NEXUS
[File created by TreeBASE:  1/16/09  15:31:12]
[Matrix accession#: M3496]

BEGIN TAXA;
	TITLE M3496;
	DIMENSIONS  NTAX=39;
	TAXLABELS
		Stylomesus_sarsi	Stylomesus_menziesi	Stylomesus_hexapodus	Contrarimesus_curtispinis	Ischnomesus_bispinosus	Ischnomesus_antarcticus	Contrarimesus_franklinae	Fortimesus_zuluensis	Fortimesus_thomsoni	Gracilimesus_tenuispinis	Fortimesus_scabriusculus	Fortimesus_robustus	Haplomesus_quadrispinosus_sensu_Brandt	Haplomesus_quadrispinosus	Haplomesus_bifurcatus	Fortimesus_profundicolus	Haplomesus_ornatus	Gracilimesus_orientalis	Gracilimesus_insignis	Gracilimesus_gorbunovi_sensu_Svavarsson	Gracilimesus_gorbunovi	Fortimesus_gigas	Fortimesus_consanguineus	Fortimesus_cornutus	Fortimesus_concinnus	Fortimesus_brevispinis	Cornuamesus_longiramus	Cornuamesus_biscayensis	Gracilimesus_tropicalis	Gracilimesus_corniculatus	Gracilimesus_angustus	Gracilimesus_celticensis	Gracilimesus_hanseni	Heteromesus_spinosus	Heteromesus_similis	Heteromesus_schmidtii	Heteromesus_granulatus	Heteromesus_frigidus	Heteromesus_drachi
	;
END;

BEGIN CHARACTERS;
	TITLE M3496;
	DIMENSIONS NCHAR=79;
	FORMAT SYMBOLS = " 1 2 3 4 5 " MISSING = ? GAP = - ;
	CHARSTATELABELS
		1 Per.5_fusion / freely_articulated fused_with_pereonite_6, 2 Per.6_fusion / freely_articulated fused_with_pereonite_7, 3 Per.7_fusion / freely_articulated fused_with_pleonite_1, 4 PL_1_fusion / freely_articulated fused_with_pleotelson, 5 Haplomesus_segmental_artic. / 'articulation absent- no segmental demarcation' 'articulation present- rudimentary- non-functional', 6 Per.7_length / length_not_reduced_compared_to_pereonite_6 'length reduced- less than pereonite 6' pereonite_7_completely_undeveloped, 7 Head_anterolat._projections / antennae_emerging_directly_from_head squat_anterolat._projections_supporting_antennae robust_rounded_anterolat._projections_supporting_antennae 'long- thin anterolat. projections supporting antennae', 8 Head_lobe_on_ventrolateral_margin / absent_in_lateral_view present_in_lateral_view, 9 Spination_pattern_on_pers.1_to_4 / identical_for_males_and_females sexually_dimorphic, 10 Per.1_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 11 Per.1_female_anterolat._spines_length / 'elongate- length near width of pereonite 1' 'short- length near pereonite 1 length' 'tiny- length distinctly less than pereonite 1 length', 12 Per.1_female_paired_lateral_spines / no_lateral_spines_or_tubercles paired_lateral_simple_spines paired_lateral_pedestal_spines paired_lateral_rook_spines paired_lateral_tubercles, 13 Per.2_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 14 Per.2_female_paired_lateral_spines / no_lateral_spines_or_tubercles paired_lateral_simple_spines paired_lateral_pedestal_spines paired_lateral_rook_spines paired_lateral_tubercles, 15 Per.3_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 16 Per.3_female_paired_lateral_spines / no_lateral_spines_or_tubercles paired_lateral_simple_spines paired_lateral_pedestal_spines paired_lateral_rook_spines paired_lateral_tubercles, 17 Per.4_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 18 Per.4_female_paired_anterolat._spines_or_tubercles / no_anterolat._spines_or_tubercles anterolat._simple_spines anterolat._pedestal_spines anterolat._rook_spines anterolat._tubercles, 19 Per.4_female_with_posterolat._spines_or_tubercles / posterolat._spines_or_tubercles_absent posterodorsal_spines_or_tubercles, 20 Per.4_female_anterolat._spines_length / 'elongate- length near width of pereonite 1' 'short- length near pereonite 1 length' 'tiny- length distinctly less than pereonite 1 length', 21 Per.5_anterodorsal_spines_or_tubercles / anterodorsal_spines_or_tubercles_absent anterodorsal_rook_spines anterodorsal_pedestal_spines anterodorsal_simple_spines anterodorsal_tubercles, 22 Per.5_posterodorsal_spines_or_tubercles / no_posterodorsal_spines_or_tubercles posterodorsal_simple_spines posterodorsal_rook_spines posterodorsal_pedestal_spines posterodorsal_tubercles, 23 Per.5_spines_or_tubercles_lateral_row / no_lateral_row_spines_or_tubercles lateral_row_spines lateral_row_tubercles, 24 Per.6_dorsal_spines_or_tubercles / no_dorsal_spines_or_tubercles dorsal_simple_spines dorsal_rook_spines dorsal_pedestal_spines dorsal_tubercles, 25 PL_1_lateral_spines_or_tubercles / no_lateral_spines_or_tubercles lateral_simple_spines lateral_pedestal_spines lateral_rook_spines lateral_tubercles, 26 PLtn._dorsal_surface_axial_ridge / 'smooth arched- not separated from lateral fields ' 'weakly vaulted- separated from lateral fields by shallow elongate concavities' 'strongly vaulted- separated from lateral fields by distinct indented lines', 27 PLtn._axial_ridge_length / axial_ridge_not_truncated axial_ridge_truncated, 28 PLtn._axial_ridge_distal_tip / medially_smoothly_arched 'medially incised- bilobed posteriorly' 'medially indented- hollow posteriorly', 29 PLtn._at_uropod_insertion / produced_from_margin indented_into_margin inflected, 30 PLtn._with_dorsal_spines_or_tubercles / dorsal_spines_or_tubercles_absent dorsal_tubercles dorsal_rook_spines dorsal_simple_spines dorsal_pedestal_spines, 31 PLtn._spines_or_tubercles_posterolat._marg. / spines_or_tubercles_absent spines_or_tubercles_present, 32 PLtn._type_of_spines_posterolat._marg. / simple_spines pedestal_spines rook_spines tubercles, 33 PLtn._terminal_margin_with_spines_or_tubercles / spines_or_tubercles_absent tubercles paired_simple_spines paired_pedestal_spines, 34 Antennula_num.__articles / two_or_less_articles 3_articles 4_articles 5_articles 6_or_more_articles, 35 Antennula_article_2_length / subequal_to_article_1_length less_than_2_times_article_1_length approximately_2_times_article_1_length greater_than_three_times_article_1_length, 36 Antennula_article_2_insertion / inserting_on_article_1_anteriorly anterodorsally dorsally, 37 Antennula_article_2_shape / 'straight- not curved at proximal insertion' strongly_curved_at_proximal_insertion, 38 Antennula_article_2_ventromedial_setae / 2_or_less_ventromedial_setae 3_ventromedial_setae 4_ventromedial_setae 5_ventromedial_setae 6_ventromedial_setae, 39 Antennula_article_2_distal_tip_shape / 'produced distally beyond insertion of next article- distal article inserting subapically' 'not produced distally- next article inserting apically', 40 Antennula_article_2_lateral_margin / robust_setae_absent row_robust_setae one_robust_seta, 41 Antennula_distal_articles_length / 'elongate- longer than article 2' subequal_to_article_2 'small- shorter than article 2' 'tiny- less than 0.3 article 2 length', 42 Antennula_article_3_length / article_3_longer_than_article_4 similar_length_to_article_4 'terminal- no distal articles', 43 Antennula_article_3_shape / 'elongate and tubular- much longer than wide' 'short and tubular- length near width' 'squat- wider than long', 44 Antennula_terminal_article_length / shorter_than_penultimate_article longer_than_penultimate_article, 45 Antenna_article_3_cuticle_smooth / smooth granulated, 46 Antenna_female_article_3_ventromedial_spines / no_ventromedial_spines ventromedial_spines, 47 Labrum_knobs / absent indistinct_low_projection present, 48 Maxilliped_palp_articles_width / article_2_wider_than_article_3 'articles 2-3 width subequal' article_3_wider_than_article_2, 49 Maxilliped_epipod_spines / epipod_spines_present epipod_spines_absent, 50 Per.bases_proximal_part_of_shaft / projecting_dorsally_above_shoulder dorsal_projection_absent, 51 Per.bases_proximal_shoulder / spines simple_spines bifid_spines rook_spines, 52 Per.I_shape / weakly_carposubchelate strongly_carposubchelate, 53 Per.I_merus_dorsal_surface / without_robust_setae with_distinctly_robust_setae, 54 Per.I_carpus_palm_vent._marg. / palm_or_distinctly_wider_part_absent proximally_expanded medially_expanded medially_expanded, 55 Per.I_carpus_vent._marg._robust_elongate_setae / 1_elongate_robust_seta 2_elongate_robust_setae row_similar_robust_setae 'fine setae only (no robust seta)', 56 Per.I_carpus_vent._marg._short_proximal_setae / several_setae_proximal_to_elongate_setae one_seta_proximal_to_elongate_seta setae_proximal_to_elongate_setae_absent, 57 Per.I_carpus_vent._marg._distal_setae_on_palm / several_distal_setae_on_palm 1_distal_seta_on_palm distal_setae_on_palm_absent, 58 Per.I_propodus_vent._marg._setae / several_robust_setae robust_setae_absent one_robust_seta, 59 Per.VII_presence / 'present- shorter than Per.VI' absent, 60 Pereopods_II, 61 PL.II_operculum_shape / broadly_triangular 'flap-like- much longer than wide' 'narrow proximal neck- almost circular posteriorly' 'narrow proximal neck- laterally convex- broadening posteriorly to rounded angles- posterior margin weakly convex' 'narrow proximal neck- laterally convex- broadening posteriorly to rounded angles- posterior margin weakly concave', 62 PL.II_operculum_setae / plumose_setae_absent plumose_setae_present, 63 Uropods_num.__articles / one_article 'two articles- protopod distinct', 64 Female_uropods_length_relative_to_pleotelson / extending_near_posterior_margin_of_pleotelson 'minute- not extending near posterior margin of pleotelson' 'elongate- extending beyond posterior margin of pleotelson', 65 Uropods_female_length_or_width / less_than_five_times_as_long_as_wide greater_than_or_equal_to_five_times_as_long_as_wide, 66 Uropods_female_length_or_length_of_pleotelson / less_than_0.4_length_of_pleotelson greater_than_or_equal_to_0.4_length_of_pleotelson, 67 Per.1_male_anterolat._spines_length / 'extremely elongate- length distinctly greater than width of pereonites' 'elongate- length near width of pereonite 1' 'short- length near length of pereonite 1' 'tiny- length distinctly less than length of pereonite 1', 68 Per.2_male_paired_anterolat._spines / no_paired_anterolat._spines 1_pair_of_anterolat._simple_spines 1_pair_of_anterolat._pedestal_spines 1_pair_of_anterolat._rook_spines 1_pair_of_anterolat._tubercles, 69 Per.3_male_paired_anterolat._spines / no_paired_anterolat._spines 1_pair_of_anterolat._simple_spines 1_pair_of_anterolat._pedestal_spines 1_pair_of_anterolat._rook_spines 1_pair_of_anterolat._tubercles, 70 Per.4_male_with_paired_anterolat._spines / no_paired_anterolat._spines 1_pair_of_anterolat._simple_spines 1_pair_of_anterolat._pedestal_spines 1_pair_of_anterolat._rook_spines 1_pair_of_anterolat._tubercles, 71 Per.4_male_length_of_anterolat._spines / 'elongate- length near width of pereonite 1' 'short- length near length of pereonite 1' 'tiny- length distinctly less than length of pereonite 1', 72 Per.4_male_posterolat._spines_or_tubercles / no_posterolat._spines_or_tubercles posterolat._spines_or_tubercles, 73 Antenna_flagellum_form_male / 'decreasing in width distally- tubular- longer than wide' 'some segments expanded- wider than long- with more setae on distal margins than juveniles or females', 74 PL.I_male_setae_on_distal_margin / setae_absent simple_setae blunt_setae plumose_setae, 75 PL.II_male_position_of_setae_on_protopod / setae_on_lateral_or_distal_margin setae_on_distal_margin_only setae_absent, 76 PL.II_stylet_length / 'short- reaching end of protopod' 'elongate- reaching beyond end of protopod', 77 PL.II_stylet_tip_shape / 'thread-like' thick_and_blunt thin_with_medial_blade, 78 Uropods_male_length_relative_to_pleotelson / not_extending_near_posterior_margin_of_pleotelson 'extending near posterior margin of pleotelson (+ or - 10pct)' extending_beyond_posterior_margin_of_pleotelson, 	79 Uropods_male_length_or_pleotelson_length / less_than_0.5_length_of_pleotelson greater_than_0.5_length_of_pleotelson
	;
	MATRIX
		Stylomesus_sarsi                              1122-231????????????11111211211-1?321221????2??12??31112111242232?4211-12211132
		Stylomesus_menziesi                           1122-13?1-----------11111111211-1533122132112-??2??3?11213?1--2----111-1?213122
		Stylomesus_hexapodus                          1122-231?131111111-111111111211-113313?14???21?22213?2121133412322-------------
		Contrarimesus_curtispinis                     22222111?121121511-111111211111-154311213212?1?322?2211331?1322312?????????????
		Ischnomesus_bispinosus                        1111-1111131111111-111111111211-15421221211111112?13221112112?23113111-11223131
		Ischnomesus_antarcticus                       1111-1111-----------111??1112?1-?542112111121-?1221312211111--2---3112221422231
		Contrarimesus_franklinae                      22222211?2111111123111111211111-15431321311121?22212211321114223??222231?2111??
		Fortimesus_zuluensis                          22221211?521555551-151111211211-1?431??1311?21?22??211231111??1111-------------
		Fortimesus_thomsoni                           2222111??-----------15253211211-3?????2??????-??221211131?12--1----11232?1???11
		Gracilimesus_tenuispinis                      22221111?1211111121111111111212115431121311111?122?2111323?1421311-------------
		Fortimesus_scabriusculus                      2222111??1-1111111-211211211211-354312223212?1?22??2?12311?1221211-------------
		Fortimesus_robustus                           2222111??135155151-251111211211-153312213111?1?32??2212311??421211??????121111?
		Haplomesus_quadrispinosus_sensu_Brandt        22221111?-----------11111322211-1543122131112-?2221211122321--1---1121-11213111
		Haplomesus_quadrispinosus                     22221211?121111111-111111311211-15431321311121?32212111221113?1111-------------
		Haplomesus_bifurcatus                         22221211?-----------11111313211-1?431221??1?1-?32??2?1122112--1---2121-11213111
		Fortimesus_profundicolus                      2222121??(25)22(25)25221-251111211221-344314223112?1?32??2212311??411111-------------
		Haplomesus_ornatus                            22221211?-----------11111322211-1543122131112-?3221211122111--1---2121-112???11
		Gracilimesus_orientalis                       2222111??-----------1111121121211542112131111-?12??21112232?--1---211211?211121
		Gracilimesus_insignis                         22221211?-----------1111121121211543122131111-?12212111323?1--1---11121112???31
		Gracilimesus_gorbunovi_sensu_Svavarsson       2222121111211111122111111111212115431121311111?22212111323114111111112111211121
		Gracilimesus_gorbunovi                        2222111??121111111-111111111212115431?21221111??2?????????????11?1-------------
		Fortimesus_gigas                              2222121?1131111111-111111211211-35431-223111?1?32??2112311??3112114111-11211111
		Fortimesus_consanguineus                      22221111?132121211-111111211211-154312223111???32??2212311??421111-------------
		Fortimesus_cornutus                           2222121??-----------11111211221-3543152231121-?12??221231121--1---2221-1?211111
		Fortimesus_concinnus                          2222111??-----------55253211241-354312212112?-?32??2112311??--1---252212?211111
		Fortimesus_brevispinis                        2222111??5325252523??1111211221-354312223112???22???????????221211-------------
		Cornuamesus_longiramus                        222211412121111111-114111211112115431321311211?11122111213115113222222211212332
		Cornuamesus_biscayensis                       2222124??-----------11111211112115431-2131112-?11??21123111?--1---222231?213331
		Gracilimesus_tropicalis                       222213112131111111-11111121121211?431121?????1?12212?112133?411111411231?2???21
		Gracilimesus_corniculatus                     22221111?-----------1111121121221543112132111-?1221211132331--1---2111-12212121
		Gracilimesus_angustus                         22221311?-----------111112112121144?112132121-?32212?1122331--1---3111-1?1???21
		Gracilimesus_celticensis                      222213112121111111-111111211212214431121321211?12212?1122331411111211221?111121
		Gracilimesus_hanseni                          2222131111211111122111111211212215431121321111?12212211123314211112112112211121
		Heteromesus_spinosus                          1222-11??4234343123211244311152244432211421122322143?41211123213?2-------------
		Heteromesus_similis                           1222-112?????????????????3?11????4????2?????2?122123?31211?3??1???4221-211???31
		Heteromesus_schmidtii                         1222-11?1532555151-111111311111-12432211433-22322123?41221123213?2-------------
		Heteromesus_granulatus                        1222-11??531111111-111111311111-12432311433-21322123?41211133213114??????????21
		Heteromesus_frigidus                          1222-11?1131111111-111111311111-12432211433-21322113?31211133213?(12)4112?21112332
		Heteromesus_drachi                            1222-112?-----------111??3111?1-?34323214121?-122123?31211?1--1---4221-11112331
	;
END;

BEGIN ASSUMPTIONS;
OPTIONS DEFTYPE = unord PolyTcount = MINSTEPS ; 
 END;