#NEXUS Begin taxa; Dimensions ntax=283; Taxlabels Acanthixalus_sonjae Acanthixalus_spinosus Afrixalus_brachycnemis Afrixalus_delicatus Afrixalus_dorsalis_1 Afrixalus_dorsalis_2 Afrixalus_dorsalis_3 Afrixalus_dorsimaculatus_1 Afrixalus_dorsimaculatus_2 Afrixalus_enseticola Afrixalus_equatorialis Afrixalus_fornasini Afrixalus_fulvovittatus_1 Afrixalus_fulvovittatus_2 Afrixalus_knysae Afrixalus_lacteus Afrixalus_lacustris Afrixalus_laevis Afrixalus_morerei Afrixalus_nigeriensis Afrixalus_osorioi Afrixalus_paradorsalis_1 Afrixalus_paradorsalis_2 Afrixalus_paradorsalis_3 Afrixalus_quadrivittatus_1 Afrixalus_quadrivittatus_2 Afrixalus_spinifrons Afrixalus_stuhlmanni Afrixalus_sylvaticus Afrixalus_vibekensis Afrixalus_vittiger_1 Afrixalus_vittiger_2 Afrixalus_weidholzi Afrixalus_wittei Arthroleptis_adelphus Arthroleptis_affinis Arthroleptis_bioko Arthroleptis_brevipes Arthroleptis_fichika Arthroleptis_francei Arthroleptis_nikeae Arthroleptis_palava Arthroleptis_perreti Arthroleptis_poecilonotus Arthroleptis_reichei Arthroleptis_schubotzi Arthroleptis_stenodactylus Arthroleptis_sylvaticus Arthroleptis_taeniatus Arthroleptis_tanneri Arthroleptis_variabilis Arthroleptis_xenodactyloides Astylosternus_batesi Astylosternus_diadematus Astylosternus_laticephalus Astylosternus_occidentalis Astylosternus_schioetzi Balebreviceps_hillmani Breviceps_adspersus Breviceps_branchi Breviceps_fuscus Breviceps_gibbosus Breviceps_macrops Breviceps_montanus Breviceps_mossambicus Breviceps_namaquensis Callulina_dawida Callulina_hanseni Callulina_kanga Callulina_kisiwamsitu Callulina_kreffti Callulina_laphami Callulina_meteora Callulina_shengena Callulina_sp_Rubeho Callulina_sp_lowland Callulina_stanleyi Cardioglossa_annulata Cardioglossa_elegans Cardioglossa_escalerae Cardioglossa_gracilis Cardioglossa_leucomystax Cardioglossa_manengouba Cardioglossa_melanogaster Cardioglossa_nigromaculata Cardioglossa_occidentalis Cardioglossa_oreas Cardioglossa_pulchra Cardioglossa_trifasciata Chlorolius_koehleri Cryptothylax_greshoffii Hemisus_marmoratus Hemisus_sudanensis Heterixalus_alboguttatus Heterixalus_andrakata Heterixalus_betsileo Heterixalus_boettgeri Heterixalus_carbonei Heterixalus_luteostriatus Heterixalus_madagascariensis Heterixalus_punctatus Heterixalus_rutenbergi Heterixalus_tricolor Heterixalus_variabilis Hyperolius_acuticeps Hyperolius_ademetzi Hyperolius_adspersus Hyperolius_argus Hyperolius_balfouri_1 Hyperolius_balfouri_2 Hyperolius_baumanni Hyperolius_benguellensis Hyperolius_bobirensis Hyperolius_bocagei Hyperolius_bolifambae Hyperolius_burgessi Hyperolius_camerunensis Hyperolius_castaneus Hyperolius_chlorosteus Hyperolius_cinereus Hyperolius_cinnamomeoventris_1 Hyperolius_cinnamomeoventris_2 Hyperolius_cinnamomeoventris_3 Hyperolius_concolor_1 Hyperolius_concolor_2 Hyperolius_concolor_3 Hyperolius_cystocandicans Hyperolius_dartevellei Hyperolius_dintelmanni Hyperolius_discodactylus Hyperolius_endjami_1 Hyperolius_endjami_2 Hyperolius_friedemanni Hyperolius_frontalis Hyperolius_fusciventris_1 Hyperolius_fusciventris_2 Hyperolius_guttulatus_1 Hyperolius_guttulatus_2 Hyperolius_howelli Hyperolius_igbettensis Hyperolius_inyangae Hyperolius_jackie Hyperolius_jacobseni Hyperolius_kivuensis_1 Hyperolius_kivuensis_2 Hyperolius_kivuensis_3 Hyperolius_kuligae Hyperolius_lamottei Hyperolius_langi Hyperolius_lateralis Hyperolius_lupiroensis Hyperolius_mariae Hyperolius_marmoratus_marmoratus Hyperolius_marmoratus_swynnertoni Hyperolius_marmoratus_taeniatus Hyperolius_minitissimus Hyperolius_mitchelli Hyperolius_molleri Hyperolius_montanus Hyperolius_mosaicus Hyperolius_nasutus Hyperolius_nitidulus_1 Hyperolius_nitidulus_2 Hyperolius_occidentalis Hyperolius_ocellatus_1 Hyperolius_ocellatus_2 Hyperolius_parallelus_alborufus Hyperolius_parallelus_marginatus Hyperolius_parallelus_parallelus Hyperolius_parallelus_pyrrhodictyon Hyperolius_pardalis Hyperolius_parkeri Hyperolius_phantasticus Hyperolius_picturatus Hyperolius_pictus Hyperolius_platyceps Hyperolius_poweri Hyperolius_pseudoargus Hyperolius_pusillus Hyperolius_quinquevittatus Hyperolius_raymondii Hyperolius_reesi Hyperolius_riggenbachi Hyperolius_rubrovermiculatus Hyperolius_rwandae Hyperolius_semidiscus Hyperolius_sp_nov_1 Hyperolius_sp_nov_2 Hyperolius_spatzi Hyperolius_spinigularis Hyperolius_steindachneri Hyperolius_substriatus Hyperolius_sylvaticus Hyperolius_tanneri Hyperolius_thomensis Hyperolius_torrentis Hyperolius_tuberculatus Hyperolius_tuberilinguis Hyperolius_veithi Hyperolius_viridiflavus_destefanii Hyperolius_viridiflavus_glandicolor Hyperolius_viridiflavus_ngorongoriensis Hyperolius_viridiflavus_ommatostictus Hyperolius_viridiflavus_pantherinus Hyperolius_viridiflavus_pitmani Hyperolius_viridiflavus_sp Hyperolius_viridiflavus_variablilis Hyperolius_viridis Hyperolius_zonatus Kassina_arboricola Kassina_cassinoides Kassina_cochranae Kassina_decorata Kassina_fusca Kassina_kuvangensis Kassina_lamottei Kassina_maculosa Kassina_schioetzi Kassina_senegalensis Leptodactylodon_axillaris Leptodactylodon_bicolor Leptodactylodon_boulengeri Leptodactylodon_bueanus Leptodactylodon_erythrogaster Leptodactylodon_mertensi Leptodactylodon_ornatus Leptodactylodon_ovatus Leptodactylodon_perreti Leptodactylodon_polyacanthus Leptodactylodon_ventrimarmoratus Leptopelis_argenteus Leptopelis_aubryi Leptopelis_aubryioides Leptopelis_bocagii Leptopelis_boulengeri Leptopelis_brevirostris Leptopelis_calcaratus Leptopelis_fiziensis Leptopelis_flavomaculatus Leptopelis_gramineus Leptopelis_karissimbensis Leptopelis_kivuensis Leptopelis_macrotis Leptopelis_millsoni Leptopelis_modestus Leptopelis_natalensis Leptopelis_nordequatorialis Leptopelis_notatus Leptopelis_ocellatus Leptopelis_palmatus Leptopelis_parkeri Leptopelis_rufus Leptopelis_spiritusnoctis Leptopelis_susanae Leptopelis_vannutellii Leptopelis_vermiculatus Leptopelis_viridis Leptopelis_yaldeni Morerella_cyanophthalma Nyctibates_corrugatus Opisthothylax_immaculatus Paracassina_kounhiensis Paracassina_obscura Phlyctimantis_boulengeri Phlyctimantis_keithae Phlyctimantis_leonardi Phlyctimantis_maculata Phlyctimantis_verrucosus Phrynomantis_microps Probreviceps_durirostris Probreviceps_loveridgei Probreviceps_macrodactylus Probreviceps_rungwensis Probreviceps_sp_Kigogo Probreviceps_sp_Nguru Probreviceps_sp_Rubeho Probreviceps_sp_Udzungwa Probreviceps_uluguruensis Scotobleps_gabonicus Semnodactylus_wealii Spelaeophryne_methneri Tachycnemis_seychellensis Trichobatrachus_robustus ; End; Begin trees; Translate 1 Acanthixalus_sonjae, 2 Acanthixalus_spinosus, 3 Afrixalus_brachycnemis, 4 Afrixalus_delicatus, 5 Afrixalus_dorsalis_1, 6 Afrixalus_dorsalis_2, 7 Afrixalus_dorsalis_3, 8 Afrixalus_dorsimaculatus_1, 9 Afrixalus_dorsimaculatus_2, 10 Afrixalus_enseticola, 11 Afrixalus_equatorialis, 12 Afrixalus_fornasini, 13 Afrixalus_fulvovittatus_1, 14 Afrixalus_fulvovittatus_2, 15 Afrixalus_knysae, 16 Afrixalus_lacteus, 17 Afrixalus_lacustris, 18 Afrixalus_laevis, 19 Afrixalus_morerei, 20 Afrixalus_nigeriensis, 21 Afrixalus_osorioi, 22 Afrixalus_paradorsalis_1, 23 Afrixalus_paradorsalis_2, 24 Afrixalus_paradorsalis_3, 25 Afrixalus_quadrivittatus_1, 26 Afrixalus_quadrivittatus_2, 27 Afrixalus_spinifrons, 28 Afrixalus_stuhlmanni, 29 Afrixalus_sylvaticus, 30 Afrixalus_vibekensis, 31 Afrixalus_vittiger_1, 32 Afrixalus_vittiger_2, 33 Afrixalus_weidholzi, 34 Afrixalus_wittei, 35 Arthroleptis_adelphus, 36 Arthroleptis_affinis, 37 Arthroleptis_bioko, 38 Arthroleptis_brevipes, 39 Arthroleptis_fichika, 40 Arthroleptis_francei, 41 Arthroleptis_nikeae, 42 Arthroleptis_palava, 43 Arthroleptis_perreti, 44 Arthroleptis_poecilonotus, 45 Arthroleptis_reichei, 46 Arthroleptis_schubotzi, 47 Arthroleptis_stenodactylus, 48 Arthroleptis_sylvaticus, 49 Arthroleptis_taeniatus, 50 Arthroleptis_tanneri, 51 Arthroleptis_variabilis, 52 Arthroleptis_xenodactyloides, 53 Astylosternus_batesi, 54 Astylosternus_diadematus, 55 Astylosternus_laticephalus, 56 Astylosternus_occidentalis, 57 Astylosternus_schioetzi, 58 Balebreviceps_hillmani, 59 Breviceps_adspersus, 60 Breviceps_branchi, 61 Breviceps_fuscus, 62 Breviceps_gibbosus, 63 Breviceps_macrops, 64 Breviceps_montanus, 65 Breviceps_mossambicus, 66 Breviceps_namaquensis, 67 Callulina_dawida, 68 Callulina_hanseni, 69 Callulina_kanga, 70 Callulina_kisiwamsitu, 71 Callulina_kreffti, 72 Callulina_laphami, 73 Callulina_meteora, 74 Callulina_shengena, 75 Callulina_sp_Rubeho, 76 Callulina_sp_lowland, 77 Callulina_stanleyi, 78 Cardioglossa_annulata, 79 Cardioglossa_elegans, 80 Cardioglossa_escalerae, 81 Cardioglossa_gracilis, 82 Cardioglossa_leucomystax, 83 Cardioglossa_manengouba, 84 Cardioglossa_melanogaster, 85 Cardioglossa_nigromaculata, 86 Cardioglossa_occidentalis, 87 Cardioglossa_oreas, 88 Cardioglossa_pulchra, 89 Cardioglossa_trifasciata, 90 Chlorolius_koehleri, 91 Cryptothylax_greshoffii, 92 Hemisus_marmoratus, 93 Hemisus_sudanensis, 94 Heterixalus_alboguttatus, 95 Heterixalus_andrakata, 96 Heterixalus_betsileo, 97 Heterixalus_boettgeri, 98 Heterixalus_carbonei, 99 Heterixalus_luteostriatus, 100 Heterixalus_madagascariensis, 101 Heterixalus_punctatus, 102 Heterixalus_rutenbergi, 103 Heterixalus_tricolor, 104 Heterixalus_variabilis, 105 Hyperolius_acuticeps, 106 Hyperolius_ademetzi, 107 Hyperolius_adspersus, 108 Hyperolius_argus, 109 Hyperolius_balfouri_1, 110 Hyperolius_balfouri_2, 111 Hyperolius_baumanni, 112 Hyperolius_benguellensis, 113 Hyperolius_bobirensis, 114 Hyperolius_bocagei, 115 Hyperolius_bolifambae, 116 Hyperolius_burgessi, 117 Hyperolius_camerunensis, 118 Hyperolius_castaneus, 119 Hyperolius_chlorosteus, 120 Hyperolius_cinereus, 121 Hyperolius_cinnamomeoventris_1, 122 Hyperolius_cinnamomeoventris_2, 123 Hyperolius_cinnamomeoventris_3, 124 Hyperolius_concolor_1, 125 Hyperolius_concolor_2, 126 Hyperolius_concolor_3, 127 Hyperolius_cystocandicans, 128 Hyperolius_dartevellei, 129 Hyperolius_dintelmanni, 130 Hyperolius_discodactylus, 131 Hyperolius_endjami_1, 132 Hyperolius_endjami_2, 133 Hyperolius_friedemanni, 134 Hyperolius_frontalis, 135 Hyperolius_fusciventris_1, 136 Hyperolius_fusciventris_2, 137 Hyperolius_guttulatus_1, 138 Hyperolius_guttulatus_2, 139 Hyperolius_howelli, 140 Hyperolius_igbettensis, 141 Hyperolius_inyangae, 142 Hyperolius_jackie, 143 Hyperolius_jacobseni, 144 Hyperolius_kivuensis_1, 145 Hyperolius_kivuensis_2, 146 Hyperolius_kivuensis_3, 147 Hyperolius_kuligae, 148 Hyperolius_lamottei, 149 Hyperolius_langi, 150 Hyperolius_lateralis, 151 Hyperolius_lupiroensis, 152 Hyperolius_mariae, 153 Hyperolius_marmoratus_marmoratus, 154 Hyperolius_marmoratus_swynnertoni, 155 Hyperolius_marmoratus_taeniatus, 156 Hyperolius_minitissimus, 157 Hyperolius_mitchelli, 158 Hyperolius_molleri, 159 Hyperolius_montanus, 160 Hyperolius_mosaicus, 161 Hyperolius_nasutus, 162 Hyperolius_nitidulus_1, 163 Hyperolius_nitidulus_2, 164 Hyperolius_occidentalis, 165 Hyperolius_ocellatus_1, 166 Hyperolius_ocellatus_2, 167 Hyperolius_parallelus_alborufus, 168 Hyperolius_parallelus_marginatus, 169 Hyperolius_parallelus_parallelus, 170 Hyperolius_parallelus_pyrrhodictyon, 171 Hyperolius_pardalis, 172 Hyperolius_parkeri, 173 Hyperolius_phantasticus, 174 Hyperolius_picturatus, 175 Hyperolius_pictus, 176 Hyperolius_platyceps, 177 Hyperolius_poweri, 178 Hyperolius_pseudoargus, 179 Hyperolius_pusillus, 180 Hyperolius_quinquevittatus, 181 Hyperolius_raymondii, 182 Hyperolius_reesi, 183 Hyperolius_riggenbachi, 184 Hyperolius_rubrovermiculatus, 185 Hyperolius_rwandae, 186 Hyperolius_semidiscus, 187 Hyperolius_sp_nov_1, 188 Hyperolius_sp_nov_2, 189 Hyperolius_spatzi, 190 Hyperolius_spinigularis, 191 Hyperolius_steindachneri, 192 Hyperolius_substriatus, 193 Hyperolius_sylvaticus, 194 Hyperolius_tanneri, 195 Hyperolius_thomensis, 196 Hyperolius_torrentis, 197 Hyperolius_tuberculatus, 198 Hyperolius_tuberilinguis, 199 Hyperolius_veithi, 200 Hyperolius_viridiflavus_destefanii, 201 Hyperolius_viridiflavus_glandicolor, 202 Hyperolius_viridiflavus_ngorongoriensis, 203 Hyperolius_viridiflavus_ommatostictus, 204 Hyperolius_viridiflavus_pantherinus, 205 Hyperolius_viridiflavus_pitmani, 206 Hyperolius_viridiflavus_sp, 207 Hyperolius_viridiflavus_variablilis, 208 Hyperolius_viridis, 209 Hyperolius_zonatus, 210 Kassina_arboricola, 211 Kassina_cassinoides, 212 Kassina_cochranae, 213 Kassina_decorata, 214 Kassina_fusca, 215 Kassina_kuvangensis, 216 Kassina_lamottei, 217 Kassina_maculosa, 218 Kassina_schioetzi, 219 Kassina_senegalensis, 220 Leptodactylodon_axillaris, 221 Leptodactylodon_bicolor, 222 Leptodactylodon_boulengeri, 223 Leptodactylodon_bueanus, 224 Leptodactylodon_erythrogaster, 225 Leptodactylodon_mertensi, 226 Leptodactylodon_ornatus, 227 Leptodactylodon_ovatus, 228 Leptodactylodon_perreti, 229 Leptodactylodon_polyacanthus, 230 Leptodactylodon_ventrimarmoratus, 231 Leptopelis_argenteus, 232 Leptopelis_aubryi, 233 Leptopelis_aubryioides, 234 Leptopelis_bocagii, 235 Leptopelis_boulengeri, 236 Leptopelis_brevirostris, 237 Leptopelis_calcaratus, 238 Leptopelis_fiziensis, 239 Leptopelis_flavomaculatus, 240 Leptopelis_gramineus, 241 Leptopelis_karissimbensis, 242 Leptopelis_kivuensis, 243 Leptopelis_macrotis, 244 Leptopelis_millsoni, 245 Leptopelis_modestus, 246 Leptopelis_natalensis, 247 Leptopelis_nordequatorialis, 248 Leptopelis_notatus, 249 Leptopelis_ocellatus, 250 Leptopelis_palmatus, 251 Leptopelis_parkeri, 252 Leptopelis_rufus, 253 Leptopelis_spiritusnoctis, 254 Leptopelis_susanae, 255 Leptopelis_vannutellii, 256 Leptopelis_vermiculatus, 257 Leptopelis_viridis, 258 Leptopelis_yaldeni, 259 Morerella_cyanophthalma, 260 Nyctibates_corrugatus, 261 Opisthothylax_immaculatus, 262 Paracassina_kounhiensis, 263 Paracassina_obscura, 264 Phlyctimantis_boulengeri, 265 Phlyctimantis_keithae, 266 Phlyctimantis_leonardi, 267 Phlyctimantis_maculata, 268 Phlyctimantis_verrucosus, 269 Phrynomantis_microps, 270 Probreviceps_durirostris, 271 Probreviceps_loveridgei, 272 Probreviceps_macrodactylus, 273 Probreviceps_rungwensis, 274 Probreviceps_sp_Kigogo, 275 Probreviceps_sp_Nguru, 276 Probreviceps_sp_Rubeho, 277 Probreviceps_sp_Udzungwa, 278 Probreviceps_uluguruensis, 279 Scotobleps_gabonicus, 280 Semnodactylus_wealii, 281 Spelaeophryne_methneri, 282 Tachycnemis_seychellensis, 283 Trichobatrachus_robustus ; tree TREE1 = [&R] 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