#NEXUS

[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on September 18, 2025; 23:54 GMT

TreeBASE (cc) 1994-2008

Study reference:
Zhang J., Meng S., Wen J., & Rao G. 2013. Phylogenetic Relationships and Character
Evolution of Rhodiola (Crassulaceae) based on Nuclear Ribosomal ITS, trnL-F, and
psbA-trnH Sequences. Systematic Botany, .

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S14270]

BEGIN TAXA;
        TITLE  Taxa;
        DIMENSIONS NTAX=78;
        TAXLABELS
            Aeonium_pseudourbicum
            Aeonium_undulatum
            Greenovia_aurea
            Hylotelephium_spectabile
            Hylotelephium_tatarinowii
            Kalanchoe_densiflora
            Kalanchoe_tomentosa
            Meterostachys_sikokiana
            Orostachys_japonica
            Orostachys_spinosa
            Phedimus_aizoon
            Phedimus_kamtschaticus
            Pseudosedum_Sp.
            Pseudosedum_longidentatum
            Rhodiola_alsia
            Rhodiola_alterna
            Rhodiola_amabilis
            Rhodiola_angusta
            Rhodiola_atsaensis
            Rhodiola_brevipetiolata
            Rhodiola_bupleuroides
            Rhodiola_calliantha
            Rhodiola_chrysanthemifolia
            Rhodiola_coccinea
            Rhodiola_crenulata
            Rhodiola_cretinii
            Rhodiola_discolor
            Rhodiola_dumulosa
            Rhodiola_eurycarpa
            Rhodiola_fastigiata
            Rhodiola_forrestii
            Rhodiola_gannanica
            Rhodiola_henryi
            Rhodiola_heterodonta
            Rhodiola_himalensis
            Rhodiola_hobsonii
            Rhodiola_humilis
            Rhodiola_integrifolia_Alaska_Allen
            Rhodiola_integrifolia_Canada_Allen
            Rhodiola_ishidae
            Rhodiola_kirilowii
            Rhodiola_liciae
            Rhodiola_macrocarpa
            Rhodiola_nepalica
            Rhodiola_nobilis
            Rhodiola_ovatisepala
            Rhodiola_ovatisepala_v_ovati
            Rhodiola_prainii
            Rhodiola_purpureoviridis_R
            Rhodiola_quadrifida
            Rhodiola_rhodantha_Wyoming_Allen
            Rhodiola_rosea
            Rhodiola_rosea_Austria
            Rhodiola_rosea_Canada_Allen
            Rhodiola_sachalinensis
            Rhodiola_sacra
            Rhodiola_scabrida
            Rhodiola_semenovii_Janssen_2205
            Rhodiola_serrata
            Rhodiola_sexifolia
            Rhodiola_sinuata
            Rhodiola_smithii
            Rhodiola_smithii_2
            Rhodiola_sp.
            Rhodiola_stapfii
            Rhodiola_tangutica
            Rhodiola_tibetica
            Rhodiola_tieghemii
            Rhodiola_wallichiana
            Rhodiola_yunnanensis
            Sedum_bulbiferum
            Sedum_japonicum
            Sedum_mexicanum
            Sedum_obtrullatum
            Sedum_oreades
            'Sedum zentaro-tashiroi'
            Sinocrassula_densirosulata
            Sinocrassula_indica
            ;
END;

BEGIN TREES;
      TITLE ITS_trnlf_psbA_B;
      LINK TAXA = Taxa;
      TREE Fig._1 = [&R] ((Kalanchoe_densiflora:0.03925595245486826,Kalanchoe_tomentosa:0.02015914390026065):0.04441755834381328,((((Aeonium_pseudourbicum:0.003910978573354345,(Aeonium_undulatum:0.008451008254109982,Greenovia_aurea:0.01322499418698786):0.003417415687578332):0.129590165305912,(Sedum_japonicum:0.3005004540501944,(Sedum_obtrullatum:0.08665163818020652,Sedum_oreades:0.04137087452034667):0.06940343179151245):0.0228739464886032):0.02169941848957681,((Sedum_mexicanum:0.05357385226468995,(Sedum_bulbiferum:0.0125644460235988,'Sedum zentaro-tashiroi':0.07286624480635487):0.04666969464417826):0.05519752250548695,((Sinocrassula_densirosulata:0.01566118544423071,Sinocrassula_indica:0.01175911287814861):0.06167608926361108,((Hylotelephium_spectabile:0.0366808251307509,Hylotelephium_tatarinowii:0.02739355460964022):0.01345828674855296,(Meterostachys_sikokiana:0.03389760502459686,(Orostachys_japonica:0.02035364908642864,Orostachys_spinosa:0.01902176465478588):0.0214130957221349):0.02280688030810585):0.01498553393460951):0.03488035762108891):0.02835777221783058):0.005241235656321341,((Phedimus_aizoon:0.002431360902655427,Phedimus_kamtschaticus:0.001890747293397597):0.1036502099902829,((Pseudosedum_longidentatum:0.005015867769033577,Pseudosedum_Sp.:0.007246716213808312):0.02422084315491672,(Rhodiola_smithii:0.001682494786627136,Rhodiola_smithii_2:0.005352753347232354):0.01667343808578437,(Rhodiola_semenovii_Janssen_2205:0.01627386996865617,((Rhodiola_atsaensis:0.00768250116869878,(Rhodiola_discolor:0.008457317824814238,Rhodiola_henryi:0.008723782091609811):0.00328262455387122):0.005759737352648486,(Rhodiola_angusta:0.008728897477238208,(Rhodiola_crenulata:0.007950211536662801,(Rhodiola_bupleuroides:0.008006049748030715,Rhodiola_serrata:0.00908901722252778):0.002783410328876112):0.004050592716353208):0.001024291550781745):0.00184961115947062):0.002397416227327859,((Rhodiola_wallichiana:0.0155379052165624,(Rhodiola_rhodantha_Wyoming_Allen:0.0144258390447665,(Rhodiola_integrifolia_Alaska_Allen:0.005588732027492729,Rhodiola_integrifolia_Canada_Allen:0.004020565717724823):0.005413153416349672):0.006797062603507101):0.002400014863128126,(Rhodiola_hobsonii:0.01028849664476052,(Rhodiola_humilis:0.01849070827320395,((Rhodiola_prainii:0.01071541620463577,Rhodiola_stapfii:0.01043768211818603):0.002550043030821029,((Rhodiola_chrysanthemifolia:0.003210320664466159,Rhodiola_sinuata:4.82205340532545E-4):0.00223378141315859,(Rhodiola_tieghemii:0.002498989378434477,(Rhodiola_ovatisepala:4.764116033728103E-4,Rhodiola_ovatisepala_v_ovati:4.719129751705295E-4):0.002540443585000193):0.01029180301205798,(Rhodiola_sacra:0.009197824702696162,(Rhodiola_alterna:0.007296860649694635,(Rhodiola_liciae:4.833306800596677E-4,Rhodiola_sexifolia:4.726777172477708E-4):0.002767491040266439):0.00214210026858381):0.001320514626606701):0.003829647662283876):0.001218919358508875):0.001279605441827139):0.00381027248086308):0.005289222543111065,((Rhodiola_tibetica:0.005991723669593923,(Rhodiola_dumulosa:0.001819121758541857,Rhodiola_gannanica:0.001100399694732234):0.006697686882882549):0.001254400374399167,(Rhodiola_scabrida:0.01249817487299603,(((Rhodiola_forrestii:0.009216869726700546,Rhodiola_yunnanensis:0.005324553655251313):0.008319095462203826,(Rhodiola_kirilowii:0.006996854828363938,(Rhodiola_sp.:0.01164331840783935,Rhodiola_sachalinensis:0.004989240604858222):0.001859070503837415):0.004067010676507986):0.001444829166879475,(Rhodiola_calliantha:0.007175003219849371,(Rhodiola_eurycarpa:0.003586027470694199,Rhodiola_macrocarpa:0.00460305491657818):0.003029613291963539,((Rhodiola_rosea:0.003035749252085051,Rhodiola_ishidae:0.003174940162355824):0.001452599955593989,(Rhodiola_coccinea:0.004364802853633796,(Rhodiola_alsia:0.003785690689504875,Rhodiola_fastigiata:0.001123972464316647,Rhodiola_tangutica:0.002479586863603527):0.002610123510577294):0.001313929349250459):0.003365340725947611,(Rhodiola_brevipetiolata:0.005256370498552081,Rhodiola_himalensis:0.005231547929886478,Rhodiola_nepalica:0.01587304785204475,(Rhodiola_nobilis:0.002465150580563632,Rhodiola_purpureoviridis_R:0.003691089063667236):0.001107262888317848,(Rhodiola_amabilis:7.33385838924494E-4,Rhodiola_cretinii:7.717969795083252E-4):0.006329638724257105,(Rhodiola_quadrifida:0.003260655218886921,(Rhodiola_heterodonta:0.009210980864488709,(Rhodiola_rosea_Canada_Allen:0.0120843981192112,Rhodiola_rosea_Austria:6.108682144615072E-4):0.003146727287097676):0.003377821308271399):0.001101085668578188):0.002716102060510154):0.002633308962176295):0.001852447499345803):0.004341016303088141):0.001173632821639851):0.02469019336431926):0.03601141288835615):0.04441755834381328);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr63652]


END;