATGTTGGCCTCCCGTGCGAGCGATTGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTTTAT-TAGCCCTCGGCGAAACACGTTGCA-GTGCGAGACGCGGTAT-CGAGAA--CTCCCAC---GCGACCCTGCTGCGCTGAAACACAA-A-ACCAAAAACCGGAGCC
		Begonia_ebolowensis          AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAACCAAATAATGGTTTCGAGAGCAAGAACAAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGT????????????????????GTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGTTGGGGCG----------------CAAGCCCCGCAT-----------TTCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TATGTAAC-T--AGAACGCGGCGCCTCTGCCCC-G-CCTGCGG-TATGC----AGGGG-CGCTGCGAGCTTGAGAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGTAGCCCCCC--GC----GCCAAAAAA-----TTC----TC-GATCTCTTC--TT-----------------GGGGTCGGGCTGGACTTT-GGT--GGGACGAGGGGG-CGATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTAGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCAC---GCGACCCTGCTGCGCTGAAACACAA-A-ACAAAAAACCGGAGCC
		Begonia_molleri              AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATAGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTG?CCATGTCAATACGGACAAAAAAT-GAAATTT?TAGTDCAGAACAACCCGCGAACCCGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAAGA-ATGTTGGGGCG----------------CAAGCCCCGCAT-----------TCCGCGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TCTTTAA-TTG-AGAACGCGGTGCCTCTACCCCCG-TTTGCGG-GATGC----AGGGG-CACTGTGAGCTTGCCAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAAGGCACGTCTGCCTGGGCGTCACGCATCGCAGCCCCCC--GC----GCCAAAAAAA-----GC----TC-AATCTCTTC--TTCC-GG----------TTGGGGTTGGGCCGGACTTT-GGTC--GGACGAAGGGG-CAATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TGGCCCTCGGCGAAATACGTTGCACGTGCGAGACGCGGTAT-CGGGAA--CTCCCAC---GCGACCCTGCTGCGCTGAAACAA-CCCAACACAAAACGAAAGAC
		Begonia_furfuracea           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTTGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAAT?C----------GACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACCCGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGCTGGGGCG----------------CACGCCCCACAT-----------TCCACGCGCACCTTAACGAACCCCCCGGCGCGAGTCGCGTCAAGGAA--TCTTTAA-TTG-AGAACGCGGCGCCTTTGCCCC-G-CCTGCGG-GATAC----AGGGG-TGCTGCGAGCTTGCCAATCA--CACACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCATCGCAGCCCCCC--GC----GCCAAAACAAA----GT---TT--GATCTCTTC--TTCTTGGAG--------TTGGGGTTGTGCTGGACTTT-GGTC--GGACGAGGGGG-TAATGTTGGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGAAATACGTTGCACGTGCGAGATGCAGTAT-CAGGAA--CTCCCAC---GCGACCCTGCTGCGCTGACACCA-CGCAACAACACACCAAAGAC
		Begonia_subalpestris         AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATAGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACCCGTTGTGCAACGAACCCCGCAAGGGGAGCGGCGAGGA-GTGTTGGG-C?----------------CACGCCCCGCAT-----------T?CGCGCGCACCTTAAC?AA?CC?CCGGC?CGAGTCGC?TC?A?GAA--TCTTT??-TTG-AGAACGCGGTGCCTCT?CCCCCG-CTTGGGG-GATG?----A?GGG-C?CTGT?AGCTTGCCAATCA--CACAC?A?TC?CGAC?A?GGATA?CTCGG?TC?CGC?TCGAT?AAGAACGTAGTGAA?TGCGATCCTTGGTGTG?ATTG?A?AATCCCG?GAAC?A?CGA?TTTTTGAACGCAAGTTGC?CCCGAAGCCTTCTGGCGGAGGG?A?GTCTGCCTGGGCGTCACTCATC?CAGCCCCCC--GC----GCCAAAAAAACCCA----------AACCTCTTC--TTCC-GG----------TTGGGGT?GGGGTGGGCACA-GACT--GGTAGAAAGGG???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????AACCAA-CTCAACCCAAAACSAAAGAC
		Begonia_cavallyensis         AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATT-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAGAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATTCTCCACCGACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGT---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------CA-ACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGCGGAGGGCACGTCTGCCTGGGCGTCACGCAT?GTAGCCCCCC--GC----ACC??AAAAT----TGC----TC-AATCCCTTCCCTTCTCGGAG--------TTGGGGTTGGGCTGGACTTT-GGTC--GGACGAGGGGG-?GA?GT?GGCCTCCCGTGCGAGCGATCGCGCGGTTGGCCTAAAGTCGAGTCCTCGGCGTCGCGCAGTGCGACCGACGGTGGTTTT----TAGCCCTCGGCGA?ATACGTTGCACG?GCGAGACGCTGTAT-AGGGGA--CCACCAC---GCGACCCTGCTGCGCTGAACCACACRCAAMAAA?AAMMCCAGAC
		Begonia_bogneri              AAGTGATAACTTTCAAATTCAGAGAA-CCCTGG-AAT--AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAAAGCAAGAACGAAAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAACCCTTCCAGCGAAACTCCAGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAATAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CGC--AACTTCAAGGTGAAAAAAGAATCAAATATTCATTAATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAACTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACTAGTTAACCATCG-TCCCCATGTGGGAATTGT-GAGG-CGTGTTGGAGCTGCGACACAAGCTTGCGCGTGCCCTG---CCTTGCATGCCTC---GCAAACCATAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TCTTTAAC-TG-AGAACGCCGCTCTCCTACCCC-GACTT-CGG-GATGY----AGGGG-TGCTGCGAACTTGCTTATCA--TGAACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACACATCGTAGCCACCCAC------ACCAAA---------GC---TTC-AAGC-TAT---------------------------------------------------GAGGTGG-TAATGTTGACCTCCCGTGAGAGCAAGTGCACGGTTGGTCTAAATTTGAGTCCTCGGCGTTGCGCAGTGCGATCGGCGGTGGTT--GCCAT-GCCCTCGGCGAATTACATTGCACTTGCGGGACGCATGTT-CGGGAA--CTCC---TTTGCGACCCTATTGTGCTGAAAAAC?A-A-ACAAAAAAGCCC?TTA
		Begonia_betsimisaraka        AAGTGAT?RCTTTCAAATTCAGAGAAACCCTGG-AATAAAAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAACAAATAATGGTTTCGAAAGCAAGAACGAAAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAACCCTTCCAGCGAAACTCCAGAAAG---GATGAAAGATAAACCTATATACATACGTATACGTACTAAAATAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTCAAAATGAACGAATTGTTGTAAATCAATT--CGC--AACTTCAAGGTGAAAAAAGAATCAAATATTCATTAATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAACTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGTGCAGAACAACCCGCGAACTAGT?AACCATCG-TCCCCATGTGGGAATTGT-GAGGC-GTGTTGGAGCTGCGGCACAAGCTTGCGCGTGCCCTG---CCTTGCATGCCT?---GCAAACCATAACGAACCCC--GGCGCGAGTTGCGTCAAGGAA--TCTTTAAC-TG-AGAACGCCGCTCTCCTAGCCC-GACTT-CGG-GATGC----AGGGG-TGCTGCGAACTTGCTTATCA--TGAACGACTCTCGACAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGTGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACACATCGTAGCCACCCAC------ACCAAA---------GC---TTGCAAGCTTAT---------------------------------------------------GAGGTGG-CAATGTTGGCCTCCCGTGACAGCAAGTGCACGGTTGGTCTAAATTTGAGTCCTCGGCGTTGCGCAGTGCGATCGGCGGTGGTT--GCCAT-GCCCTCGGCGAATTACATTGCACTTGCGGGACGCATGTT-CGGGAA--CTCC---TTTGCGACCCTATTGCACTGAACAAC?A-A-ACA???AASCCC?TTA
		Begonia_squamulosa           AAGTGATAACTTTCAAATTCAGAGAAACCCTGG-AATG-AAAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAAGAAATAATGGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCTTTCCAGCGAAACTCCGGAAAG---GATGAAAAAGAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAAT-TTTTTTTC-AATGATATAGTGAAAATGAACGAATTGTTGTAAATCAATT--CCC--AACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACAAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGT???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????CAACAA-CWCAACAAACCACBAAAGAC
		Begonia_eleagnifolia         AAGTGAGTACTTTCAAATTCAGAGAAACCCTGG-AATG-ATAAATGGGCAATCCTGAGCCAAATCCGGTTTTCCGAAAAGAAATAATGGTTTCGAAAGCAAGAACGAGAAAAACAAAAAGGATAGGTGCAGAGACTCAATGGAAGCTGTTCCAACAAATGGAGTTGACTGCAGTTGCGTTAGTAAAGGAATCCAAATCCTTCCAGCGAAACTCCGGAAAG---GATGAAAAATAAACCTATATACATACGTATACGTACTAAAAAAGTATTTCAAATCA---GATTAATGACGACTCGAATGTTTTTTTC-AATGATATAGTGAAAATGAACGAATTGTTGTAAATCAATT--CCC--AACTTCAAGTTGAAAAAAGAATCAAATATTCATTGATTAAATATCATTCACTCCATCATAATCTGATAGATCCGTTGAAGAATTGATTC-AATCAGACGAGAATAGAATAAAGATAGAG--TCCCATTCTACATGTCAATACGGACAAAAA-T-GAAATTTATAGT???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????CAACAA-C?CAACAACCCACVAAAGAC
	;
END;

BEGIN CODONS;
CODONPOSSET * CodonPositions = 
 N: 1 - 1230 ; 
 CODESET * UNTITLED = Universal: all ; 
 END;

BEGIN ASSUMPTIONS;
CHARSET trnL = 1 - 524 ; 
 CHARSET ITS = 525 - 1203 ; 
 CHARSET Morphology = 1204 - 1230 ; 
 CHARPARTITION all = trnL: 1 - 524 , ITS: 525 - 1203 , Morphology: 1204 - 1230 ; 
 CHARPARTITION ITSvtrnL = trnL: 1 - 524 , ITS: 525 - 1203 , Default: 1204 - 1230 ; 
 CHARPARTITION ITSvmorph = Default: 1 - 524 , ITS: 525 - 1203 , Morphology: 1204 - 1230 ; 
 CHARPARTITION trnLvmorph = trnL: 1 - 524 , Default: 525 - 1203 , Morphology: 1204 - 1230 ; 
 OPTIONS DEFTYPE = unord PolyTcount = MINSTEPS ; 
 END;