#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 18, 2025; 19:12 GMT TreeBASE (cc) 1994-2008 Study reference: Simmons M.P., Bacon C.D., Cappa J., & Mckenna M.J. 2011. Phylogeny of Celastraceae subfamilies Cassinoideae and Tripterygioideae Inferred from Morphological Characters and Nuclear and Plastid Loci. Systematic Botany, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S11387] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=246; TAXLABELS Acanthothamnus_aphyllus Allocassine_laurifolia Apatophyllum_flavovirens Apatophyllum_teretifolium Astrocassine_fusiformis_2917 Astrocassine_fusiformis_2918 Astrocassine_mastophorus Astrocassine_pleurostyloides Brassiantha_hedraiantheroides Brassiantha_pentamera Brexia_madagascariensis Canotia_holacantha Canotia_wendtii Cassine_peragua Cassine_schinoides Catha_edulis_1490_90 Catha_edulis_3016 Celastrus_angulatus Celastrus_flagellaris Celastrus_hindsii Celastrus_hypoleucus Celastrus_orbiculatus Celastrus_rosthornianus Celastrus_scandens Celastrus_strigillosus Celastrus_vulcanicolus Crossopetalum_decussatum Crossopetalum_eucymosum Crossopetalum_gaumeri Crossopetalum_ilicifolium Crossopetalum_managuatillo Crossopetalum_parviflorum Crossopetalum_rhacoma Crossopetalum_riparium Crossopetalum_uragoga Denhamia_bilocularis Denhamia_celastroides Denhamia_cunninghamii Denhamia_disperma Denhamia_fournieri Denhamia_obscura Denhamia_oleaster Denhamia_pittosporoides Denhamia_silvestris Denhamia_viridissima Dicarpellum_baillonianum Dicarpellum_pancheri Dinghoua_globularis_2628 Dinghoua_globularis_4739 Elaeodendron_australe_var._angustifolia Elaeodendron_australe_var._integrifolium Elaeodendron_cunninghamii Elaeodendron_curtipendulum Elaeodendron_matabelicum Elaeodendron_melanocarpum Elaeodendron_orientale Elaeodendron_pininsulare Elaeodendron_vitiense Elaeodendron_xylocarpum Empleuridium_juniperinum Euonymus_aff_carnosus Euonymus_aff_hamiltonianus Euonymus_alatus Euonymus_americanus Euonymus_australianus Euonymus_europaeus Euonymus_fimbriatus Euonymus_fortunei Euonymus_grandiflorus Euonymus_hamiltonianus_1791 Euonymus_hamiltonianus_1792 Euonymus_japonicus Euonymus_lanceolatus Euonymus_latifolius Euonymus_macropterus Euonymus_melananthus Euonymus_myrianthus Euonymus_nanus Euonymus_nitidus Euonymus_oxyphyllus Euonymus_phellomanus Euonymus_semenovii Euonymus_sp Euonymus_sp_sect_Euonymus Euonymus_tingens Euonymus_vagans Euonymus_verrucosus Fraunhofera_multiflora Glyptopetalum_palawanense Glyptopetalum_rhytidophyllum Gyminda_latifolia Gyminda_tonduzii Gymnosporia_arbutifolia Gymnosporia_cassinoides_2201 Gymnosporia_cassinoides_3525 Gymnosporia_crenata Gymnosporia_divaricata Gymnosporia_diversifolia Gymnosporia_engleriana Gymnosporia_harveyana Gymnosporia_inermis Gymnosporia_linearis Gymnosporia_littoralis Gymnosporia_mossambicensis Gymnosporia_palauica Gymnosporia_pertinax Gymnosporia_polyacantha Gymnosporia_pyracantha Gymnosporia_pyria Gymnosporia_royleana Gymnosporia_senegalensis_3030 Gymnosporia_senegalensis_4_93 Gymnosporia_verrucosa Gymnosporia_wallichiana Hartogiopsis_trilobocarpa_1259 Hartogiopsis_trilobocarpa_1278 Haydenia_gentryi_1866 Haydenia_gentryi_2267 Haydenia_haberiana Haydenia_urbaniana_3269 Haydenia_urbaniana_9169 Hedraianthera_porphyropetala_4370 Hedraianthera_porphyropetala_4544 Hexaspora_pubescens Hypsophila_dielsiana Hypsophila_halleyana Kokoona_sp Lauridia_reticulata Lauridia_tetragona Lepidobotrys_staudtii Lepuropetalon_spathulatum Lophopetalum_arnhemicum Lydenburgia_abbottii Lydenburgia_cassinoides Macgregoria_racemigera Maurocenia_frangula Maytenus_abbottii Maytenus_aff_obtusifolia Maytenus_aquifolium Maytenus_boaria Maytenus_buxifolia Maytenus_cordata Maytenus_disticha Maytenus_distichophylla Maytenus_domingensis Maytenus_elliptica Maytenus_elongata Maytenus_evonymoides Maytenus_filliformis Maytenus_floribunda Maytenus_grenadensis Maytenus_ilicifolia Maytenus_magellanica_753 Maytenus_magellanica_8438 Maytenus_oleosa Maytenus_peduncularis Maytenus_procumbens Maytenus_rigida Maytenus_robusta Maytenus_salicifolia Maytenus_spinosa Maytenus_texana Maytenus_undata_3028 Maytenus_undata_64700 Maytenus_verticillata Maytenus_woodsonii Menepetalum_salicifolium Menepetalum_schlechteri Microtropis_discolor Microtropis_fokienensis_229 Microtropis_fokienensis_37 Microtropis_japonica Microtropis_sp Microtropis_tetragona Microtropis_triflora Monimopetalum_chinense_38 Monimopetalum_chinense_9138 Mortonia_diffusa Mortonia_greggii Mortonia_hidalgensis Mortonia_scabrella Mystroxylon_aethiopicum_subsp._macrocarpum Orthosphenia_mexicana_1040 Orthosphenia_mexicana_19104 Parnassia_faberi Parnassia_glauca Parnassia_laxmannii Parnassia_palustris_2600 Parnassia_palustris_Fay Parnassia_tenella Parnassia_trinervis Paxistima_canbyi Paxistima_myrsinites Peripterygia_marginata Platypterocarpus_tanganyikensis Plenckia_populnea_12780 Plenckia_populnea_21279 Plenckia_populnea_8611 Pleurostylia_capensis Pleurostylia_leucocarpa Pleurostylia_opposita Polycardia_lateralis Polycardia_libera Polycardia_phyllanthoides Pottingeria_acuminata Psammomoya_choretroides Psammomoya_ephedroides Psammomoya_implexa Pseudocatha_mandenensis_2996 Pseudocatha_mandenensis_2997 Pseudosalacia_streyi Pterocelastrus_echinatus Pterocelastrus_tricuspidatus Quetzalia_occidentalis Quetzalia_schiedeana Quetzalia_stipitata Robsonodendron_eucleiforme Ruptiliocarpon_caracolito Rzedowskia_tolantonguensis_9025 Rzedowskia_tolantonguensis_9518 Salaciopsis_glomerata Salaciopsis_longistyla Salaciopsis_sparsiflora Schaefferia_angustifolia Schaefferia_argentinensis Schaefferia_cuneifolia Schaefferia_frutescens Schaefferia_pilosa Schaefferia_shrevei Schaefferia_stenophylla Siphonodon_australis Siphonodon_celastrineus Siphonodon_membranaceus Siphonodon_pendulus Stackhousia_monogyna Torralbasia_cuneifolia Tripterococcus_brunonis Tripterygium_regelii Wimmeria_bartletti Wimmeria_concolor Wimmeria_persicifolia Zinowiewia_concinna Zinowiewia_costaricensis_10927 Zinowiewia_costaricensis_1147 Zinowiewia_matudai Zinowiewia_rubra ; END; BEGIN TREES; TITLE Combined; LINK TAXA = Taxa; TREE PAUP_1 = [&R] ((Lepidobotrys_staudtii:0.013492,Ruptiliocarpon_caracolito:0.020015):0.0451575,((Lepuropetalon_spathulatum:0.064591,(Parnassia_faberi:0.015219,(Parnassia_laxmannii:0.013,((Parnassia_tenella:0.00709,Parnassia_trinervis:0.008107):0.005798,(Parnassia_glauca:0.019102,(Parnassia_palustris_Fay:4.05E-4,Parnassia_palustris_2600:6.16E-4):0.016771):0.002244):6.13E-4):0.002939):0.016056):0.098973,((Pottingeria_acuminata:0.035951,((Mortonia_diffusa:0.004386,Mortonia_hidalgensis:0.004068):0.004563,(Mortonia_greggii:0.004265,Mortonia_scabrella:0.014777):0.006917):0.060103):0.007617,(((Zinowiewia_concinna:0.002093,Zinowiewia_costaricensis_10927:0.002012,Zinowiewia_costaricensis_1147:0.001199,Zinowiewia_matudai:0.001408,Zinowiewia_rubra:0.001116):0.028795,((Quetzalia_occidentalis:7.07E-4,(Quetzalia_schiedeana:0.0,Quetzalia_stipitata:6.84E-4):4.61E-4):0.007609,((Microtropis_discolor:0.005384,(Microtropis_sp:0.003934,Microtropis_tetragona:6.49E-4):0.00381):0.006062,(Microtropis_japonica:0.003245,(Microtropis_triflora:0.002073,(Microtropis_fokienensis_37:0.0,Microtropis_fokienensis_229:0.0):0.001575):7.27E-4):0.011318):0.003677):0.010755):0.03365,((Monimopetalum_chinense_38:0.004361,Monimopetalum_chinense_9138:0.0):0.055462,(((Peripterygia_marginata:0.016978,(Siphonodon_celastrineus:0.009273,(Siphonodon_membranaceus:0.00563,(Siphonodon_australis:5.49E-4,Siphonodon_pendulus:0.004041):0.001964):0.006169):0.02579,(Crossopetalum_managuatillo:0.01133,(Crossopetalum_decussatum:0.014509,(Crossopetalum_ilicifolium:0.0,Crossopetalum_parviflorum:4.9E-4):0.010315,(Crossopetalum_riparium:0.005688,(Crossopetalum_gaumeri:0.003762,Crossopetalum_uragoga:0.007531,(Crossopetalum_eucymosum:0.001192,Crossopetalum_rhacoma:0.004239):0.004424):0.002612):0.008199):0.006938):0.014539):0.001447,(Hexaspora_pubescens:0.041488,(((Brassiantha_hedraiantheroides:0.00523,Brassiantha_pentamera:0.004443):0.007016,(Hedraianthera_porphyropetala_4370:6.95E-4,Hedraianthera_porphyropetala_4544:4.47E-4):0.011909):0.01007,((Dicarpellum_baillonianum:0.004224,Dicarpellum_pancheri:0.00199):0.006141,(Hypsophila_dielsiana:0.00396,Hypsophila_halleyana:0.002428):0.005282):0.013098):0.006207,((Denhamia_cunninghamii:0.016161,Denhamia_fournieri:0.004517,(Denhamia_bilocularis:0.003575,Denhamia_disperma:0.004932,Denhamia_silvestris:0.004022):0.005579):0.007297,(Denhamia_oleaster:0.007811,(Denhamia_celastroides:0.001511,Denhamia_pittosporoides:0.002484,(Denhamia_obscura:0.004114,Denhamia_viridissima:0.001935):6.5E-4):0.001354):0.00847):0.01698,((Tripterococcus_brunonis:0.028836,(Macgregoria_racemigera:0.034809,Stackhousia_monogyna:0.033421):0.010867):0.07582,(((Dinghoua_globularis_2628:0.001361,Dinghoua_globularis_4739:8.55E-4):0.005049,(Menepetalum_salicifolium:0.006175,Menepetalum_schlechteri:0.008144):0.006311):0.010953,((Apatophyllum_flavovirens:0.009849,Apatophyllum_teretifolium:0.006347):0.006168,(Psammomoya_ephedroides:0.004944,(Psammomoya_choretroides:0.00473,Psammomoya_implexa:0.00307):0.002152):0.011101):0.020525):0.006809):0.00548):0.007766):0.003958,(((Robsonodendron_eucleiforme:0.009717,((Empleuridium_juniperinum:0.021237,(Maytenus_abbottii:0.00783,Pseudosalacia_streyi:0.005011):0.001974):0.001167,((Pterocelastrus_echinatus:0.001137,Pterocelastrus_tricuspidatus:0.005393):0.00528,((Maytenus_filliformis:0.005756,(Maytenus_cordata:0.006131,(Maytenus_peduncularis:0.001991,Mystroxylon_aethiopicum_subsp._macrocarpum:0.0):0.007168):0.002274):0.002736,(Maytenus_oleosa:0.001273,(Maytenus_undata_3028:0.001921,(Maytenus_procumbens:9.7E-4,Maytenus_undata_64700:0.0):0.002077):0.003921):0.004438):0.001281):0.00116):0.001469):0.015394,(((Kokoona_sp:0.023178,Lophopetalum_arnhemicum:0.0238):0.013179,(Brexia_madagascariensis:0.010252,Polycardia_libera:0.01266,(Polycardia_lateralis:0.010468,Polycardia_phyllanthoides:0.007434):0.002839):0.013241,((((Astrocassine_fusiformis_2917:0.0,Astrocassine_fusiformis_2918:0.0):0.011005,(Astrocassine_mastophorus:0.00285,Astrocassine_pleurostyloides:0.004063):0.003916):0.002121,((Hartogiopsis_trilobocarpa_1259:0.0,Hartogiopsis_trilobocarpa_1278:0.0):0.005211,(Pleurostylia_capensis:0.004757,(Pleurostylia_leucocarpa:0.001372,Pleurostylia_opposita:0.005003):3.62E-4):0.00465):0.004595):0.012247,((Elaeodendron_orientale:0.004881,Elaeodendron_vitiense:0.004996):0.005704,((Pseudocatha_mandenensis_2996:0.0,Pseudocatha_mandenensis_2997:0.0):0.012663,((Elaeodendron_matabelicum:0.001996,Elaeodendron_xylocarpum:0.005202):0.007786,((Elaeodendron_melanocarpum:0.001336,(Elaeodendron_australe_var._angustifolia:3.37E-4,Elaeodendron_australe_var._integrifolium:3.45E-4):3.23E-4):0.005226,(Elaeodendron_pininsulare:0.004274,(Elaeodendron_cunninghamii:0.004415,Elaeodendron_curtipendulum:0.004176):0.001163):2.95E-4):0.006193):0.003597):0.00107):0.006177):0.007725):0.005912,((Salaciopsis_sparsiflora:0.010294,(Salaciopsis_glomerata:0.001897,Salaciopsis_longistyla:0.002771):0.003488):0.024833,(((Catha_edulis_1490_90:0.0,Catha_edulis_3016:0.0):0.007171,((Lauridia_reticulata:0.007143,(Allocassine_laurifolia:0.002679,Lauridia_tetragona:0.005577):7.06E-4):0.001773,(Cassine_peragua:0.006348,Cassine_schinoides:0.007539,Maurocenia_frangula:0.007716):9.71E-4):0.00134):0.002512,((Lydenburgia_abbottii:0.003244,(Lydenburgia_cassinoides:0.010049,Platypterocarpus_tanganyikensis:0.011945):0.001159):2.04E-4,(Gymnosporia_divaricata:0.002003,((Gymnosporia_harveyana:0.0,Gymnosporia_mossambicensis:0.0):0.002274,(Gymnosporia_pyracantha:0.001862,Gymnosporia_verrucosa:0.001916):0.001664,(Gymnosporia_engleriana:0.002206,Gymnosporia_polyacantha:4.68E-4,(Gymnosporia_arbutifolia:6.51E-4,Gymnosporia_royleana:3.29E-4):0.002176,((Gymnosporia_linearis:0.001238,Gymnosporia_senegalensis_3030:0.003359):0.00346,(Gymnosporia_pyria:0.001068,Gymnosporia_senegalensis_4_93:0.001204):9.34E-4):0.005194,((Gymnosporia_cassinoides_2201:0.0,Gymnosporia_cassinoides_3525:3.4E-4):0.007458,(Gymnosporia_diversifolia:0.004244,(Gymnosporia_inermis:0.001999,(Gymnosporia_littoralis:3.22E-4,Gymnosporia_wallichiana:3.26E-4):3.21E-4,(Gymnosporia_palauica:0.001144,(Gymnosporia_crenata:0.0,Gymnosporia_pertinax:6.65E-4):0.001314):0.00111):0.001027):0.001465):0.001015):0.001792):0.002939):0.00116):0.00304):0.013446):0.002024):0.00124):0.004572,((((Haydenia_gentryi_1866:0.0,Haydenia_gentryi_2267:0.0):0.002501,(Haydenia_haberiana:0.002866,(Haydenia_urbaniana_3269:6.58E-4,Haydenia_urbaniana_9169:8.55E-4):0.003338):0.001192):0.010517,((Gyminda_latifolia:0.003777,Gyminda_tonduzii:0.004805):0.012873,(((Orthosphenia_mexicana_1040:0.001006,Orthosphenia_mexicana_19104:8.26E-4):0.004185,(Rzedowskia_tolantonguensis_9025:0.0,Rzedowskia_tolantonguensis_9518:0.00188):0.009244):0.01107,(Schaefferia_argentinensis:0.012399,Schaefferia_cuneifolia:0.007557,Schaefferia_stenophylla:0.008954,(Schaefferia_angustifolia:0.001427,Schaefferia_frutescens:7.04E-4):0.00828,(Schaefferia_pilosa:0.004109,Schaefferia_shrevei:0.001302):0.004983):0.015764):0.003793):0.008427):0.002178,((((Maytenus_verticillata:0.003451,Maytenus_woodsonii:0.006229):6.8E-4,(Maytenus_disticha:0.001331,(Maytenus_boaria:0.005083,(Maytenus_magellanica_753:0.0,Maytenus_magellanica_8438:0.0):0.003716):0.001914):0.004116):0.013843,(Maytenus_texana:0.01852,((Fraunhofera_multiflora:0.016644,(Plenckia_populnea_8611:0.0,Plenckia_populnea_12780:0.001685,Plenckia_populnea_21279:0.002034):0.005018):0.005452,(Maytenus_spinosa:0.017574,(Maytenus_distichophylla:0.003009,Maytenus_evonymoides:0.002162):0.002639,(Maytenus_aquifolium:0.005116,Maytenus_floribunda:0.004771,Maytenus_ilicifolia:0.003463,Maytenus_aff_obtusifolia:0.010733,(Maytenus_buxifolia:0.004488,Maytenus_domingensis:0.002875,Maytenus_elliptica:3.32E-4,Maytenus_elongata:7.48E-4):0.003116,((Maytenus_grenadensis:0.005876,Maytenus_rigida:0.007297):0.002438,(Maytenus_robusta:8.71E-4,Maytenus_salicifolia:7.73E-4):0.003441):0.00381):0.003698):0.004861):0.002176):0.005435):0.007284,((Tripterygium_regelii:0.003457,(Celastrus_vulcanicolus:0.011832,((Celastrus_angulatus:0.00133,Celastrus_hypoleucus:0.001349):0.012159,((Celastrus_rosthornianus:0.007895,Celastrus_scandens:0.010016):0.001621,(Celastrus_flagellaris:0.002044,Celastrus_strigillosus:7.78E-4,(Celastrus_hindsii:3.39E-4,Celastrus_orbiculatus:8.14E-4):0.002153):0.006225):0.001715):0.0015):0.001012):0.017854,(((Paxistima_canbyi:0.002682,Paxistima_myrsinites:0.00417):0.009864,(Wimmeria_persicifolia:0.007807,(Wimmeria_bartletti:0.002046,Wimmeria_concolor:0.003024):0.006369):0.019814):0.00276,((Acanthothamnus_aphyllus:0.016493,(Canotia_holacantha:0.007462,Canotia_wendtii:0.004115):0.008783):0.014268,(((Euonymus_lanceolatus:8.85E-4,Euonymus_melananthus:0.0):0.00539,(Euonymus_americanus:0.00943,(Euonymus_latifolius:0.004003,(Euonymus_fimbriatus:0.003603,(Euonymus_macropterus:9.32E-4,Euonymus_oxyphyllus:0.001877):0.0):0.001108):0.002579):0.001142):6.77E-4,(Torralbasia_cuneifolia:0.0,(((Glyptopetalum_palawanense:0.010589,Glyptopetalum_rhytidophyllum:0.001612):0.007886,(Euonymus_vagans:0.002222,(Euonymus_sp:0.005839,(Euonymus_fortunei:0.001278,Euonymus_japonicus:6.76E-4):0.002789):0.003251):0.007645):5.43E-4,((Euonymus_verrucosus:0.009374,((Euonymus_alatus:0.002579,Euonymus_phellomanus:0.003801):0.004763,(Euonymus_aff_carnosus:0.005314,Euonymus_grandiflorus:0.004293):0.010835):3.2E-4):0.002436,((Euonymus_australianus:0.008739,(Euonymus_nanus:0.015584,(Euonymus_semenovii:0.001685,Euonymus_tingens:0.002905):0.005416):0.001482):4.07E-4,((Euonymus_nitidus:0.001478,(Euonymus_myrianthus:0.00149,Euonymus_sp_sect_Euonymus:5.97E-4):4.8E-4):0.004958,(Euonymus_europaeus:0.001343,(Euonymus_hamiltonianus_1791:0.0,(Euonymus_hamiltonianus_1792:0.0,Euonymus_aff_hamiltonianus:0.0):2.04E-4):0.004105):0.011943):3.19E-4):5.48E-4):0.001277):0.0):6.51E-4):0.007474):8.7E-4):0.00261):0.00174):3.38E-4):0.005573):0.006845):0.018647):0.01261):0.003823):0.010004):0.0451575); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr43817] END;