#NEXUS [written Thu Jan 07 18:37:21 SGT 2021 by Mesquite version 3.61 (build 927) at NoPsechrids.local/10.1.28.245] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=58; TAXLABELS Dichobune Cebochoerus Choeropotamus Siamotherium Anthracokeryx_tenuis Anthracokeryx_thailandicus Microbunodon Heptacodon Anthracotherium_chaimanei Anthracotherium_magnum Myaingtherium Bothriogenys_orientalis Bothriogenys_fraasi Bothriogenys_gorringei Bothriogenys_andrewsi Brachyodus_aequatorialis Brachyodus_onoideus Brachyodus_depereti Bothriodon Aepinacodon Elomeryx_crispus Elomeryx_borbonicus Afromeryx Sivameryx Merycopotamus_nanus Merycopotamus_dissimilis Libycosaurus_anisae Libycosaurus_bahri Epirigenys Morotochoerus Kenyapotamus_coryndonae Kenyapotamus_ternani Chororatherium Chorora_hippopotamine Archaeopotamus_qeshta Archaeopotamus_harvardi Hexaprotodon Palaeochoerus Kenyasus Perchoerus Amphimeryx Khirtaria Gobiohyus Mixtotherium Lophiomeryx Homacodon Xiphodon Diplobune Gujaratia Bunophorus Dacrytherium Paroxacron Entelodon Indohyus Merycoidodon Archaeomeryx Amphiraghatherium Hallebune ; END; BEGIN CHARACTERS; TITLE Hippopotamidae_late_Miocene_Baynunah_Formation; DIMENSIONS NCHAR=164; FORMAT DATATYPE = STANDARD RESPECTCASE GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4"; CHARSTATELABELS 1 number_of_lower_incisors / three two one, 2 lower_incisor_morphology / not_caniniform at_least_one_caniniform_lower_incisor, 3 relative_dimensions_of_lower_incisors / all_of_equal_size one_or_two_more_developped, 4 most_developped_incisor / 'I/2 ' 'I/3 ' 'I/1', 5 transverse_section_of_lower_incisors_crowns / strongly_irregular about_rounded, 6 lower_incisor_cervix_morphology_ / no_deep_indentation 'indented cervix, indentation as long as the root diameter on the cervix' deep_identation_longer_than_the_root_diameter_at_cervix, 7 crown_of_lower_I1 / straight__ 'spatulate (with convex mesial and distal border) ', 8 'presence of a median lingual pillar (lingual rib) on lower i1' / yes no, 9 Lower_canine_cross_setcion_at_cervix / subcircular elliptical_, 10 cristids_on_lower_canine_enamel_caps / none one_distal 'two, one mesial and one distal' one_mesial, 11 wear_on_lower_canine / distal_wear_facet_contact_with_canine_ mesial_wear_facet_contact_with_I3_, 12 groove_on_labial_side_of_lower_canine_ / no yes, 13 groove_on_lingual_side_of_lower_canine / no yes, 14 lower_canine_in_male / 'fang-like' premolariform incisiform, 15 crown_of_lower_canine_in_male / small_near_premolar_size at_least_twice_the_premolar_size 'prolonged growth to ever-growing ', 16 'p/1 roots' / one two, 17 paraconid_on_lower_premolars / no yes, 18 'elongated p/3 ' / 'no (shorter or equal than M/1 lenght)' 'yes (longer than M/1 lenght)', 19 'three lobed P/3' / no yes, 20 accessory_cusp_on_the_preprotocristid_of_lower_premolars / none one_or_more, 21 'lingual contour at cervix of P/4 in occlusal view' / convex_to_straight concave, 22 'labial wall on P/3 or P/4' / convex concave, 23 'Orientation of postprotocristid on P/3' / distal distolingual distolabial, 24 'high cingulid on labial face of P/3' / no yes, 25 'preprotocristid mesiolingualy curved on P/3' / no yes, 26 'mesial accessory cusp on preprotocristid on P/3' / no Shoulder_like_structure_on_lateral_view_of_preprotocristid_of_lower_premolars complete_accessory_cusp, 27 Change_in_the_orientation_of_the_preprotocristid_mesialy_to_the_junction_of_accessory_mesiolingual_crest_on_lower_premolars / no yes, 28 'Orientation of the endoprotocristid on P/4' / absent separated_from_postprotocristid_at_the_protoconid_apex_and_then_strait_and_distolingual fused_with_postprotocristid_in_part_and_then_curved_mesiolingually, 29 'entoconid on P/3' / never at_least_on_some_specimen, 30 'P/3 hypoconid' / no yes, 31 'distolingual cingulid on P/4 in lingual view' / forming_a_continuous_wall_lingually_until_the_distostylid_ reaching_the_level_of_the_distal_basin_and_keeping_be_shallow_until_the_distostylid 'reaching the level of distal basin and then being high when joigning the distostylid (distolingual notch of cingulid)', 32 'presence of a preentocristid on P/3 and/or on P/4' / no yes, 33 'mesiolingual secondary cristid on P/4 (cristid connecting lingual margin and preprotocristid)' / no yes, 34 'labial cingulid form a V (indentated) on P/4 before to reach the distal cingulid' / no_ yes_, 35 'marked postprotofossid on P/4' / absent present, 36 'postectoprotocristid on P/4' / no_ yes_, 37 'hypoconid on P/4' / no_ 'yes (even incipient) ', 38 'ectoprotofossid on p/4' / absent frequent, 39 premetacristid_on_lower_molars / strong reduced_or_missing_, 40 'paraconid on lower molars, almost on unworn specimens' / yes no, 41 lower_molar_trigonid_ / equal_in_height_with_talonid higher_than_talonid, 42 connection_between_premetacristid_and_preprotocristid_on_lower_molars / yes_ no_, 43 postectoprotocristid__on_lower_molars / absent 'reduced in the valley to fully developped at least on M/1', 44 'postprotofossid on lower molars at least on m/3' / no_ yes_, 45 'postmetacristid on M1-2' / curving_toward_postprotocristid_forming_a_transverse_bridge_with_it orientated_toward_the_centre_of_the_tooth forming_a_rounded_postmetaconulid_not_preferentiallyorientated joins_prehypocristid, 46 ectoprotofossid_on_lower_molars / absent present, 47 Ectometafossid__on_lower_molars / yes_ no_, 48 endometacristid__on_lower_molars / no_or_slightly_expressed_much_more_like_an_enamel_fold present, 49 postectometacristid_on_lower_molars / lightly_marked_to_absent always_present_and_well_marked, 50 premetafossid_on_lower_molars / present absent, 51 preentocristid_ / absent present, 52 preentocristid_connects / endohypocristid prehypocristid__toward_the_cuspid_apex prehypocristid_toward_its_mesial_extremity, 53 postectoentocristid__on_lower_molars / absent_ present_but_more_like_a_keel_on_cusp present_and_well_individualized_from_the_cusp, 54 ectoentocristid / present absent, 55 postentocristid__on_lower_molars / absent present_, 56 prehypocristid_dividing_in_two_mesial__arms_on_lower_molars / yes_ no_, 57 'prehypocristid inflated (not sallient when unworn) ) in transverse valley of lower molars' / no_ 'yes (even to form a conulid) ', 58 prehypocristid_reaches / 'median part of transverse valley (eventually labial)' lingual_part_of_transverse_valley labial_part_of_the_transverse_valley, 59 main_arm_of_prehypocristid_connects / 'trigonid distal walls (junction between cristids from metaconid and protoconid)' postmetafossid lingual_margin_of_transverse_valley postmetacristid, 60 posthypocristid_joins / nothing_or_distostyle postentocristid postectoentocristid, 61 endohypocristid_on_lower_molars / absent present, 62 posthypofossid_on_lower_molars / absent present, 63 entostylid__on_lower_molars_that_could_sometimes_be_linked_to_an_entocristylid / never frequently_present, 64 ectostylid__on_lower_molars / no_cingulid_ a_shallow_and_constant_cingulid_in_front_of_the_transverse_valley 'frequently developped cingulid in a /some stylid at least on m1', 65 ectocrystilid__on_lower_molars_ / no_ yes_even_if_variable, 66 'cingulid surrounding m/3 hypoconulid' / no_specimen_exhibiting_such_extension occasionally_bordering_the_labial_wall_, 67 'ectohypocristulid on M/3' / absent not_complete present_joigning_the_summit_of_hypoconulid, 68 'distostylid on M/1-M/2' / median lingual none, 69 'mesial part of Loop-like hypoconulid' / open_ pinched_, 70 entoconulid / no yes, 71 Number_of_upper_incisors / 3_ 2, 72 central_upper_incisor / 'morphologically similar to I2/I3 ' 'peg-like, morphologically different from others ' caniniform, 73 'I3/ reduced in size compared to I1/' / no_ yes_, 74 upper_canine_morphology / strong_with_circular_or_elliptic_cross_section_ 'strong and laterally compressed (blade-like)' premolariform, 75 canine_size_root / equivalent_to_slightly_longer_than_the_crown_ at_least_twice_the_size_of_the_crown_ prolonged_to_continous_growth_of_root_ prolonged_and_continuous_growth_of_crown, 76 dimorphic_upper_canine / no_ yes_, 77 'diastem C-P1/' / yes no, 78 'diastem P1/-P2/' / no yes, 79 number_of_upper_premolar / 4_ 5_ 3, 80 distolabial_crests_of_upper_premolars / simple with_a_maximum_of_two_accessory_cusps with_more_than_two_accessory_cusps_at_least_on_one_premolar, 81 'number of mesial crests on P1-3' / one two, 82 'disto-lingual basin in P2' / yes no, 83 'accessory cusp on disto-lingual cingulum of P3/' / none one_cingular_style 'protocone (surrounded by a cingulum)', 84 'Metacone on P3/' / no yes, 85 P3_root_pattern / 'one mesial root, two distal root not fused' one_mesial_root_and_fused_distal_ones, 86 P4_paracone / simple_with_crest complex_with_fossa very_complex_with_more_fossae, 87 'orientation of preparacrista on P4/' / mesial labial, 88 'postprotocrista on P4/' / absent present, 89 'Postprotocrista on P4/ joins' / base_of_paracone distostyle metastyle, 90 'preprotocrista on P4/ joins�' / mesiostyle base_of_the_paracone_then_mesiostyle parastyle, 91 'postectoprotocrista on P4/' / absent present, 92 P4_protocone_ / rounded crescentic, 93 'in lingual view protocone of P4/ is' / displaced_mesially median_, 94 'P4/ mesial margin' / concave convexe, 95 'strong development of distostyle on P4/' / no yes, 96 'mesial accessory cusp on P4/ that can be linked to mesiostyle' / no yes, 97 P4_metacone / absence presence, 98 P4_paracone_higher_than_the_protocone / slightly_higher_than_protocone much_higher_than_protocone, 99 P4_endoparacrista / absence_ presence, 100 'distal accessory cusp on postprotocrista of P4/ that can be linked to distostyle' / no yes, 101 height_of_lingual_cingulum_compared_to_unworn_protocone_height__on_upper_molars / one_third_ half_ no_cingulum, 102 'mesio-distal ribs development of labial cusps of upper molars' / almost_half_the_molar_lenght_ 'pinched (inferior to one third of molar lenght)' 'enlarged (superior to half the molar lenght) ', 103 postectoprotocrista / absent pr�sent, 104 postprotocrista / present absent, 105 Protocone_and_metaconule_junction_on_upper_molars / none_ 'premetaconule-postectoprotocrista ' 'premetaconule-postprotocrista ' postprotocrista_and_lingual_part_of_metaconule, 106 Premetacristule_divided_in_two_mesial_arms / no_ yes_, 107 ectometacristule_on_upper_molars / absent present_at_least_on_M1 not_frequent_and_only_on_M2_or_M3_, 108 postmetafossule / absent present, 109 secondary_cristule_labial_to_metaconule_eventually_an_endometacristule_or_enamel_knob / no_ yes_, 110 Distostyle_on_upper_molar / yes no, 111 distostyle_position_on_upper_molars_levels / metaconule_ metacone_, 112 secondary_ectometafossule_lingual_to_ectometacristule / absent_or_very_light present_mesially_at_least_on_M1_linked_to_ectometacristule, 113 paraconule_on_upper_molars / present_ absent, 114 'M2/ paraconule when present' / similar_in_size_with_protocone smaller_than_protocone, 115 postparacristule_extend_to_connect_ / none base_of_the_paracone transverse_valley, 116 preparacrista_connects_the_parastyle_ / 'no, separated by a groove ' yes_lingually yes_labially, 117 endoparacrista_on_upper_molars / absence presence, 118 ectoparafossa_on_upper_molars / no yes, 119 Ectocristyle / frequently_present absent, 120 premetacrista_and_postparacrista_connect / no_connection 'direct connection, forming a centrocrista' 'connection to mesostyle (direct, or indirect through ectocristyle)', 121 endometacrista_and_endometacristule_forming_a_transverse_crest_ / absence presence, 122 parastyle_development / enamel_knob smaller_or_equal_than_mesostyle larger_than_mesostyle, 123 premetacristule_invade_labial_part_of_the_transverse_valley / no yes, 124 position_of_metaconule_on_upper_molar / labial_side_of_the_protocone distal_side_of_the_protocone, 125 'M2/ metaconule' / similar_in_size_with_protocone smaller_than_protocone, 126 mesostyle_on_upper_molars / no yes, 127 mesostyle / enamel_knob half_to_the_size_of_labial_cusp larger_than_labial_cusp, 128 metastyle / reduced_to_enamel_knob_or_absent fully_developped_, 129 root_fusion_on_upper_molars / four_roots_with_occasional_fusion_close_to_cervix_the_apices_always_remaining_free fully_fused_lingual_roots three_roots, 130 lingual_cingulum_on_upper_molars / no yes developped_in_entostyle, 131 'hypocone on upper molars (at least M2/' / yes no, 132 'shape of M1/' / triangular quadrate, 133 'shape of M3/' / triangular quadrate, 134 'M3/ size' / 'Larger than M2/' 'equal in size with M2/' 'reduced (less than 60%)', 135 Mesiolingual_style_on_upper_molar_mesial_cingulum / no_ yes_, 136 symphysis_morphology_in_sagittal_section / elliptic dorsally_concave ventrally_concave, 137 'diastem C-P/1' / absent pr�sent, 138 bone_fusion_at_symphysis_in_adult_specimens / no_ yes_, 139 maximal_thickness_of_the_symphysis_in_sagittal_section / in_the_middle_part_ in_the_rostral_part_ in_the_nucal_part, 140 symphysis_extension / 'extends nuchally between C and P/1' 'extend nuchally between P/1 and P/3' 'extends nuchally to P/3', 141 number_and_position_of_main_external_foramen / numerous_ only_one_below_the_anterior_part_of_of_the_premolar_row_ 'two, one below the anterior part and the other below the posterior ', 142 'transverse constriction of mandible at C-p/1 diastema' / no_ yes_, 143 'P/1-P/2 diastema' / absent_ present_, 144 'P/2-P/3 diastema' / yes no, 145 Opening_of_internal_choanes / 'at M3/ ' 'nucal to M3/ ', 146 enamel_ornementation / no yes, 147 Schmelzmuster_composed_of_ / two_layers_ three_layers_ one_layer, 148 inner_radial_enamel_ / absent_ present_, 149 HSB_percent_of_Schmelzmuster / absent_ less_than_75%_ more_than_76%, 150 Outer_radial_enamel_ / less_or_equal_to_20%_ more_than_20%_, 151 Hsb_zone / thin_with_bands_always_less_than_100�m_ 'large (equal or more than 100�m) ', 152 'regular aspect (constant width)' / yes_ no_, 153 'HSB varaible (SD>20)' / no_ yes_, 154 HSB_angle_with_EDJ_ / '>70� ' '<70 ��', 155 orientation_of_HSB / straight_ bent_, 156 'HSB definition (decussation angle and size of transition zone)' / clear_ blurry_, 157 division_of_HSB / anastomosis_ bifurcation no_division, 158 HSB_configuration / curved_ transverse_, 159 synchronous_prism_undulation_on_horizontal_section / no_ yes_but_few__ yes_but_more_or_equal_to_4, 160 IPM_in_inner_portion / closed_sheath_ Inter_row_sheets_, 161 IPM_in_middle_portion / closed_sheath_ Inter_row_sheets_ no_IPM, 162 IPM_in_outer_portion / _closed_sheath no_IPM_, 163 prism_angle_with_EDJ_ / equal_or_more_than_60�_ less_than_60_�_ tend_to_diminish_in_the_inner_part, 164 prism_diameter / mean_between_3_and_3.9_�m small_diameter_mean_below_3_�m_ large_diameter_mean_above_or_equal_to_4 ; MATRIX Dichobune 000-00??12?0010010000010020000000000100010000000010-000100001002000000???????1000?1?0000-0001100000000010000000001010011020010-02001020000?120100001200110010??0??02 Cebochoerus 000-000?10?0020100000000000001000000100100010100000-100100000102002000???20001000020010110000100000002002001000001010001020100-0001111020121?01100002000000000000001 Choeropotamus 000-???????0020001000000010100000010000100001000000-001000000012000000???0????000100?0011001010010000100201110000110010201010111?111110010010011?1001111100001000002 Siamotherium 0???00??0000?00000000000000100000010010100000000001010010000110000200000001?0?0001(0 1)01100-000010000001000210000100111010100010100011110000021200101002000000001000002 Anthracokeryx_tenuis 000-00001?1000000100000000010010000000010100000110101001000010000020000001{2 3}?01000100?10????10???00000010111000100111010202010110??11100?1021(1 2)11001?????????????????? Anthracokeryx_thailandicus ???????????????0000000000001001000100001000000011012100100000001002000????????????00?101000100000000?00021{0 1}000100121010201010111011110011021211101?????????????????? Microbunodon 000-00001110000001000010000100100000000100000001101211010000000100200000013110000100?1110001010000000100200000100121010201010111011110011121211101002101000000000000 Heptacodon 000-????110000100000002000010000001010110000000110101111000110000000000?001110000110?1111011000000000010101000110111010202010121?211101001211001?1?????????????????? Anthracotherium_chaimanei ?0100101????????0?0000000001000010101001001100011010101010011110001001???01101000110?1111011000?000000101110101101210102010101110211101????{0 1}???101?????????????????? Anthracotherium_magnum 00100100000000100000000000010000101010010011000110102010100(0 1)110000200100001?110001101111111?000000000010111010(0 1)10121010202010111021110100101201101002000000001200000 Myaingtherium 000-000?0?????1100000000000100?000101001001100001110100100001000002001???????????????101001100000000?2101110000001100111000101001211101010?2?101?1?????????????????? Bothriogenys_orientalis ???????????????0000011201211001010100001000001011011101100010102101100??????0000(0 1)000?11100011010000000101110?00001110102010101100111100010211?1111?????????????????? Bothriogenys_fraasi ??????????0????0000010201112001?10100011000000011011101100011001110100???20?00001000?11101010000000?0010111000100121010202010111?11110011020111011?????????????????? Bothriogenys_gorringei {0 1}00?0011120001000000110012120010101000110000001(0 1)10122001000(1 2)0111110100000????????????1110101000000001110(1 2)1100010011101020201011101111001102121101111100001102??01?10 Bothriogenys_andrewsi ??????????????????????????????????????01010000100011210100120001110?00??????????????????????????????0?????00???00??1?102?2???111?1????0??????????1?????????????????? Brachyodus_aequatorialis 11110?1?1?0??1000000111011020011(0 1)0100011010000101111200000(0 1)200011111001212000000{0 1}000?1110101000000010111001000100121010202010111011111011120110111?????????????????? Brachyodus_onoideus 111100111200010000000100110200101010001101000010111121010112000011010012120000001000?111010100000001011100000010012101020201011101111{0 1}011120110111202000011001201010 Brachyodus_depereti 21110?--?????100000000001102001100100011010000101112200101120000110100???????00????0?1111001000000000100000{0 1}00100121010202010111?11?1?0111?0?101?1?????????????????? Bothriodon 0010000012000100000010001201000000100011000000011012200100120001?0011000020001000100?1111001010000000110011000000022011201010120011110011121111001?????????????????? Aepinacodon 0?10?0??????00100000100011010020011001110100001110122001011200011??1??00020001000100?111000111000000011000000000002201120101011(0 1)?11110011121101001?????????????????? Elomeryx_crispus 0010001012000100000(0 1)101010010011101000010000000110102001011210011101{0 1}(0 1)00011100000000?111100100000000011011110010012201120201011101111002101?111111?????????????????? Elomeryx_borbonicus 00100010120001000001101012010011101000110000001(0 1)111220010112000211011000011100000000?111(0 1)0010000000001(0 1)0100100100122011201010111021110021021111111112000011001111110 Afromeryx {0 2}???????1????010000110101201001?011000110100000110122001000000011?01100?0?1100011000?1111001000000000110000000101--1011202010111021110021021210111001100011011101111 Sivameryx 000-????110?00110001101012010010001000010000010110122001012(0 2)0101110110??????00?11000?1110001000000000110000000100112011201010111?211100210202101?1210---------112110 Merycopotamus_nanus 000-0???110100110001101012010010001010110100001011121(0 1)010120000211011000001?0001100011111001010000000100000000101--20112010101111211100210011101110011111111211111?2 Merycopotamus_dissimilis 000-0???11010021000110101201001000101011010000101112100101200002110110???0110001100011111001010000000100002000101--2011201010111121110021101200111111111111121112112 Libycosaurus_anisae (0 1)012100(0 1)11010021000110101201001000101011010000101112100101200002110110???020?0121000?1111001110000000100002000101--20112010101111?1110021020210111111111111121112110 Libycosaurus_bahri 201210011101002100011010120100100010101101000010111210010120000211011000102?0012100011111001010000000100002000101--201120101011111111002112021(0 1)111210---------112120 Epirigenys ????0111????????0?00010111-1002(0 1)1111100100(0 1)(0 1)00011012100010000112?1?100???????????????1111001101100011010(1 2)11010010120010201010110?111100??????????1002000011021101110 Morotochoerus ????????????????0???01????-1??2?0{0 1}11?10101111101101{0 1}100110000112102000??????????????????????????????(0 1)200201100000110010(0 2)00010100?111100??????????1?????????????????? Kenyapotamus_coryndonae ?0??12011?00102?000?112110-11121011(0 1)110100(0 1)1(1 3)101(0 1)012101110000112112000?????????10?10110110011011001112(0 1)020(0 1)100(0 1)10110110(0 2)000101000211110??1?2?0?(0 1)?120201011100121101{0 2} Kenyapotamus_ternani 00101???1??010200?00?12?10-??0????????0100(0 1)1(1 3)?01(0 1)01210111000011?1?2000?????????(0 1)0?10?2011001101?00111210201100(0 1)10110110(0 2)000101000211110{0 1}11020010?1?????????????????? Chororatherium ????????120??02?0??0??????-???????????110101?111001201?110??01???1{0 2}?00?????????20?101??110011?1?0011?200200100(0 1)00110110(0 2)00010100021???0??????????1?????????????????? Chorora_hippopotamine ?0??1???1200102?0??0??????-1???10?10111101013111001(1 2)0111103001121??0?0???03????20?20120110011111?0110200200100(0 1)0???0010(0 2)00010100021???0??????????1?????????????????? Archaeopotamus_qeshta 001012011?0010200?00112110-11121011011110111311?001201111030?11211?00000103???0?0?201??110011?1?0??10200200100(0 1)011-0011000010(0 1)00?21?1?0211(0 1)200?0?1?????????????????? Archaeopotamus_harvardi 001212011?0010200000112110-111210110111101113111001201(0 1)110300112112000000031010(1 2)002012011001101100110200200100(0 1)0(0 1)--0{0 1}10(0 1 2)00010100021111021102001(0 1)11?????????????????? Hexaprotodon 001212011?0010200000112110-111210110111101113111001(1 2)01(0 1)11030(0 1)1121120000000310(0 1)0(1 2)002012011001101100110200200100(0 1)0(0 1)--0{0 1}10(0 1 2)00010100021111021102001(0 1)11111111111121212111 Palaeochoerus 001(0 2)0100130110200000000000-000-00000101100110000110-1001101000010000000?1021(0 1)00000101000-0001101100022010011100001000010100100-01011110211?1?001?00020000000(0 1)0000000 Kenyasus 001(0 2)0100130100210000000000-000-00000101100012001000-11110010100100000001102?01000?00?10110010100100022010011000101001010100100-01011100201?1?00100112000000001001000 Perchoerus 001(0 2)0?0013011020000000?0??-0?0?00000101100110000100-?1?1001010010000000??02?(0 1)1000?00?000-000010010002201001000?001000011000100-0?011110?11??2?11?0?????????????????? Amphimeryx 0011000110?0020011101020000001-0000010110100301001120111013100000001100??2???1000?2100012201110000002100200001-00022011201110110201111020001211000210---------000002 Khirtaria ????????????????0000000000010000001010111001-0111110001102001000001{0 2}0{0 1}??????????????????????????????02000010100001101011100110-02110021??????????1?????????????????? Gobiohyus ????????????????100000001001002001001000100000001110000100001001001000????????????2?01011001010000001001000001-001010111020100-02111110??????????0?????????????????? Mixtotherium ????????1200001011100020000111-?0000100010000110110-101100010102000100???01?10000?2101111001000010000100200001-001020002020111112211110?1?????11?011100000000??00002 Lophiomeryx 001200??1????20011100020000101-000001011010030100112010101300002000110---2?00120011000112201110000000100200001-01--2011201110111?111100010?0201110?????????????????? Homacodon ???????????????????0??????????????????10010000100110000100001002000000?????????00?10000110001100000000103000000000010001020000-02101020??????????0?????????????????? Xiphodon ????????????????11101020000211010000101101000010110-200101100001000110???????001010111111201010000002100000001-000220012011101102211111???????01?01111000100(0 1)1100012 Diplobune 001000011210010001100000110211010000101001000010110-1101112200000001000212001000012011112201010000002100000001-001020012011101112011100100012001001111000{0 1}0001000012 Gujaratia 000-000?1210010010000000000000000000001011000010010-00110000000100100000011?11000120?00112000100010000003000001001101011100010-0201001000011??0100210---------000001 Bunophorus ????????????????1000000000000?200000001011000010110-001100000001000000???????????????000--001000000002003000001001000011000010-02100020??????????0?????????????????? Dacrytherium 0???00??1210020001000000110201210010100111000000010-1101012000100001000??20?100000201110-201011000010100000001-001020112021101112011100000?11001?0?????????????????? Paroxacron 000-000?1????2?111101020000201000000100110000010010-0101012000000001-1???????000012101111201010001002101000001-00002001201101111201112020011100000210---------000002 Entelodon 001(0 1)0001000000110100000000000000000010101100101101100011000010000000-00000??1100012010010-000100000012010-00000001001010000010-0110102020021100001111110000021?00002 Indohyus ????????????????0??000????01??0000001011100000101110000102001000001200?????????00?001000--00000001100200201001-001101011100110-0211?100????????????????????????????? Merycoidodon ????????010000100000002011011100101010010100-00001122101013200120001??00001?0000100000112201000000002000200001-01--2001202110111?21110010??1200110?????????????????? Archaeomeryx 000-?0??1??002001110102000010100000010010000301001120101013000000001?000010?0120012{0 1}?0110201000000000101200001-01--1010202111111?111100?1??1?011?0?????????????????? Amphiraghatherium ????????1????200111000000000112101101101000000101110000110001012002000???????0010121010110010010110012102011000001110{0 1}1(1 2)0001010021111000?021201(0 1)?1?????????????????? Hallebune ???????????????011101020000011?10?001010010000101110000100001001002000?????????00?21?101100100001?0012103000000001110111000(0 1)10-0?10{0 1}010???????11?0?????????????????? ; END; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE = UNORD POLYTCOUNT = MINSTEPS; TYPESET * UNTITLED = unord: 1- 164; END; BEGIN NOTES; TEXT TAXON = 1 TEXT = 'D leporinaMaterial in UM collection, but for anterior dentition character state in Sudre 1978 and some character infered from D. Jehenni (lBrunet and Sudre 1980)'; TEXT TAXON = 2 TEXT = 'C campichiiISEM and in Sudre 1978 (skull)'; TEXT TAXON = 3 TEXT = 'Choeropotamus depereti Sthelin 1908, Euzet (Gard, MP17) Sudre 1978 UM Euz3080 md d P/3-M/3UM EUZ3081 M sup dUM EUZ 40 md g P/3-M/2et Hooker et Thomas A NEW SPECIES OF AMPHIRHAGATHERIUM(CHOEROPOTAMIDAE, ARTIODACTYLA, MAMMALIA) Palaeontology, Vol. 44, Part 5, 2001, pp. 827�853, 1 plet Sudre 1978FROM THE LATE EOCENE HEADON HILL FORMATIONOF SOUTHERN ENGLAND AND PHYLOGENY OFENDEMIC EUROPEAN `ANTHRACOTHERIOIDS'''; TEXT TAXON = 4 TEXT = 'S krabienseMat�riel Krabi, ThailandeDucrocq 1999'; TEXT TAXON = 5 TEXT = '= A ulnifer, Pondaung fm Myanmar cr�ne complet AMNH 20017'; TEXT TAXON = 6 TEXT = 'Mat�riel de KrabiTF2639 md l P/1-M/2TF2638 skull r and l P3/-M3/TF2656 M/3 rDucrocq 1999'; TEXT TAXON = 7 TEXT = 'M minimumMat�riel de La Milloque (MP28)Poitiers'; TEXT TAXON = 8 TEXT = 'H occidentalismateriel de l''AMNH attention �galement utilisation de H. curtus pour dent inf'; TEXT TAXON = 9 TEXT = 'Materiel de Wai Lek, thailandTF2637 md l P/3-M/3TF2636 holotype skull r and l C, P1/-M3/ Ducrocq 1999'; TEXT TAXON = 10 TEXT = Mandibule_de_MoissacM_sup_de_DigoinM_sup_des_MillesBritish_Museum_palaisLyon_Phosporites_du_QuercyNassiet_Landes; TEXT TAXON = 11 TEXT = 'M kenyapotamoidespeut �tre = A panganTsubamoto 2011'; TEXT TAXON = 12 TEXT = 'Thailand, Bang Pu Dam Eocene supTF 2634TF 2633Ducrocq 1999'; TEXT TAXON = 13 TEXT = 'Schmidt 1917, Ducrocq 1997, material from SNM'; TEXT TAXON = 14 TEXT = 'Schmidt 1917, Ducrocq 1997, material from SNM'; TEXT TAXON = 15 TEXT = 'Schmidt 1917, Ducrocq 1997, material from SNM'; TEXT TAXON = 16 TEXT = 'Mc Innes 1951, Dineur 1981material from KNM'; TEXT TAXON = 17 TEXT = Dineur_1981; TEXT TAXON = 18 TEXT = 'Pickford 1991, Dineur 1981, material in collection at NHM London'; TEXT TAXON = 19 TEXT = B_velaunus; TEXT TAXON = 20 TEXT = AMNH; TEXT TAXON = 21 TEXT = Mat�riel_de_Detan_Dverce; TEXT TAXON = 22 TEXT = 'Geais 1934, Hellmund 1991, Kostopoulos et al. 2012, Material at FSL'; TEXT TAXON = 23 TEXT = Af_zelteni; TEXT TAXON = 24 TEXT = S_paleindicus; TEXT TAXON = 25 TEXT = 'Potwar plateau Chinji Fm, Pakistan Lihoreau eta al. 2007'; TEXT TAXON = 27 TEXT = 'Mat�riel de l''ONM, Tunis'; TEXT TAXON = 29 TEXT = E_lokonensis; TEXT TAXON = 30 TEXT = M_ugandanensis; TEXT TAXON = 33 TEXT = 'C. roobii, nov. gen. nov. sp.'; TEXT TAXON = 36 TEXT = A_harvardi; TEXT TAXON = 37 TEXT = H_garyam; TEXT TAXON = 38 TEXT = ref__Ginsburg_1974_table_ronde_sur_les_phospho_du_Quercy_Paleovertebrata; TEXT TAXON = 39 TEXT = 'Photo from Runsinga attention enamel microstructure d''un autre hyotheriinae (Aureliachoerus)'; TEXT TAXON = 41 TEXT = 'Amphimeryx murinus (Cuvier, 1822)o Temporal and geographical distributionLate Eocene, Western Europeo Origin of examined material- Escamps, Rosi�res 2 and St N�boule France (UM2) o References: (Sudre 1978, Hooker and Weidman 2000, Erfurt and Metais 2007)'; TEXT TAXON = 42 TEXT = 'casts of specimens of Khirtharia dayi, K. inflatus and K. aurea in UM2ref Orliac and ducrocq 2011and Thewissen et al. 19??'; TEXT TAXON = 44 TEXT = 'Material from UM2 from La defense and Egerkingenref: Erfurt and Metais 2007, Hooker and Wedman 2000'; TEXT TAXON = 45 TEXT = L_chalaniatifrom_material_from_GarouillasBrunet_et_Sudre_1987_Mayence; TEXT TAXON = 47 TEXT = 'X castrenseMaterial from Robiac RBN5004, 5067 Sudre 1969 et bretou Hartenberger 1974 '; TEXT TAXON = 48 TEXT = D_minor; TEXT TAXON = 49 TEXT = material_from_Pakistan_at_UM2; TEXT TAXON = 50 TEXT = 'B lysitensisand B. grangeri for P/3'; TEXT TAXON = 51 TEXT = D_ovinum; TEXT TAXON = 52 TEXT = P_valdenseMaterial_from_escamps_in_UM2; TEXT TAXON = 53 TEXT = material_from_villebramar_and_from_Quercy; TEXT TAXON = 54 TEXT = fromI_indirai_Kunmar_and_Shani_1985; TEXT TAXON = 55 TEXT = 'Specimen from the Big Badland south Dakotacollection Lavocat UM2 probably bullatuscranium +mandible, palate, P4/-M3/, P/4-M/2 and P/3-M/2'; TEXT TAXON = 56 TEXT = 'moulage d''Archaeomeryx optatus UM2 from AMNH 20321 and publication of Colbert 1941'; TEXT TAXON = 57 TEXT = A_neumarkensis_from_Geiseltal_and_A_weigelti_also_from_Geiseltal_and_Hooker_et_Thomas_2001; TEXT CHARACTER = 1 TEXT = 1_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 2 TEXT = 3_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 3 TEXT = modified_from_4_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 5 TEXT = 3_in_Orliac_et_al._2010_8_in_Boisserie_et_al._2010; TEXT CHARACTER = 6 TEXT = 4_in_Orliac_et_al._2010; TEXT CHARACTER = 11 TEXT = 5_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 12 TEXT = car_14_in_Boisserie_et_al._2010; TEXT CHARACTER = 15 TEXT = 7_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 16 TEXT = 13_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 20 TEXT = 9_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 21 TEXT = 'modified from Orliac 2010, car 52'; TEXT CHARACTER = 29 TEXT = 51_in_orliac_et_al._2010; TEXT CHARACTER = 35 TEXT = 59_in_Boisserie_et_al._2010; TEXT CHARACTER = 38 TEXT = 54_in_Orliac_et_al._2010; TEXT CHARACTER = 39 TEXT = 69_Boisserie_et_al._2010; TEXT CHARACTER = 43 TEXT = modified_from_67_in_Boisserie_et_al._2010; TEXT CHARACTER = 45 TEXT = 72_in_boisserie_et_al._2010; TEXT CHARACTER = 46 TEXT = 66_in_Boisserie_et_al._2010; TEXT CHARACTER = 49 TEXT = 71_in_Boisserie_et_al._2010; TEXT CHARACTER = 50 TEXT = 70_in_Orliac_et_al._2010; TEXT CHARACTER = 53 TEXT = modified_from_77_in_Boisserie_et_al._2010; TEXT CHARACTER = 55 TEXT = modified_from_78_in_Boisserie_et_al._2010; TEXT CHARACTER = 61 TEXT = 75_in_Orliac_et_al._2010; TEXT CHARACTER = 62 TEXT = 77_in_Orliac_et_al._2010; TEXT CHARACTER = 63 TEXT = 'seulement sur M/3 de A chaimanee'; TEXT CHARACTER = 66 TEXT = 79_in_Boisserie_et_al._2010; TEXT CHARACTER = 67 TEXT = 80_in_Boisserie_et_al._2010; TEXT CHARACTER = 69 TEXT = 29_in_Lihoreau_and_Ducrocq; TEXT CHARACTER = 74 TEXT = 6_in_Lihoreau_and_ducrocq_2007; TEXT CHARACTER = 79 TEXT = 10_modified_from_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 80 TEXT = 11_from_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 81 TEXT = 14_from_Lihoreau_and_Ducrocq; TEXT CHARACTER = 82 TEXT = 12?_in_Orliac_et_al._2010; TEXT CHARACTER = 83 TEXT = 16_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 85 TEXT = X_in_Orliac_2010; TEXT CHARACTER = 86 TEXT = a_r�ecrire_form_Orliac_et_al._in_prep_22; TEXT CHARACTER = 87 TEXT = 20_in_Orliac_et_al._2010; TEXT CHARACTER = 92 TEXT = from_orliac_in_prep_23; TEXT CHARACTER = 97 TEXT = X_in_Orliac_et_al._2010; TEXT CHARACTER = 98 TEXT = Orliac_2012_32; TEXT CHARACTER = 99 TEXT = Orliac_et_Ducrocq_2012_31; TEXT CHARACTER = 107 TEXT = Modified_from_48_in_Boisserie_et_al._2010; TEXT CHARACTER = 112 TEXT = modified_from_47_in_Boisserie_et_al._2010; TEXT CHARACTER = 117 TEXT = 21_in_Orliac_2012; TEXT CHARACTER = 118 TEXT = 36_in_orliac_in_prep_; TEXT CHARACTER = 124 TEXT = 27_in_orliac_2012; TEXT CHARACTER = 126 TEXT = 31_in_orliac_in_prep; TEXT CHARACTER = 130 TEXT = 34_in_orliac_in_prep; TEXT CHARACTER = 135 TEXT = 19_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 136 TEXT = 38_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 137 TEXT = 34_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 138 TEXT = modified_from_37_in_Lihoreau_and_Ducocq_2007; TEXT CHARACTER = 139 TEXT = 39_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 141 TEXT = 40_in_lihoreau_and_ducrocq_2007; TEXT CHARACTER = 142 TEXT = 33_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 143 TEXT = 35_in_Lihoreau_and_Ducrocq; TEXT CHARACTER = 144 TEXT = 6_in_Boisserie_et_al._2010; TEXT CHARACTER = 145 TEXT = 51_in_Lihoreau_and_Ducrocq_2007; TEXT CHARACTER = 146 TEXT = from_orliac_in_prep_26; TEXT CHARACTER = 147 TEXT = 1_in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 148 TEXT = 2_in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 149 TEXT = 3_in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 150 TEXT = 4_in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 151 TEXT = 5__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 152 TEXT = 6__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 153 TEXT = 7__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 154 TEXT = 8__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 155 TEXT = 9__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 156 TEXT = 10__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 157 TEXT = 11__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 158 TEXT = 12__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 159 TEXT = 13__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 160 TEXT = 14__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 161 TEXT = 15__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 162 TEXT = 16__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 163 TEXT = 17__in_Alloing_s�guier_et_al._soumis; TEXT CHARACTER = 164 TEXT = 18__in_Alloing_s�guier_et_al._soumis; END; BEGIN PAUP; outgroup gujaratia homacodon bunophorus; END; BEGIN MESQUITECHARMODELS; ProbModelSet * UNTITLED = 'Mk1 (est.)': 1- 164; END; BEGIN TREES; Title Hippopotamidae_late_Miocene_Baynunah_Formation; ID 0176dc6b633e47; LINK Taxa = Taxa; TRANSLATE [0] n0 Dichobune, [1] n1 Cebochoerus, [2] n2 Choeropotamus, [3] n3 Siamotherium, [4] n4 Anthracokeryx_tenuis, [5] n5 Anthracokeryx_thailandicus, [6] n6 Microbunodon, [7] n7 Heptacodon, [8] n8 Anthracotherium_chaimanei, [9] n9 Anthracotherium_magnum, [10] n10 Myaingtherium, [11] n11 Bothriogenys_orientalis, [12] n12 Bothriogenys_fraasi, [13] n13 Bothriogenys_gorringei, [14] n14 Bothriogenys_andrewsi, [15] n15 Brachyodus_aequatorialis, [16] n16 Brachyodus_onoideus, [17] n17 Brachyodus_depereti, [18] n18 Bothriodon, [19] n19 Aepinacodon, [20] n20 Elomeryx_crispus, [21] n21 Elomeryx_borbonicus, [22] n22 Afromeryx, [23] n23 Sivameryx, [24] n24 Merycopotamus_nanus, [25] n25 Merycopotamus_dissimilis, [26] n26 Libycosaurus_anisae, [27] n27 Libycosaurus_bahri, [28] n28 Epirigenys, [29] n29 Morotochoerus, [30] n30 Kenyapotamus_coryndonae, [31] n31 Kenyapotamus_ternani, [32] n32 Chororatherium, [33] n33 Chorora_hippopotamine, [34] n34 Archaeopotamus_qeshta, [35] n35 Archaeopotamus_harvardi, [36] n36 Hexaprotodon, [37] n37 Palaeochoerus, [38] n38 Kenyasus, [39] n39 Perchoerus, [40] n40 Amphimeryx, [41] n41 Khirtaria, [42] n42 Gobiohyus, [43] n43 Mixtotherium, [44] n44 Lophiomeryx, [45] n45 Homacodon, [46] n46 Xiphodon, [47] n47 Diplobune, [48] n48 Gujaratia, [49] n49 Bunophorus, [50] n50 Dacrytherium, [51] n51 Paroxacron, [52] n52 Entelodon, [53] n53 Indohyus, [54] n54 Merycoidodon, [55] n55 Archaeomeryx, [56] n56 Amphiraghatherium, [57] n57 Hallebune; TREE Strict = (n45,((n48,n49),(n52,(n41,n53))),(n0,(n42,(((n56,n57),(n43,((n47,(n50,n54)),(n46,(n51,(n40,(n44,n55))))))),((n1,(n38,(n37,n39))),(n2,(n3,((n4,(n10,(n7,(n8,n9)))),(n5,(n6,((n22,(n23,(n24,(n25,(n26,n27))))),((n18,n19),(n20,n21),((n14,n15,n16,n17),(n12,n13,(n11,(n28,(n29,n30,n31,(n32,(n33,(n34,(n35,n36)))))))))))))))))))))[% ] [% ] [% setBetweenBits = selected ]; END; Begin MESQUITE; MESQUITESCRIPTVERSION 2; TITLE AUTO; tell ProjectCoordinator; timeSaved 1610015841143; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; setID 0 5271987024600702022; tell It; attachments ; endTell; endTell; getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters; tell It; setID 0 1848080691471385991; mqVersion 361; checksumv 0 3 3788879246 null getNumChars 164 numChars 164 getNumTaxa 58 numTaxa 58 short true bits 2305843009213693967 states 15 sumSquaresStatesOnly 26846.0 sumSquares -5.995191823955604E19 longCompressibleToShort false usingShortMatrix true NumFiles 1 NumMatrices 1; mqVersion; endTell; getWindow; tell It; suppress; setResourcesState true false 141; setPopoutState 300; setExplanationSize 0; setAnnotationSize 0; setFontIncAnnot 0; setFontIncExp 0; setSize 1257 709; setLocation 8 23; setFont SanSerif; setFontSize 10; getToolPalette; tell It; endTell; setActive; desuppress; endTell; getEmployee #mesquite.charMatrices.BasicDataWindowCoord.BasicDataWindowCoord; tell It; showDataWindow #1848080691471385991 #mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindowMaker; tell It; getWindow; tell It; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 0 -72; setLocation 8 23; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.arrow; endTell; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.arrow; colorCells #mesquite.charMatrices.NoColor.NoColor; colorRowNames #mesquite.charMatrices.TaxonGroupColor.TaxonGroupColor; colorColumnNames #mesquite.charMatrices.CharGroupColor.CharGroupColor; colorText #mesquite.charMatrices.NoColor.NoColor; setBackground White; toggleShowNames on; toggleShowTaxonNames on; toggleTight off; toggleThinRows off; toggleShowChanges on; toggleSeparateLines off; toggleShowStates on; toggleReduceCellBorders off; toggleAutoWCharNames on; toggleAutoTaxonNames off; toggleShowDefaultCharNames off; toggleConstrainCW on; toggleBirdsEye off; toggleShowPaleGrid off; toggleShowPaleCellColors off; toggleShowPaleExcluded off; togglePaleInapplicable on; togglePaleMissing off; toggleShowBoldCellText off; toggleAllowAutosize on; toggleColorsPanel off; toggleDiagonal on; setDiagonalHeight 80; toggleLinkedScrolling on; toggleScrollLinkedTables off; endTell; getWindow; tell It; forceAutosize; endTell; hideWindow; getEmployee #mesquite.charMatrices.AlterData.AlterData; tell It; toggleBySubmenus off; endTell; getEmployee #mesquite.charMatrices.ColorByState.ColorByState; tell It; setStateLimit 9; toggleUniformMaximum on; endTell; getEmployee #mesquite.charMatrices.ColorCells.ColorCells; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.categ.StateNamesStrip.StateNamesStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.AnnotPanel.AnnotPanel; tell It; togglePanel off; endTell; getEmployee #mesquite.charMatrices.CharReferenceStrip.CharReferenceStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.QuickKeySelector.QuickKeySelector; tell It; autotabOff; endTell; getEmployee #mesquite.charMatrices.SelSummaryStrip.SelSummaryStrip; tell It; showStrip off; endTell; getEmployee #mesquite.categ.SmallStateNamesEditor.SmallStateNamesEditor; tell It; panelOpen true; endTell; endTell; endTell; endTell; end;