#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 18, 2025; 22:11 GMT TreeBASE (cc) 1994-2008 Study reference: Shaw B., Crandall-stotler B.J., Vana J., Stotler R.E., Von konrat M., Engel J.J., Davis E., Long D., Sova P., & Shaw A. 2014. Phylogenetic Relationships and Morphological Evolution in a Major Clade of Leafy Liverworts (Phylum Marchantiophyta, order Jungermanniales): suborder Jungermanniineae. Systematic Botany, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S16128] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=169; TAXLABELS Acrobolbus_ochrophyllus Acroscyphella_phoenicorhiza Alobiellopsis_parvifolius Anastrophyllopsis_subcomplicata Anthelia_julacea Arnellia_fennica Austrolophozia_paradoxa Balantiopsis_diplophylla Balantiopsis_splendens Bazzania_exempta Blepharidophyllum_vertebrale Bragginsella_anomala Calypogeia_muelleriana Cephalozia_bicuspidata Cephalozia_conchata Cephaloziella_divaricata Cephaloziella_hirta Clandarium_gottscheanum Clandarium_xiphophyllum Cryptocoleopsis_imbricata Cuspidatula_monodon Delavayella_serrata Endogemma_caespiticium Eocalypogeia_quelpartensis Eremonotus_myriocarpus Geocalyx_caledonicus Geocalyx_graveolens Goebelobryum_unguiculatum Gongylanthus_ericetorum Gongylanthus_liebmanianus Gongylanthus_liebmannianus_FATOL774 Gottschelia_schizopleura Gymnocolea_inflata Gymnomitrion_africanum Gymnomitrion_alpinum Gymnomitrion_commutatum Gymnomitrion_concinnatum Gymnomitrion_corallioides Gymnomitrion_pacificum Gymnomitrion_revolutum Gymnomitrion_rubidum Gymnomitrion_verrucosum Gyrothyra_underwoodiana Harpanthus_flotovianus Harpanthus_scutatus Herzogobryum_atrocapillum Herzogobryum_vermiculare Hygrobiella_laxifolia Isotachis_lyallii Jackiella_curvata Jackiella_javanica Jungermannia_atrovirens Jungermannia_exsertifoliaSspCordifolia Jungermannia_konstantinovae Jungermannia_polaris_L1280_France Jungermannia_polaris_L1281_Russia Jungermannia_polaris_L1420_Norway Jungermannia_pumila Leiomylia_anomala Lethocolea_glossophylla Liochlaena_lanceolata Liochlaena_subulata Lophocolea_heterophylla Lophozia_ventricosa Marsupella_aquatica Marsupella_arctica Marsupella_emarginata Marsupella_funckii Marsupella_sphacelata Marsupella_stoloniformis Marsupella_yakushimensis Mesoptychia_badensis Mesoptychia_bantriensis Mesoptychia_gillmanii Mesoptychia_heterocolpos Mesoptychia_morrisoncola Mesoptychia_polymorpha Mesoptychia_rutheana Mesoptychia_sahlbergii Metacalypogeia_alternifolia Mizutania_riccardioides Mnioloma_fuscum Mylia_taylorii Nardia_assamica Nardia_breidleri Nardia_compressa_FATOL538_Russia Nardia_compressa_L1429_Norway Nardia_geoscyphus_FATOL528_Russia Nardia_geoscyphus_IBC49_USA Nardia_geoscyphus_L1075_comb Nardia_insecta Nardia_japonica Nardia_lescurii Nardia_scalaris Nardia_succulenta Nothogymnomitrion_erosum Notoscyphus_lutescens Odontoschisma_sphagni Pedinophyllum_interruptum Plagiochila_porelloides Poeltia_campylata Prasanthus_suecicus Pseudolepicolea_quadrilaciniata Pseudolepicolea_trollii Saccogyna_viticulosa Saccogynidium_australe Saccogynidium_decurvum Scapania_nemorea Scapania_nimbosa Schistochila_laminigera Solenostoma_abyssinicum Solenostoma_appressifolium Solenostoma_atrorevolutum Solenostoma_callithrix Solenostoma_chenianum Solenostoma_clavellatum Solenostoma_comatum Solenostoma_confertissimum Solenostoma_coniflorum Solenostoma_crenuliforme Solenostoma_cryptogynum Solenostoma_dulongense Solenostoma_faurieanum Solenostoma_flagellatum Solenostoma_fusiforme Solenostoma_gracillimum Solenostoma_grosseverrucosum Solenostoma_handelii Solenostoma_hasskarlianum Solenostoma_hyalinum Solenostoma_infuscum Solenostoma_inundatum Solenostoma_kashyapii Solenostoma_lanigerum Solenostoma_macrocarpum Solenostoma_marginatum Solenostoma_montanum Solenostoma_obovatum Solenostoma_obscurum Solenostoma_orbiculatum Solenostoma_parvitextum Solenostoma_patoniae Solenostoma_polyrhizoides Solenostoma_pseudopyriflorum Solenostoma_pusillum Solenostoma_pyriflorum Solenostoma_rosulans Solenostoma_rubripunctatum Solenostoma_rubrum Solenostoma_rufiflorum Solenostoma_sanguinolentum Solenostoma_schusterianum Solenostoma_speciosum Solenostoma_sphaerocarpum Solenostoma_subacutum Solenostoma_subellipticum Solenostoma_totipapillosum Solenostoma_truncatum Southbya_nigrella Southbya_tophacea Syzygiella_autumnalis Syzygiella_colorata Syzygiella_nipponica Syzygiella_sonderi Syzygiella_tasmanica Syzygiella_teres Trichotemnoma_corrugatum Tylimanthus_laxus Zoopsis_nitida ; END; BEGIN TREES; TITLE Leafy_Liverworts_MLT_analysis_of_10_loci_for_169_taxa; LINK TAXA = Taxa; TREE bestREP1_169taxa = [&R] ((((((((((((((((((((Solenostoma_speciosum:0.0058794,(Solenostoma_atrorevolutum:0.0073998,((((Solenostoma_kashyapii:0.00537024,(Solenostoma_dulongense:0.00211957,Solenostoma_lanigerum:0.00576347):4.878E-4):3.1811E-4,(Solenostoma_abyssinicum:0.00270851,Solenostoma_confertissimum:0.00287984):0.00282032):6.4912E-4,(Solenostoma_parvitextum:0.00456728,Solenostoma_chenianum:0.00897526):6.7336E-4):0.00182864,(((Solenostoma_grosseverrucosum:0.00109094,Solenostoma_subacutum:0.00277506):0.00433868,((((Solenostoma_coniflorum:9.0712E-4,((Solenostoma_rufiflorum:2.6055E-4,Solenostoma_inundatum:3.2352E-4):0.00114538,Solenostoma_totipapillosum:0.00146462):0.00147439):0.00446072,(Solenostoma_appressifolium:0.00415809,(Solenostoma_pseudopyriflorum:0.00520014,(Solenostoma_faurieanum:0.00303585,Solenostoma_pyriflorum:0.00333263):0.00482421):0.00213779):9.388E-4):0.00284761,(Solenostoma_clavellatum:0.00519811,Solenostoma_sanguinolentum:0.0061899):0.00166811):1.0E-8,Solenostoma_patoniae:0.00350747):7.0544E-4):0.00111837,(Solenostoma_pusillum:4.3843E-4,Solenostoma_sphaerocarpum:1.0E-8):0.0049096):1.9511E-4):0.00165849):0.00158808):0.00116098,Solenostoma_orbiculatum:0.01088951):0.00167983,Solenostoma_macrocarpum:0.00560405):0.0019707,Solenostoma_cryptogynum:0.01941645):0.01255916,((((Solenostoma_obscurum:1.0E-8,Solenostoma_obovatum:1.0E-8):1.0E-8,Solenostoma_subellipticum:0.00358309):0.00577129,Solenostoma_flagellatum:0.00592024):0.00846293,Solenostoma_schusterianum:0.01177245):0.00774275):0.00166191,((Solenostoma_fusiforme:0.008517,Solenostoma_gracillimum:0.01209418):0.00280253,(Solenostoma_rubrum:7.3181E-4,Solenostoma_handelii:7.8705E-4):0.01281127):0.01823492):0.00178979,((Solenostoma_crenuliforme:0.00461287,(((Solenostoma_rubripunctatum:0.01166597,(Solenostoma_truncatum:0.01484183,Solenostoma_montanum:0.0169525):9.8854E-4):9.8901E-4,((Solenostoma_rosulans:0.00584695,Solenostoma_comatum:0.01525183):0.0017257,(Solenostoma_infuscum:0.00893329,Solenostoma_marginatum:0.01674623):5.4551E-4):7.5011E-4):7.3465E-4,(Solenostoma_hyalinum:0.00432888,Solenostoma_callithrix:0.00418271):0.00354586):0.00302852):0.00875721,(Solenostoma_hasskarlianum:0.03672786,Solenostoma_polyrhizoides:0.00409311):0.01272039):0.0086355):0.02565973,((((((Gymnomitrion_corallioides:0.00623207,(Gymnomitrion_rubidum:0.00840775,(Gymnomitrion_commutatum:0.0063007,((Gymnomitrion_africanum:0.00328413,Gymnomitrion_revolutum:0.00525481):0.0031836,Gymnomitrion_verrucosum:0.00559764):0.00338583):0.0051298):7.9334E-4):0.00122446,Gymnomitrion_concinnatum:0.00984028):0.00400616,(Gymnomitrion_alpinum:0.0079217,Gymnomitrion_pacificum:0.00319606):0.00549015):0.00957461,(Prasanthus_suecicus:0.01107648,Cryptocoleopsis_imbricata:0.01161066):0.00591442):9.1029E-4,(((Marsupella_stoloniformis:0.01898,(Marsupella_arctica:0.00565588,(Marsupella_emarginata:0.00773268,(Marsupella_yakushimensis:0.00586809,Marsupella_aquatica:0.00693665):0.00167241):0.00243772):0.00310518):0.00235058,(Marsupella_funckii:0.00527107,Marsupella_sphacelata:0.00595883):0.01061773):0.00397344,Poeltia_campylata:0.02146279):0.0044612):0.00413333,((Nardia_insecta:0.01068167,(Nardia_succulenta:0.01575286,Nardia_assamica:0.01692811):0.01099731):0.00414156,((Nardia_japonica:0.00437926,Nardia_compressa_L1429_Norway:0.00689386):0.00253894,(((Nardia_geoscyphus_FATOL528_Russia:0.00450412,Nardia_lescurii:0.00370321):0.01111663,(Nardia_scalaris:0.01062357,((Nardia_compressa_FATOL538_Russia:0.00920971,Nardia_breidleri:0.00983101):0.00243738,Nardia_geoscyphus_L1075_comb:0.01166171):4.422E-4):0.00236459):4.4919E-4,Nardia_geoscyphus_IBC49_USA:0.00875672):0.00103833):0.00271823):0.01109488):0.002727):0.00292093,Endogemma_caespiticium:0.06941099):0.01921514,(((Harpanthus_flotovianus:0.01226014,Harpanthus_scutatus:0.01330747):0.0382607,Gyrothyra_underwoodiana:0.07856256):0.00392245,Arnellia_fennica:0.05435706):0.00483199):0.00155687,Saccogyna_viticulosa:0.0440459):0.00284385,((((Jungermannia_atrovirens:0.01568474,(((Jungermannia_exsertifoliaSspCordifolia:0.01326692,(Jungermannia_polaris_L1420_Norway:0.00670715,(Jungermannia_pumila:0.00721492,Jungermannia_polaris_L1281_Russia:0.00528324):0.00326408):0.00312058):0.00472353,Jungermannia_konstantinovae:0.00331285):0.00162851,Jungermannia_polaris_L1280_France:0.01724233):0.00683919):0.01981662,Eremonotus_myriocarpus:0.03286582):0.01191121,(Delavayella_serrata:0.03373748,(Liochlaena_subulata:0.00106598,Liochlaena_lanceolata:0.00111962):0.03088789):0.01372532):0.00409519,(((((Mesoptychia_bantriensis:0.00120312,Mesoptychia_gillmanii:0.00229164):0.01230006,Mesoptychia_rutheana:0.00538936):0.00213175,((Mesoptychia_polymorpha:0.01618501,Mesoptychia_morrisoncola:0.04440614):0.00742585,Mesoptychia_badensis:0.04553802):0.00254204):0.00585955,Mesoptychia_sahlbergii:0.00793984):0.00921657,Mesoptychia_heterocolpos:0.02404451):0.01013396):0.01365138):0.00549962,(Hygrobiella_laxifolia:0.02219427,(Anthelia_julacea:0.00922795,Pseudolepicolea_trollii:0.02029625):0.00894081):0.04346505):0.00240294,(((Jackiella_curvata:0.00394456,Jackiella_javanica:0.00337822):0.09354573,Notoscyphus_lutescens:0.07770611):0.01594343,(((((Gongylanthus_liebmannianus_FATOL774:0.0157495,Gongylanthus_liebmanianus:0.01195518):0.01965117,Gongylanthus_ericetorum:0.04004022):0.00759419,(Southbya_tophacea:0.01796112,Southbya_nigrella:0.02493352):0.03798679):0.03828193,(Geocalyx_caledonicus:0.00885626,Geocalyx_graveolens:0.01013691):0.02752503):0.00875747,((Eocalypogeia_quelpartensis:0.07867614,Metacalypogeia_alternifolia:0.04491085):0.00613659,(Calypogeia_muelleriana:0.04139058,(Mizutania_riccardioides:0.14087713,Mnioloma_fuscum:0.06206419):0.016716):0.03169876):0.01375098):0.00122681):0.00200579):0.01217807,((Trichotemnoma_corrugatum:0.03930205,(Acroscyphella_phoenicorhiza:0.02844142,(Isotachis_lyallii:0.02108117,(Balantiopsis_splendens:0.01586268,Balantiopsis_diplophylla:0.01233392):0.00729842):0.01343296):0.01677514):0.01693966,((Clandarium_xiphophyllum:0.01173918,Clandarium_gottscheanum:0.01414399):0.00165341,Blepharidophyllum_vertebrale:0.00764422):0.03220204):0.00375285):0.00286863,(Lethocolea_glossophylla:0.08729751,((Saccogynidium_australe:0.00654093,Saccogynidium_decurvum:0.01051028):0.05737176,((Austrolophozia_paradoxa:0.02774449,Goebelobryum_unguiculatum:0.04387277):0.03310489,(Acrobolbus_ochrophyllus:0.01123754,Tylimanthus_laxus:0.01668374):0.03679087):0.0064688):0.00703196):0.0135897):0.0252879,((Cuspidatula_monodon:0.01933,(Syzygiella_tasmanica:0.01121044,((Syzygiella_teres:0.01148625,((Syzygiella_nipponica:0.01045684,Syzygiella_autumnalis:0.01241524):0.00774322,Syzygiella_colorata:0.0130272):0.00232762):0.00103973,Syzygiella_sonderi:0.02070445):0.00698392):0.005133):0.02502447,(((Nothogymnomitrion_erosum:0.02514747,((Anastrophyllopsis_subcomplicata:0.0237681,Gottschelia_schizopleura:0.04002195):0.00402358,(Gymnocolea_inflata:0.03653301,((Scapania_nemorea:0.01645603,Scapania_nimbosa:0.01609476):0.01586486,Lophozia_ventricosa:0.01516466):0.00676852):0.00214435):0.00263947):0.0068439,((Cephaloziella_hirta:0.00511815,Cephaloziella_divaricata:0.01393416):0.07643186,(Herzogobryum_atrocapillum:0.00328349,Herzogobryum_vermiculare:0.00312093):0.04717163):0.00672562):0.02418022,((Cephalozia_conchata:0.010845,Cephalozia_bicuspidata:0.01134703):0.07341509,(Alobiellopsis_parvifolius:0.04670296,Odontoschisma_sphagni:0.075194):0.00760012):0.03311545):0.01665687):0.01742038):0.00877726,((Zoopsis_nitida:0.05194648,Bazzania_exempta:0.03704069):0.04568974,(Pseudolepicolea_quadrilaciniata:0.08727624,((Pedinophyllum_interruptum:0.02108007,Plagiochila_porelloides:0.02800625):0.01760522,(Bragginsella_anomala:0.02381589,Lophocolea_heterophylla:0.02856586):0.0351935):0.01896662):0.00570582):0.01138846):0.01245426,(Mylia_taylorii:0.01786765,Leiomylia_anomala:0.01240312):0.08965266):0.071762425,Schistochila_laminigera:0.071762425); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr76499] END;