#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on July 11, 2025; 14:52 GMT TreeBASE (cc) 1994-2008 Study reference: Kolicka M., Dabert M., Olszanowski Z., & Dabert J. 2020. Sweet or salty? The origin of freshwater gastrotrichs (Gastrotricha, Chaetonotida) revealed by molecular phylogenetic analysis. Cladistics, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S25116] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=183; TAXLABELS Arenotus_strixinoi Aspidiophorus_ophiodermus Aspidiophorus_paramediterraneus Aspidiophorus_polystictos Aspidiophorus_sp._3_TK_2012 Aspidiophorus_sp._nov._2_MOTU_48 Aspidiophorus_sp._nov._3_MOTU_34 Aspidiophorus_sp._nov._3_MOTU_63 Aspidiophorus_sp._nov._MOTU_35 Aspidiophorus_tentaculatus Aspidiophorus_tetrachaetus Bifidochaetus_arcticus Bifidochaetus_sp._MOTU_61 Cephalionotus_kisielewskii Chaetonotus_acanthocephalus Chaetonotus_acanthodes_1 Chaetonotus_acanthodes_2 'Chaetonotus acanthodes MOTU_59' Chaetonotus_aemilianus Chaetonotus_aff._bombardus_MOTU_12 Chaetonotus_aff._bombardus_MOTU_13 Chaetonotus_aff._bombardus_MOTU_14 Chaetonotus_aff._bombardus_MOTU_16 Chaetonotus_aff._daphnes_2_MOTU_64 Chaetonotus_aff._daphnes_MOTU_46 Chaetonotus_aff._daphnes_MOTU_47 Chaetonotus_aff._euhystrix_MOTU_11 Chaetonotus_aff._euhystrix_MOTU_62 Chaetonotus_aff._gelidus_2_MOTU_29 Chaetonotus_aff._gelidus_MOTU_20 Chaetonotus_aff._gelidus_MOTU_22 Chaetonotus_aff._gelidus_MOTU_23 Chaetonotus_aff._gelidus_MOTU_27 Chaetonotus_aff._haideri_2_MOTU_50 Chaetonotus_aff._haideri_3_MOTU_51 Chaetonotus_aff._haideri_MOTU_52 Chaetonotus_aff._haideri_MOTU_53 Chaetonotus_aff._maximus_MOTU_40 Chaetonotus_aff._maximus_MOTU_49 Chaetonotus_aff._microchaetus_MOTU_45 Chaetonotus_aff._oculiefer_MOTU_24 Chaetonotus_aff._oculifer_MOTU_30 Chaetonotus_aff._oculifer_MOTU_31 Chaetonotus_aff._persimilis_6_2 Chaetonotus_aff._persimilis_MOTU_1 Chaetonotus_aff._persimilis_MOTU_2 Chaetonotus_aff._persimilis_MOTU_6 Chaetonotus_aff._sphagnophilus_MOTU_26 Chaetonotus_aff._subtilis_3_MOTU_37 Chaetonotus_aff._subtilis_3_MOTU_58 Chaetonotus_aff._subtilis_4_MOTU_36 Chaetonotus_aff._subtilis_5_MOTU_39 Chaetonotus_aff._subtilis_MOTU_57 Chaetonotus_antrumus Chaetonotus_armatus Chaetonotus_bisacer Chaetonotus_bombardus Chaetonotus_borealis 'Chaetonotus borealis MOTU_5' Chaetonotus_cf_maximus_1 Chaetonotus_cf_maximus_2 Chaetonotus_cf._dispar Chaetonotus_cf._hornsundi_MOTU_8 Chaetonotus_cf._hystrix Chaetonotus_cf._laroides Chaetonotus_cf._novenarius Chaetonotus_cf._oculifer Chaetonotus_cf._similis_1 Chaetonotus_cf._similis_2 Chaetonotus_cf._sphagnophilus Chaetonotus_daphnes Chaetonotus_gelidus Chaetonotus_heideri_1 Chaetonotus_heideri_2 Chaetonotus_heteracanthus 'Chaetonotus heterolepidodermus MOTU_28' 'Chaetonotus heterolepidodermus MOTU_28_2' Chaetonotus_hornsundi 'Chaetonotus hornsundi MOTU_21' 'Chaetonotus hornsundi MOTU_9' Chaetonotus_hystrix_1 Chaetonotus_jaceki 'Chaetonotus jaceki MOTU_17' 'Chaetonotus jaceki MOTU_18' Chaetonotus_laroides Chaetonotus_mariae Chaetonotus_microchaetus Chaetonotus_neptuni_1 Chaetonotus_neptuni_2 Chaetonotus_persimilis Chaetonotus_persimilis_2 'Chaetonotus persimilis MOTU_10' 'Chaetonotus persimilis MOTU_25' 'Chaetonotus persimilis MOTU_3' 'Chaetonotus persimilis MOTU_3_2' 'Chaetonotus persimilis MOTU_4' 'Chaetonotus persimilis MOTU_7' Chaetonotus_polyspinosus Chaetonotus_schultzei Chaetonotus_semovitus Chaetonotus_similis Chaetonotus_sp._1 Chaetonotus_sp._1_aff_armatus Chaetonotus_sp._2 Chaetonotus_sp._2_MOTU_19 Chaetonotus_sp._2_MOTU_38 Chaetonotus_sp._2_aff_acanthocephalus Chaetonotus_spec._A Chaetonotus_subtilis 'Chaetonotus subtilis MOTU_56' Chaetonotus_uncinus Dactylopodola_mesotyphle Dasydytes_Prodasydytes_elongatus_1 Dasydytes_Prodasydytes_elongatus_2 Dasydytes_Prodasydytes_papaveroi Dasydytes_carvalhoae Diuronotus_aspetos Draculiciteria_tesellata_MT63 Draculiciteria_tesellata_TK142 Gnathostomula_armata Gnathostomula_axi Halichaetonotus_aculifer Halichaetonotus_euromarinus Halichaetonotus_paradoxus Halichaetonotus_schromi Halichaetonotus_sp._2 Halichaetonotus_sp._4 Haltidytes_squamosus Heterolepidoderma_acidophilum Heterolepidoderma_aff._ocellatum_MOTU_32 Heterolepidoderma_loricatum Heterolepidoderma_macrops Heterolepidoderma_ocellatum Heterolepidoderma_sinus Heterolepidoderma_sp._1 Heterolepidoderma_sp._2 Heterolepidoderma_sp._3 Heteroxenotrichula_squamosa Ichthydium_skandicum 'Ichthydium skandicum MOTU_54' 'Ichthydium skandicum MOTU_55' Ichthydium_spec._nov._MOTU_33 Ichthydium_squamigerum Kijanebalola_devestiva_1 Kijanebalola_devestiva_2 Lepidochaetus_brasilense Lepidochaetus_brasilense_2 Lepidochaetus_zelinkai_1 Lepidochaetus_zelinkai_2 Lepidodermella_aff._squamata_MOTU_44 Lepidodermella_cf._polaris_MOTU_41 Lepidodermella_intermedia Lepidodermella_minor_minor Lepidodermella_polaris 'Lepidodermella polaris MOTU_42' 'Lepidodermella polaris MOTU_43' Lepidodermella_squamata_1 Lepidodermella_squamata_2 Macrodasys_sp._1 Megadasys_sp._1 Musellifer_delamarei Musellifer_reichardti Neodasys_chaetonotoideus Neodasys_cirritus Neodasys_uchidai Neogossea_acanthocolla Neogossea_antennigera_1 Neogossea_antennigera_2 Ornamentula_paraensis Polymerurus_nodicaudus_1 Polymerurus_nodicaudus_2 Polymerurus_nodicaudus_3 Polymerurus_nodicaudus_4 Polymerurus_rhomboides_1 Polymerurus_rhomboides_2 Polymerurus_rhomboides_3 Stylochaeta_fusiformis Stylochaeta_scirtetica Xenotrichula_cf._intermedia Xenotrichula_intermedia Xenotrichula_punctata Xenotrichula_sp._TK121 Xenotrichula_velox ; END; BEGIN TREES; TITLE Gastrotricha; LINK TAXA = Taxa; TREE PAUP_1 = [&R] ((Gnathostomula_axi:0.203284,Gnathostomula_armata:0.353679):4.5428655,(((Xenotrichula_sp._TK121:0.94357,Xenotrichula_punctata:0.865726):2.099708,(Xenotrichula_velox:2.539125,(Heteroxenotrichula_squamosa:2.655139,(Xenotrichula_intermedia:0.55632,Xenotrichula_cf._intermedia:0.484706):1.063133):0.106143):0.380312):0.828337,(((Dactylopodola_mesotyphle:1.734956,(Macrodasys_sp._1:2.188232,Megadasys_sp._1:1.687583):0.272129):0.451407,(Neodasys_cirritus:0.386116,(Neodasys_chaetonotoideus:0.061954,Neodasys_uchidai:0.014856):0.031492):1.629964):0.848076,(Diuronotus_aspetos:1.842387,((Draculiciteria_tesellata_MT63:0.100916,Draculiciteria_tesellata_TK142:0.100294):1.230714,((Aspidiophorus_polystictos:0.271648,(Aspidiophorus_paramediterraneus:0.418323,(Aspidiophorus_sp._3_TK_2012:0.177038,Aspidiophorus_tentaculatus:0.093098):0.155269):0.083558):0.186798,(Halichaetonotus_schromi:0.109459,(((Cephalionotus_kisielewskii:0.229896,Heterolepidoderma_loricatum:0.216536):6.69999E-6,((Heterolepidoderma_sp._1:0.006615,Heterolepidoderma_sinus:0.049417):0.201835,((Halichaetonotus_sp._4:0.094057,Halichaetonotus_euromarinus:0.051871):0.071143,(Halichaetonotus_aculifer:0.041051,(Halichaetonotus_paradoxus:0.045454,Halichaetonotus_sp._2:0.072887):0.017381):0.035687):0.013095):0.017464):0.042049,(((((Bifidochaetus_arcticus:0.005643,Bifidochaetus_sp._MOTU_61:0.002363):0.158564,(Lepidodermella_minor_minor:0.05762,Lepidodermella_intermedia:0.088302):0.057499):0.023401,((Lepidochaetus_brasilense:0.066111,Lepidochaetus_brasilense_2:0.06371):0.030659,(Lepidochaetus_zelinkai_1:0.00418,Lepidochaetus_zelinkai_2:0.00398):0.054357):0.054823):0.012274,((((Chaetonotus_acanthocephalus:0.061209,Chaetonotus_sp._2_aff_acanthocephalus:0.077583):0.080707,(Chaetonotus_armatus:0.093303,Chaetonotus_mariae:0.194639):0.015091):0.024478,(Chaetonotus_sp._2:0.095864,(Chaetonotus_acanthodes_2:0.0,(Chaetonotus_acanthodes_1:0.0,'Chaetonotus acanthodes MOTU_59':0.0):0.0):0.123748):0.021296):0.007726,((Chaetonotus_heideri_1:0.094502,Chaetonotus_heideri_2:0.065837):0.070846,((Chaetonotus_sp._1_aff_armatus:0.089761,(Chaetonotus_aff._haideri_MOTU_53:0.002161,Chaetonotus_aff._haideri_MOTU_52:0.008053):0.063454):0.017392,(Aspidiophorus_tetrachaetus:0.119464,('Ichthydium skandicum MOTU_54':0.04702,(Ichthydium_skandicum:9.52E-4,'Ichthydium skandicum MOTU_55':0.0):0.030262):0.06772):0.032047):0.001802):0.003828):0.022075):0.037455,((Chaetonotus_cf._dispar:0.071342,(Chaetonotus_neptuni_1:0.019017,Chaetonotus_neptuni_2:0.021099):0.110829):0.063743,((((Chaetonotus_cf._novenarius:0.061685,(Chaetonotus_sp._2_MOTU_38:0.009183,(Chaetonotus_aff._subtilis_4_MOTU_36:0.003203,Chaetonotus_aff._subtilis_3_MOTU_37:0.003288):0.004089):0.046716):0.028978,(Chaetonotus_aff._maximus_MOTU_40:0.079787,(Chaetonotus_persimilis_2:7.05E-4,(Chaetonotus_aff._sphagnophilus_MOTU_26:0.037307,'Chaetonotus persimilis MOTU_25':0.00535):0.0):0.042425):0.036347):0.007235,((Chaetonotus_schultzei:0.143019,(Aspidiophorus_sp._nov._2_MOTU_48:0.067189,(Aspidiophorus_sp._nov._3_MOTU_63:0.107599,(Aspidiophorus_sp._nov._3_MOTU_34:0.055187,Aspidiophorus_sp._nov._MOTU_35:0.042162):0.056223):0.018228):0.071323):0.01028,((Chaetonotus_semovitus:0.002741,Chaetonotus_uncinus:0.002614):0.148592,(((Chaetonotus_aff._subtilis_3_MOTU_58:0.023018,Chaetonotus_aff._subtilis_MOTU_57:0.02772):0.082424,(Chaetonotus_cf._sphagnophilus:0.052276,(Chaetonotus_subtilis:0.00244,'Chaetonotus subtilis MOTU_56':7.64E-4):0.044645):0.015526):0.004678,((((Kijanebalola_devestiva_1:0.0,Kijanebalola_devestiva_2:0.0):0.101253,(Neogossea_acanthocolla:0.052208,Neogossea_antennigera_2:0.005326):6.57782E-8):0.006744,(Neogossea_antennigera_1:0.00538,(Polymerurus_nodicaudus_1:0.071563,(Polymerurus_rhomboides_1:1.9304E-6,Chaetonotus_aff._persimilis_6_2:0.031113):0.135498):0.125499):3.2103E-7):0.038806,((Chaetonotus_heteracanthus:0.092992,(Chaetonotus_bisacer:0.168918,((Polymerurus_rhomboides_3:0.065879,Polymerurus_rhomboides_2:0.057088):0.05,(Polymerurus_nodicaudus_2:0.061993,(Polymerurus_nodicaudus_4:0.045877,Polymerurus_nodicaudus_3:0.072499):0.022842):0.040373):0.030601):0.003688):0.042277,((Dasydytes_carvalhoae:0.195123,(Ornamentula_paraensis:0.090519,Haltidytes_squamosus:0.046761):0.029612):0.012465,((Stylochaeta_fusiformis:0.031144,Stylochaeta_scirtetica:0.041129):0.040915,(Dasydytes_Prodasydytes_papaveroi:0.063144,(Dasydytes_Prodasydytes_elongatus_1:0.022595,Dasydytes_Prodasydytes_elongatus_2:0.019399):0.10432):0.057076):0.02386):0.026712):0.006468):0.027685):0.025798):0.021782):0.009036):0.007466,(((Chaetonotus_spec._A:0.062558,(Chaetonotus_polyspinosus:0.045853,Arenotus_strixinoi:0.066294):0.008744):0.065659,((Chaetonotus_aff._daphnes_2_MOTU_64:0.054144,Heterolepidoderma_sp._3:0.078386):0.024,(Aspidiophorus_ophiodermus:0.058609,(Chaetonotus_aff._daphnes_MOTU_46:0.04317,(Chaetonotus_daphnes:0.010127,Chaetonotus_aff._daphnes_MOTU_47:0.015808):0.031188):0.02373):0.007568):0.01595):0.032196,((Heterolepidoderma_macrops:0.107637,((Chaetonotus_laroides:8.77E-4,Chaetonotus_cf._laroides:0.0):0.068603,((Chaetonotus_cf._similis_2:0.044225,(Chaetonotus_similis:0.025132,Chaetonotus_cf._similis_1:0.024976):0.035127):0.041464,((Chaetonotus_aff._maximus_MOTU_49:0.022439,(Chaetonotus_cf_maximus_1:0.025464,Chaetonotus_cf_maximus_2:0.022109):0.00982):0.036987,((Chaetonotus_microchaetus:0.025209,Chaetonotus_aff._microchaetus_MOTU_45:0.036126):0.029223,(Lepidodermella_squamata_1:0.015733,(('Lepidodermella polaris MOTU_42':7.84E-4,Lepidodermella_cf._polaris_MOTU_41:0.004294):0.011742,((Lepidodermella_polaris:0.002521,'Lepidodermella polaris MOTU_43':4.21E-4):0.008521,(Lepidodermella_aff._squamata_MOTU_44:0.018916,Lepidodermella_squamata_2:0.016379):0.003218):0.001258):0.018146):0.118766):0.0324):0.008557):0.012225):0.009476):0.006435,(((((Chaetonotus_aff._euhystrix_MOTU_62:0.037014,Chaetonotus_aff._euhystrix_MOTU_11:0.02123):0.014913,(Chaetonotus_cf._hystrix:0.038809,Chaetonotus_aemilianus:0.039529):0.017128):0.008491,(Chaetonotus_aff._bombardus_MOTU_16:0.049364,(Chaetonotus_bombardus:0.071502,(Chaetonotus_aff._bombardus_MOTU_14:0.017158,(Chaetonotus_aff._bombardus_MOTU_13:0.021661,Chaetonotus_aff._bombardus_MOTU_12:0.023095):0.017491):0.043583):0.018746):0.02679):0.006125,((Chaetonotus_aff._persimilis_MOTU_1:0.017573,Chaetonotus_aff._persimilis_MOTU_2:0.012099):0.005381,(Chaetonotus_aff._persimilis_MOTU_6:0.011001,(Chaetonotus_hystrix_1:0.002618,((Chaetonotus_hornsundi:0.00133,'Chaetonotus hornsundi MOTU_9':4.31E-4):0.017137,(('Chaetonotus persimilis MOTU_10':0.007648,(Chaetonotus_borealis:0.003388,'Chaetonotus borealis MOTU_5':0.0):0.002373):0.005716,((Chaetonotus_cf._hornsundi_MOTU_8:0.005902,(Chaetonotus_persimilis:0.00498,'Chaetonotus persimilis MOTU_3':0.002309):8.93E-4):0.002516,('Chaetonotus persimilis MOTU_3_2':0.001462,('Chaetonotus persimilis MOTU_7':0.006063,'Chaetonotus persimilis MOTU_4':0.0):0.004418):0.001571):5.22E-4):0.003215):0.0):0.001328):0.018275):0.041671):0.112268,((Chaetonotus_sp._2_MOTU_19:0.066208,((Chaetonotus_antrumus:0.012788,Chaetonotus_sp._1:0.011905):0.001525,('Chaetonotus jaceki MOTU_18':0.008356,(Chaetonotus_jaceki:0.002145,'Chaetonotus jaceki MOTU_17':0.002456):0.003271):0.006347):0.061449):0.016219,(((Ichthydium_squamigerum:0.024246,Ichthydium_spec._nov._MOTU_33:0.047851):0.077959,((Chaetonotus_aff._gelidus_MOTU_20:0.019047,(Chaetonotus_gelidus:0.010064,'Chaetonotus hornsundi MOTU_21':0.008512):0.009212):0.007201,(Chaetonotus_aff._gelidus_MOTU_22:0.020357,(Chaetonotus_aff._haideri_2_MOTU_50:0.089681,((Chaetonotus_aff._haideri_3_MOTU_51:0.0,Musellifer_reichardti:3.252922):0.086933,(Chaetonotus_aff._oculifer_MOTU_30:0.0,Musellifer_delamarei:4.11349):0.173705):0.002296):0.138296):1.50734E-7):0.032328):0.041758,((Chaetonotus_aff._oculiefer_MOTU_24:0.002548,(Chaetonotus_aff._gelidus_MOTU_23:0.001082,Heterolepidoderma_sp._2:0.024246):0.003994):0.05621,((Chaetonotus_aff._oculifer_MOTU_31:0.030003,Chaetonotus_cf._oculifer:0.033497):0.036999,((Chaetonotus_aff._subtilis_5_MOTU_39:0.00558,Chaetonotus_aff._gelidus_2_MOTU_29:0.004628):0.032424,((Chaetonotus_aff._gelidus_MOTU_27:0.009384,('Chaetonotus heterolepidodermus MOTU_28':4.4E-4,'Chaetonotus heterolepidodermus MOTU_28_2':0.0):0.015546):0.013707,(Heterolepidoderma_acidophilum:0.021366,(Heterolepidoderma_ocellatum:0.012414,Heterolepidoderma_aff._ocellatum_MOTU_32:0.019221):0.00881):0.027493):0.006108):0.014087):0.017956):0.029805):0.029385):0.00971):0.00529):0.003162):0.020925):0.00985):0.009113):0.039531):0.010267):0.081463):1.72507):0.261787):0.073172):0.059141):4.5428655); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr125808] END;