#NEXUS

[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on September 18, 2025; 20:38 GMT

TreeBASE (cc) 1994-2008

Study reference:
Albert V., Scheen A., & Scheen A. 2009. Molecular phylogenetics of the Leucas group
(Lamioideae; Lamiaceae). Systematic Botany, 34(1): 173-181.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S2243]

BEGIN TAXA;
        TITLE  'combined trnL-trnF region, trnS-trnG spacer, and rps16 intron';
        DIMENSIONS NTAX=83;
        TAXLABELS
            Acrotome_angustifolia
            Acrotome_fleckii
            Acrotome_hispida
            Acrotome_inflata_1
            Acrotome_inflata_2
            Acrotome_pallescens
            Ballota_acetabulosa
            Ballota_pseudodictamnus
            Isoleucas_arabica
            Lamium_album_ssp_crinitum
            Lamium_tomentosum
            Leonotis_grandis
            Leonotis_leonurus
            Leonotis_myricifolia
            Leonotis_myrothamnifolia
            Leonotis_nepetifolia_var_africana
            Leonotis_nepetifolia_var_nepetifolia
            Leonotis_ocymifolia_var_raineriana_1
            Leonotis_ocymifolia_var_raineriana_2
            Leonotis_ocymifolia_var_schinzii
            Leucas_abyssinica_var_argyrophylla
            Leucas_abyssinica_var_brachycalyx
            Leucas_abyssinica_var_sidamoensis
            Leucas_aequistylosa
            Leucas_argentea
            Leucas_aspera
            Leucas_biflora
            Leucas_bracteosa
            Leucas_calostachys
            Leucas_capensis
            Leucas_ciliata
            Leucas_cuneifolia
            Leucas_decemdentata
            Leucas_deflexa
            Leucas_densiflora
            Leucas_discolor
            Leucas_flagellifera
            Leucas_glabrata
            Leucas_glabrata_var_glabrata
            Leucas_grandis
            Leucas_inflata
            Leucas_jamseii
            Leucas_kishenensis
            Leucas_lanata
            Leucas_lavandulifolia_1
            Leucas_lavandulifolia_2
            Leucas_martinicensis
            Leucas_masaiensis
            Leucas_milanjiana
            Leucas_minimifolia
            Leucas_mwingensis
            Leucas_neufliseana
            Leucas_nubica
            Leucas_oligocephala
            Leucas_sexdentata
            Leucas_spiculifolia
            Leucas_stachydiformis
            Leucas_urticifolia
            Leucas_usagarensis
            Leucas_virgata
            Leucas_volkensii_var_parviflora
            Leucas_zeylanica
            Marrubium_vulgare
            Moluccella_laevis
            Moluccella_spinosa
            Otostegia_aucheri_1
            Otostegia_aucheri_2
            Otostegia_erlangeri
            Otostegia_fruticosa
            Otostegia_hildebrandtii
            Otostegia_integrifolia
            Otostegia_limbata
            Otostegia_modesta_1
            Otostegia_modesta_2
            Otostegia_persica
            Otostegia_somala
            Otostegia_tomentosa_ssp_ambigens_1
            Otostegia_tomentosa_ssp_ambigens_2
            Phlomis_milingensis
            Phlomis_purpurea
            Roylea_cinerea
            Scutellaria_hirta
            Scutellaria_sieberi
            ;
END;

BEGIN TREES;
      TITLE Tb9485;
      LINK TAXA = 'combined trnL-trnF region, trnS-trnG spacer, and rps16 intron';
      TREE Fig._1 = [&R] ((Scutellaria_hirta,Scutellaria_sieberi),(((Roylea_cinerea,(((Marrubium_vulgare,(Ballota_acetabulosa,Ballota_pseudodictamnus)),((Otostegia_aucheri_1,Otostegia_aucheri_2),(Moluccella_laevis,Moluccella_spinosa))),(Lamium_tomentosum,Lamium_album_ssp_crinitum))),((((Leucas_aequistylosa,(Leucas_stachydiformis,(((Leucas_nubica,(Leucas_usagarensis,((((Acrotome_hispida,(Acrotome_pallescens,(Leucas_sexdentata,(Acrotome_angustifolia,(Acrotome_inflata_2,(Acrotome_fleckii,Acrotome_inflata_1)))))),((Leonotis_myrothamnifolia,(Leonotis_myricifolia,((Leonotis_grandis,(Leucas_grandis,(Leonotis_nepetifolia_var_nepetifolia,Leonotis_nepetifolia_var_africana))),(Leonotis_ocymifolia_var_schinzii,(((Leonotis_ocymifolia_var_raineriana_1,Leucas_deflexa),(Leucas_densiflora,Leucas_volkensii_var_parviflora)),(Leucas_bracteosa,Leucas_martinicensis)))))),(Leonotis_leonurus,((Leucas_urticifolia,(Leucas_calostachys,(Leonotis_ocymifolia_var_raineriana_2,Leucas_masaiensis))),(Leucas_glabrata_var_glabrata,(Leucas_capensis,(Leucas_argentea,(Leucas_glabrata,Leucas_milanjiana)))))))),(Leucas_neufliseana,Leucas_oligocephala)),(Leucas_minimifolia,(Leucas_discolor,(Leucas_abyssinica_var_argyrophylla,(Leucas_cuneifolia,(Leucas_mwingensis,(Leucas_inflata,(Leucas_abyssinica_var_brachycalyx,(Leucas_abyssinica_var_sidamoensis,Leucas_jamseii))))))))))),(Otostegia_hildebrandtii,(Otostegia_fruticosa,(Otostegia_tomentosa_ssp_ambigens_1,(Otostegia_modesta_2,(Otostegia_modesta_1,(Otostegia_erlangeri,Otostegia_tomentosa_ssp_ambigens_2))))))),((Leucas_flagellifera,Leucas_spiculifolia),(Leucas_kishenensis,Leucas_virgata))))),(Isoleucas_arabica,Otostegia_somala)),((Leucas_ciliata,(Leucas_aspera,(Leucas_lavandulifolia_2,(Leucas_lavandulifolia_1,Leucas_zeylanica)))),(Leucas_biflora,(Leucas_lanata,Leucas_decemdentata)))),(Otostegia_persica,(Otostegia_integrifolia,Otostegia_limbata)))),(Phlomis_milingensis,Phlomis_purpurea)));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5879]


END;