#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 19, 2025; 9:53 GMT TreeBASE (cc) 1994-2008 Study reference: Sundue M.A., Islam M., & Ranker T. 2010. Systematics of grammitid ferns (Polypodiaceae): using morphology and plastid sequence data to resolve the circumscriptions of Melpomene and the polyphyletic genera Lellingeria and Terpsichore. Systematic Botany, 35. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S10671] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=177; TAXLABELS Calymmodon_gracilis Calymmodon_luerssenianus Campyloneurum_angustifolium Ceradenia_aulaeifolia Ceradenia_aulaeifolia_2 Ceradenia_fucoides Ceradenia_intonsa Ceradenia_jungermannioides Ceradenia_kalbreyeri_Colombia Ceradenia_kalbreyeri_Costa_Rica Ceradenia_pilipes_Costa_Rica Ceradenia_pilipes_Ecuador Ceradenia_sp._aff._kalbreyeri Ceradenia_sp._ined._1 Ceradenia_sp._ined._1_2 Ceradenia_sp._moran_7520 Ceradenia_spixiana Chrysogrammitis_musgraveana Cochlidium_punctatum Cochlidium_rostratum Cochlidium_seminudum Ctenopteris_heterophylla Ctenopteris_heterophylla_2 Ctenopteris_lasiostipes Ctenopteris_nutans Ctenopteris_repandula Ctenopteris_rhodocarpa Enterosora_enterosoroides Enterosora_percrassa Enterosora_percrassa_2 Enterosora_trichosora Enterosora_trifurcata_Costa_Rica Enterosora_trifurcata_Ecuador Grammitis_billardieri Grammitis_ciliata Grammitis_deplanchei Grammitis_forbesiana Grammitis_knutsfordiana Grammitis_melanoloma Grammitis_parva Grammitis_poeppigiana Grammitis_poeppigiana_2 Grammitis_pseudociliata Lellingeria_apiculata_Brazil Lellingeria_apiculata_Costa_Rica Lellingeria_bishopii Lellingeria_hellwigii Lellingeria_humilis Lellingeria_limula Lellingeria_limula_2 Lellingeria_limula_3 Lellingeria_major Lellingeria_major_2 Lellingeria_major_3 Lellingeria_myosuroides Lellingeria_myosuroides_2 Lellingeria_prionodes Lellingeria_pseudomitchellae Lellingeria_saffordii Lellingeria_schenckei Lellingeria_simacensis Lellingeria_subimpressa Lellingeria_subsessilis_Costa_Rica Lellingeria_subsessilis_Ecuador Lellingeria_suprasculpta Lellingeria_suspensa Lellingeria_suspensa_2 Melpomene_anazalea_ined. Melpomene_firma Melpomene_flabelliformis Melpomene_moniliformis Melpomene_occidentalis Melpomene_occidentalis_2 Melpomene_personata_ined. Melpomene_pseudonutans Melpomene_vernicosa Microgramma_percussa Microgramma_squamulosa Micropolypodium_achilleifolium Micropolypodium_hyalinum Micropolypodium_taenifolium Micropolypodium_zurquinum Oreogrammitis_hookeri Pecluma_alfredii Polypodium_glycyrrhiza Prosaptia_alata Prosaptia_contigua Prosaptia_obliquata Prosaptia_pubipes Scleroglossum_sulcatum Scleroglossum_sulcatum_2 Terpsichore_aff._subtilis_inded. Terpsichore_alsopteris Terpsichore_amphidasyon_2 Terpsichore_amphidasyon_3 Terpsichore_amphidasyon_Moran_7646 Terpsichore_anfractuosa_Bolivia Terpsichore_anfractuosa_Bolivia_2 Terpsichore_anfractuosa_Costa_Rica Terpsichore_anfractuosa_Ecuador Terpsichore_anfractuosa_Ecuador_2 Terpsichore_anfractuosa_Mexico Terpsichore_anfractuosa_Mexico_2 Terpsichore_anfractuosa_Peru Terpsichore_anfractuosa_Peru_2 Terpsichore_angustipes Terpsichore_angustipes_2 Terpsichore_athyrioides Terpsichore_attenuatissima Terpsichore_attenuatissima_2 Terpsichore_attenuatissima_3 Terpsichore_clathrata Terpsichore_clavigera_ined. Terpsichore_colombiense_ined. Terpsichore_colombiense_ined._2 Terpsichore_cretata Terpsichore_cuencana Terpsichore_cultrata Terpsichore_davidsmithii Terpsichore_davidsmithii_2 Terpsichore_davidsmithii_3 Terpsichore_davidsmithii_4 Terpsichore_davidsmithii_5 Terpsichore_delicatula Terpsichore_dilatata Terpsichore_dilatata_2 Terpsichore_hanekeana Terpsichore_lanigera Terpsichore_lehmanniana Terpsichore_leucosticta Terpsichore_leucosticta_2 Terpsichore_loja_ined. Terpsichore_loja_ined._2 Terpsichore_loja_ined._3 Terpsichore_longicaulis Terpsichore_longipilosa_ined. Terpsichore_longipilosa_ined._2 Terpsichore_longisetosa Terpsichore_mollissima Terpsichore_nana Terpsichore_pacifica Terpsichore_pichinchae Terpsichore_pichinchense Terpsichore_pichinchense_2 Terpsichore_pichinchense_3 Terpsichore_praeceps Terpsichore_semihirsuta Terpsichore_semihirsuta_Bolivia Terpsichore_semihirsuta_Bolivia_2 Terpsichore_semihirsuta_Colombia Terpsichore_semihirsuta_Colombia_Choco Terpsichore_semihirsuta_Ecuador Terpsichore_semihirsuta_Peru Terpsichore_semihirsuta_ined._2_Costa_Rica Terpsichore_semihirsuta_ined._3_Mexico Terpsichore_senilis Terpsichore_smithii Terpsichore_sp._Sundue_1160 Terpsichore_sp._Sundue_1167 Terpsichore_sp._aff._alsopteris_ined._1_Bolivia Terpsichore_sp._aff._alsopteris_ined._1_Bolivia_2 Terpsichore_sp._aff._alsopteris_ined._1_Bolivia_3 Terpsichore_sp._aff._alsopteris_ined._1_Bolivia_4 Terpsichore_sp._aff._alsopteris_ined._2_Costa_Rica Terpsichore_sp._aff._taxifolia_ined._Colombia Terpsichore_subscabra Terpsichore_subscabra_2 Terpsichore_subtilis Terpsichore_subtilis_2 Terpsichore_taxifolia_Brazil Terpsichore_taxifolia_Ecuador Terpsichore_vascoi_ined. Terpsichore_zeledoniana Xiphopteris_conjunctisora Zygophlebia_mathewsii_Costa_Rica Zygophlebia_mathewsii_Ecuador Zygophlebia_sectifrons ; END; BEGIN TREES; TITLE Sundue_Islam_Ranker_TB; LINK TAXA = Taxa; TREE t1 = [&R] (Campyloneurum_angustifolium,((Microgramma_percussa,Microgramma_squamulosa),((Pecluma_alfredii,Polypodium_glycyrrhiza),((Grammitis_melanoloma,(Cochlidium_seminudum,(Cochlidium_punctatum,Cochlidium_rostratum))),((Terpsichore_hanekeana,Terpsichore_lehmanniana),(((Terpsichore_lanigera,(Terpsichore_smithii,(Terpsichore_cultrata,(Terpsichore_mollissima,Terpsichore_senilis)))),(Lellingeria_pseudomitchellae,Lellingeria_schenckei)),(((Chrysogrammitis_musgraveana,((Micropolypodium_achilleifolium,Terpsichore_longisetosa),(Micropolypodium_hyalinum,(Micropolypodium_taenifolium,Micropolypodium_zurquinum)))),((((Xiphopteris_conjunctisora,(Grammitis_knutsfordiana,(Grammitis_parva,(Grammitis_forbesiana,Oreogrammitis_hookeri)))),(Ctenopteris_nutans,(Ctenopteris_rhodocarpa,(Prosaptia_obliquata,(Prosaptia_alata,(Prosaptia_contigua,Prosaptia_pubipes)))))),(Ctenopteris_lasiostipes,(Grammitis_deplanchei,(((Grammitis_poeppigiana,Grammitis_poeppigiana_2),(Grammitis_billardieri,(Grammitis_ciliata,Grammitis_pseudociliata))),(Ctenopteris_heterophylla,Ctenopteris_heterophylla_2))))),((Ctenopteris_repandula,(Scleroglossum_sulcatum,Scleroglossum_sulcatum_2)),(Calymmodon_gracilis,Calymmodon_luerssenianus)))),(((((Terpsichore_nana,(Terpsichore_pichinchense_3,(Terpsichore_pichinchense,Terpsichore_pichinchense_2))),((Terpsichore_davidsmithii_5,((Terpsichore_davidsmithii_3,Terpsichore_davidsmithii_4),(Terpsichore_davidsmithii,Terpsichore_davidsmithii_2))),((Terpsichore_athyrioides,((Terpsichore_colombiense_ined.,Terpsichore_colombiense_ined._2),(Terpsichore_clathrata,Terpsichore_clavigera_ined.))),((Terpsichore_cuencana,Terpsichore_pichinchae),((Terpsichore_dilatata,Terpsichore_dilatata_2),((Terpsichore_loja_ined._2,(Terpsichore_loja_ined.,Terpsichore_loja_ined._3)),(Terpsichore_angustipes,Terpsichore_angustipes_2))))))),(Terpsichore_anfractuosa_Mexico,((Terpsichore_anfractuosa_Peru_2,(Terpsichore_anfractuosa_Mexico_2,(Terpsichore_anfractuosa_Peru,(Terpsichore_anfractuosa_Ecuador_2,(Terpsichore_anfractuosa_Ecuador,Terpsichore_anfractuosa_Costa_Rica))))),(Terpsichore_anfractuosa_Bolivia,Terpsichore_anfractuosa_Bolivia_2)))),(((Terpsichore_cretata,(Terpsichore_subtilis,Terpsichore_subtilis_2)),(((Terpsichore_sp._aff._alsopteris_ined._2_Costa_Rica,(Terpsichore_alsopteris,((Terpsichore_leucosticta,Terpsichore_leucosticta_2),((Terpsichore_sp._Sundue_1167,(Terpsichore_longicaulis,(Terpsichore_praeceps,Terpsichore_aff._subtilis_inded.))),(Terpsichore_attenuatissima_2,(((Terpsichore_semihirsuta_Colombia_Choco,(Terpsichore_semihirsuta_Peru,(Terpsichore_semihirsuta_Ecuador,Terpsichore_semihirsuta_Colombia))),(Terpsichore_semihirsuta_ined._3_Mexico,(Terpsichore_semihirsuta,(Terpsichore_semihirsuta_ined._2_Costa_Rica,(Terpsichore_sp._Sundue_1160,Terpsichore_vascoi_ined.))))),(Terpsichore_attenuatissima,Terpsichore_attenuatissima_3))))))),(Terpsichore_sp._aff._alsopteris_ined._1_Bolivia_3,(Terpsichore_sp._aff._alsopteris_ined._1_Bolivia_4,((Terpsichore_semihirsuta_Bolivia,Terpsichore_semihirsuta_Bolivia_2),(Terpsichore_sp._aff._alsopteris_ined._1_Bolivia,Terpsichore_sp._aff._alsopteris_ined._1_Bolivia_2))))),(Terpsichore_pacifica,(Terpsichore_zeledoniana,(Terpsichore_sp._aff._taxifolia_ined._Colombia,((Terpsichore_longipilosa_ined.,Terpsichore_longipilosa_ined._2),((Terpsichore_taxifolia_Ecuador,Terpsichore_taxifolia_Brazil),(Terpsichore_amphidasyon_2,(Terpsichore_amphidasyon_Moran_7646,Terpsichore_amphidasyon_3))))))))),((Terpsichore_delicatula,(Terpsichore_subscabra,Terpsichore_subscabra_2)),((((Melpomene_firma,(Melpomene_occidentalis,Melpomene_occidentalis_2)),(Melpomene_personata_ined.,(Melpomene_vernicosa,(Melpomene_anazalea_ined.,(Melpomene_moniliformis,(Melpomene_pseudonutans,Melpomene_flabelliformis)))))),(((Lellingeria_myosuroides_2,Lellingeria_myosuroides),(Lellingeria_hellwigii,Lellingeria_saffordii)),(Lellingeria_prionodes,(Lellingeria_limula_3,(Lellingeria_limula,Lellingeria_limula_2))))),(Lellingeria_subimpressa,((Lellingeria_suprasculpta,(Lellingeria_suspensa,Lellingeria_suspensa_2)),(Lellingeria_simacensis,((Lellingeria_bishopii,(Lellingeria_subsessilis_Costa_Rica,Lellingeria_subsessilis_Ecuador)),((Lellingeria_humilis,Lellingeria_major_2),((Lellingeria_major,Lellingeria_major_3),(Lellingeria_apiculata_Costa_Rica,Lellingeria_apiculata_Brazil))))))))))),(((Zygophlebia_sectifrons,(Zygophlebia_mathewsii_Ecuador,Zygophlebia_mathewsii_Costa_Rica)),((Enterosora_trichosora,(Enterosora_percrassa,Enterosora_percrassa_2)),(Enterosora_enterosoroides,(Enterosora_trifurcata_Ecuador,Enterosora_trifurcata_Costa_Rica)))),((Ceradenia_jungermannioides,(Ceradenia_fucoides,(Ceradenia_pilipes_Ecuador,Ceradenia_pilipes_Costa_Rica))),((Ceradenia_kalbreyeri_Colombia,(Ceradenia_kalbreyeri_Costa_Rica,Ceradenia_sp._aff._kalbreyeri)),(Ceradenia_intonsa,((Ceradenia_sp._moran_7520,(Ceradenia_sp._ined._1,Ceradenia_sp._ined._1_2)),(Ceradenia_spixiana,(Ceradenia_aulaeifolia,Ceradenia_aulaeifolia_2))))))))))))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr19939] END;