NEXUS
[File created by TreeBASE:  7/26/07  21:01:39]
[Matrix accession#: M1346]

BEGIN TAXA;
	TITLE M1346;
	DIMENSIONS  NTAX=26;
	TAXLABELS
		Trillium_undulatum	Trillium_rugelii	Trillium_pusillum	Trillium_ovatum	Trillium_grandiflorum	Trillium_govanianum	Trillium_erectum	Trillium_camschatcense	Trillium_sessile	Trillium_reliquum	Trillium_recurvatum	Trillium_petiolatum	Trillium_maculatum	Trillium_lancifolium	Trillium_discolor	Trillium_decipiens	Trillium_chloropetalum	Paris_verticillata	Paris_tetraphylla	Paris_incompleta	Kinugasa_japonica	Daiswa_violacea	Daiswa_thibetica	Daiswa_polyphylla	Daiswa_fargesii	Trillium_rivale
	;
END;

BEGIN CHARACTERS;
	TITLE M1346;
	DIMENSIONS NCHAR=110;
	FORMAT SYMBOLS = " 1 2 3 4 5 6 7 8 9 A B C D E F G H I J K L M N O P Q " MISSING = ? GAP = -  ;
	CHARSTATELABELS
		1 hybrid_status / yes no, 2 flowering_time / February March April May June July August September, 3 chromosome_number / '2n=10' '2n=15' '2n=20' '2n=30' '2n=40' '2n=60' '2n=14' '2n=12' '2n=24', 4 geographical_area / east_Asia western_North_America eastern_North_America Europe, 5 genus / outgroup Trillium Paris, 6 subgenus / outgroup Trillium Phyllantherum Paris Kinugasa Daiswa, 7 section / outgroup grandiflorum erectum catesbaei recurvatum sessile maculatum paris axiparis kinugasa daiswa dunnaniana marmorata fargesiana thibetica, 8 Rafinesque / outgroup Paris Daiswa Phyllantherum Trillium Delostylis, 9 PLANT_type / herbs vines, 10 foliaceousness / scapose leafy subscapose, 11 plant_sexuality / bisexual dioecious polygamous, 12 INFLORESCENSE_type / solitary simple_umbel raceme panicle_of_racemes panicle spike cyme, 13 FLOWERS_Num / one more_than_one, 14 composition / tepals 'sepals + petals', 15 PED._vertical_posture / erect above_the_leaves horizontal below_the_leaves none, 16 root_type / rhizome bulb tuber fibrous, 17 rhizome_size / very_thin slender thick very_thick not_applicable, 18 STEM_habit / erect partially_decumbent decumbent twining, 19 vestiture / glabrous puberulent pubescent pilose papillose scabrous striate floccose, 20 stem_color / green 'yellowish-green' reddish_purple greenish_purple purplish brownish_green reddish green_over_purple purple_tinged 'green over brown-red', 21 stem_distribution / throughout geographic suffused, 22 LEAF_number / two three four five six seven eight nine ten eleven_and_up, 23 leaf_location / terminal cauline basal reduced_cauline, 24 leaf_arrangement / whorled alternate opposite spirally_inserted, 25 Num_whorls / one more_than_two none, 26 leaf_attachment / petiolate subsessile sessile, 27 'leaf$ widest point' / 'ovate (.25)' 'elliptic-ovate (.375)' 'elliptic-rhombic-oblong (.5)' 'elliptic-obovate (.675)' 'obovate (.75)', 28 'leaf$ width (LxW)' / linear narrow average broad very_broad depressed_or_transverse, 29 'leaf$ sides' / 'elliptic (curved)' 'oblong-elliptic' 'oblong (parallel)' 'oblong-rhombic' 'rhombic (straight)' 'rhombic-elliptic', 30 leaf_margin / entire undulate serrulate, 31 leaf_distribution / throughout mottled geographic suffused, 32 leaf_color / green more_or_less_purple_mottled green_w_maroon_hue small_brownish_spots white_variegation_on_veins mottled mottled_w_dk_green spots_of_lt_green multiple_shades_of_green shades_of_green_beside_pale_mid dk_green_spots_on_mottled_backg dk_green_between_veins_w_pale_m, 33 leaf_lower_surface / pubescent_on_veins scabrous_on_veins glossy dull purple purple_nerves glabrous pubescent, 34 apex_type / entire sinusus 'mid-rib, midvein or vein exten', 35 apices / acuminate acute obtuse rounded emarginate 'sub-acute' cuspidate blunt attenuate, 36 base_type / entire with_sinusus w_specialized_attachments, 37 bases / cuneate sheathing obtuse rounded attenuate hastate cordate, 38 leaf_texture / herbaceous membranous papery rugose coriaceous fleshy, 39 Num_main_nerves / many three five seven nine eleven, 40 BRACTS / absent present, 41 SEPAL_form / normal petaloid, 42 sepal_fusion / separate fused, 43 sepal_number / three four five six seven eight nine ten, 44 'sepal$ widest point' / 'ovate (.25)' 'elliptic-ovate (.375)' 'elliptic-rhombic-oblong (.5)' 'elliptic-obovate (.625)' 'obovate (.75)', 45 'sepal$ width (LxW)' / narrow average broad, 46 'sepal$ sides' / 'elliptic (curved)' 'oblong-elliptic' 'oblong (parallel)', 47 sepal_duration / persistent deciduous, 48 sepal_texture / herbaceous membranous, 49 sepal_color / reddish_purple yellow purple purplish_green red green reddish_green yellowish_green white white_w_green_veins greenish_w_red_or_purple_veins 'greenish, purple basally' 'greenish, reddish-purple basall' greensih_brown 'greenish, w red veins' greenish_w_reddish_purple_margi greenish_w_purple_margins green_w_white_veins, 50 sepal_distribution / throughout suffused marginal mottled geographic, 51 sepal_apices / acuminate acute obtuse rounded 'acuminate-rounded' 'sub-acute' cuspidate emarginate caudate blunt attenuate, 52 sepal_apex_type / entire 'mid-rib, midvein or vei', 53 PETAL_form / normal foliaceous staminoid none, 54 petal_presence / present absent, 55 petal_number / zero three four five six seven eight, 56 petal_fusion / separate fused, 57 'petal$ widest point' / 'ovate (.25)' 'elliptic-ovate (.375)' 'elliptic-rhombic-oblong (.5)' 'elliptic-obovate (.625)' 'obovate (.75)', 58 'petal$ width (LxW)' / filiform linear narrow average broad very_broad, 59 'petal$ sides' / 'elliptic (curved)' 'oblong-elliptic' 'oblong (parallel)' 'rhombic-elliptic', 60 petal_duration / persistent deciduous, 61 petal_color / white yellow cream pink red purple bronze green 'reddish-purple' 'greenish-yellow' 'purplish-black' brownish_purple 'greenish-purple' purple_w_greenish_claw yellow_w_greenish_claw green_w_purple_claw yellow_w_purple_claw 'yellow-green w purple claw' white_spotted_w_rose white_w_purple_basally white_w_basal_red_V olive purplish_green_w_purple_cl grey_purple_w_purple_veins_and, 62 petal_distribution / throughout spotted geographic, 63 anthocyanins / yes no fading_to_pink, 64 petal_transverse_posture / incurved plane recurved undulate outcurved, 65 petal_vertical_orientation / erect divergent horizontal divergent declined, 66 petal_longitudinal_posture / straight twisted, 67 petal_apices / acuminate acute obtuse rounded blunt 'sub-acute' cuspidate apiculate emarginate mucronate, 68 petal_apex_type / entire 'mid-rib, midvein or vein extens', 69 STAMEN_form / normal 'petaloid-abortive', 70 stamen_number / zero 1X 2X_sepals 3X 4X, 71 stamen_fusion / free adnate_to_petal adnate_to_sepal adnate_to_tepal, 72 stamen_transverse_posture / straight incurved slightly_recurved, 73 stamen_vertical_posture / erect spreading, 74 ANTHERS_dehiscence / introrse lateral extrorse, 75 POLLEN_shape / spherical ellipsoid irregular, 76 pollen_aperture / omniaperturate monosulcate, 77 pollen_ornamentation / granulate echinate corrugate verrucate spinulate clavate foveolate reticulate psilate gemmate rugulate, 78 CONNECTIVE_prolongation / not_prolonged acute truncate dilated round emarginate obtuse, 79 FILAMENT_color / green purple whitish_purple reddish_purple whitish_green 'red-purple-brown' pinkish_white greenish_yellow white pink brown yellow, 80 pollen_color / yellow orange olive brownish 'olive-orange' purple greyish_purple pink yellowish_pale_purple green maroon 'orange-yellow', 81 connective_color / green pink purple pinkish_purple reddish_purple brown whitish_green white 'red-purple-brown', 82 stamen_distribution / throughout pollen_different all_different, 83 PISTIL_form / normal abortive, 84 OVARY_position / inferior superior, 85 Num_locules / one three four_to_ten, 86 placentation / axilary parietal combination, 87 ovary_shape / conical 'angular-ovoid' ovoid ellipsoid obovoid 'flask-shaped' subglobose 'oval-globose' fusiform 'ovate-conical', 88 'ovary X-section shape' / round ridged angled winged, 89 ovary_ribs / zero two three four five six seven, 90 ovary_apex / truncate crowned attenuate obtuse, 91 STYLE_presence / present absent, 92 STIGMA_number / three four five six eight ten seven one, 93 shape / linear 'linear-subulate' subulate button, 94 'vertical posture (orie' / divergent erect, 95 stigma_transverse_posture / incurved straight recurved, 96 stigma_duration / deciduous persistent, 97 stigma_size / very_thin thin average thick, 98 ovary_color / yellowish_white green purple pink cream_yellow white 'reddish-purple' purplish_brown yellow_green 'red-purple-brown over green-red' green_over_purple purple_over_green purple_over_yellow purple_over_white pink_over_white green_w_purple_disk green_w_yellow_disk 'green w white disk-green w whit' yellow_w_black_disk 'red-purple-brown' black, 99 ovary_distribution / throughout spotted striped geographic, 100 stigma_color / green purple yellowish_green creamy_white yellow yellowish_white pink black 'green and purple-red' 'brownish-violet' 'red-purple-brown' brown 'purple out, green in' 'brown-purple out, yellow in' 'purple out, yellow in' 'white, basally pink' 'green out, yellow in' 'pinkish-orange' red, 101 stigma_distribution / throughout banded geographic, 102 PISTIL_distribution / throughout geographic_by_part all_different, 103 FRUIT_type / berry capsule fleshy_capsule, 104 fruit_dehisence / indehiscent basally_absicent septicidal irregular loculicidal dehiscent decay, 105 fruit_shape / conical 'angular-ovoid' ovoid globose obovoid elliptic, 106 fruit_color / green maroon greenish_yellow rusty_red greenish_white red black 'bluish-black' brown purple green_w_purple_dots 'purplish-green' 'greenish-brown' green_w_brown_ribs 'black-purple' white pink, 107 fruit_color_distribution / throughout mottled spotted striped geographic, 108 SEED_arils / incomplete absent complete, 109 endosperm_development / helobial nuclear, 	110 cotyledon_shape / strap_like leaf_like
	;
	MATRIX
		Trillium_undulatum           2(345)13222511111221311(13)121111131(12)117111413111111111H3111121(12)(34)(12)1L31421211311(12)3114277221212313(34)2121321F471311361111
		Trillium_rugelii             2(345)132235111112413116121113345111711111(23)111111111D51111211(45)11K31321?11311221111273212(12)(12)A46121?13223121131321111
		Trillium_pusillum            2(234)1322(24)611111221211(15)(13)2111(13)(12)(23)(12)1(14)(13)71314121111(12)1(12)11(6H)(13)(23)111211311113421(23)113111(12)111141(48)31212313311?13219131111311111
		Trillium_ovatum              2(1234)12222511111221311112111324611171(12)1?1(23)1111(12)1(12)1181211121(12)(34)(12)1(14)1(13)(25)21(23)11313221151517212123363212(12)3211161131311111
		Trillium_grandiflorum        2(345)13222511111221311112111(23)2(34)(46)11171(12)111311111111161(12)111215(34)11(149)1(13)(45)21(37)11313111151918212138263212(12)32121111(13)1411111
		Trillium_govanianum          2(345)312225111112212114121111(12)(23)(12)111711(12)(47)221111(23)1(23)11(34)1111121(23)(23)(23)1(689)11(23)(234)1111311132261213212??3?32211(12)32131211114A12??
		Trillium_erectum             2(345)1322351111122131111211133451117111113111111111611111211311(13469)1(12)(25)21(12)11313121111(9A)(17)??1212(34)46321?132(34)(378)1211113A1111
		Trillium_camschatcense       2(456)112235111112213111121113(23)3611171(12)131(23)1111(23)2(23)1161(23)11121(123)4(123)1112521(123)1131111111511121212(14)26321?1322(139C)(124)51311(34)(AB)(13)111
		Trillium_sessile             2(234)132364111112513111121113(13)(34)11(12)(147)71(124)141311111(12)111(6CH)(135)611121(13)(34)11(67A)11111(26)1131111111(25)211312??(37)(34)632132?2(23)81F3311(34)C1111
		Trillium_reliquum            2(23)13236411111251?(23)11121113(13)(34)112(9C)21(23)1?12111111111(3C)(25)(24)11121(35)311(26D)111122113121111?2213112??3463211132?31F33113?1111
		Trillium_recurvatum          2(234)132354111112512111121111(13)(234)112771(12)131311111(12)11161211121(14)(34)(14)1(29C)11111(23)1131211112526311213(24)46321113222121211(24)11111
		Trillium_petiolatum          2(34)122374111112513111121111(13)(345)111171(34)14141111(2345)(12)(12)1161211121(123)3(123)1(68D)11(15)(12)121131112113(12)2(123)9112123(34)62212132(23)21212(12)13?1111
		Trillium_maculatum           2(123)132374111112513111121113(123)(234)112671(1238)1(45)1311111111131(46)11121(45)(34)(12)1(26H)(13)11(12)2(234)1131211111(35)2(14)3212133(12)61213212481B11113C1111
		Trillium_lancifolium         2(1234)132354111112512111121113(12)2112A712111311111111161211121(345)331(6GN)(13)11112113121111(12)52?3?12??(24)463211132(23)A4E3311211111
		Trillium_discolor            2(34)1323641111125131(13)1121113(12345)3112B71(12)131311111(12)111612111215(34)11F32(15)12(18A)(12)13121(12)111(25)6(12)9212123363213(12)(23)22(2A)(34)B1311(34)?1111
		Trillium_decipiens           2(123)13236411111251311112111312112A212141311111(12)111612111215(34)11(6AD)11111(234)1131(12)1211?(25)619212??4263211132(23)2151211611111
		Trillium_chloropetalum       2(12)12237411111251?11112111(23)1(345)11(12)(14)71(348)1?2?11111111161(234)111215(34)(12)1(1567AD)1(12)111(2345)1131111111(57)2(14)3212??3364212(12)2243121111311111
		Paris_verticillata           2(45)1134821111122111111(567)1112(135)(23)1111711111?111(23)11111611111(34)13131(2A)12351?1131??22272???11231(34)1?31(23)?132(12)81212114F1222
		Paris_tetraphylla            2(4567)113482111112211111131113(234)(234)(126)11171211121112(123)1(123)116111421-------------131??222A1???1123131?412???2??1?1111481222
		Paris_incompleta             2(2345)1(14)34821111122111111(56789A)1111(25)2(12)1114111112111(123)1(12)11161?1421-------------131??222?(12)???112314??41(123)???2231?1111?81222
		Kinugasa_japonica            2(567)5135A21111122141111(6789)111(23)(45)(23)11117111132121(45678)(12)1(12)12(9A)11111(17)15(12)31112?1131131?1322A15172123143?21(457)31324(HI)441311?A12?2
		Daiswa_violacea              2(34)1136?31111122131111(56789)1111121135511111?111(12)1(12)111I51111(23)11131E31??1?1131??222?1C??112124??211?13242121(13)3(56)?61322
		Daiswa_thibetica             231136F31111122131711(678)111(123)1211117111122111311111611111(234)15131?12?(23)1?1131??22282???1121283?(12)13???23(GH)4(257)133(56)?81122
		Daiswa_polyphylla            2(4567)(13)136B31111122131111(5678)1111(123)(23)(123)1117111112111(234)(12)1(12)11611111(345)15131(2A)12??1?1131??22272?1?1121233?21(23)1??24515113(56)?A1322
		Daiswa_fargesii              2(2345)1136E311111221(23)1111(345)11111(34)1111711(12)(47)2(34)111(234)1111(12)811111(345)13211(AB)1(12)?11?1131??2228(46)B?63121213?213???24?1?113(56)(34)?1322
		Trillium_rivale              2(234)1222251111122131151211111(23)111171(12)(12)(47)52111121211617211212441(14J)(12)1521(12)113111311?111121213313311213226141131411112
	;
END;

BEGIN ASSUMPTIONS;
CHARSET non_morf = 1 - 8 ; 
 CHARSET insufficient = 37 64 - 67 72 77 79 - 81 85 - 86 89 - 90 93 - 94 100 105 ; 
 CHARSET subjective = 17 88 97 ; 
 CHARSET numbers = 22 43 55 70 92 ; 
 CHARSET color = 20 32 49 61 79 - 81 98 100 106 ; 
 CHARSET problematic = 33 90 103 - 104 ; 
 TAXSET OutGroup = 1 ; 
 TAXSET Paris = 2 - 9 ; 
 TAXSET sessile = 10 - 26 ; 
 TAXSET pedicellate = 19 - 26 ; 
 OPTIONS DEFTYPE = unord PolyTcount = MINSTEPS ; 
 EXSET * del_non_morf = 1 - 8 ; 
 EXSET del_insufficient = 37 64 - 67 72 77 79 - 81 85 - 86 89 - 90 93 - 94 100 105 ; 
 EXSET del_subjective = 17 88 97 ; 
 EXSET del_numbers = 22 43 55 70 92 ; 
 EXSET del_color = 20 32 49 61 79 - 81 98 100 106 ; 
 EXSET del_problematic = 33 90 103 - 104 ; 
 END;