#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on July 09, 2025; 18:00 GMT TreeBASE (cc) 1994-2008 Study reference: Lambkin C., Sinclair B., Pape T., Courtney G.W., Skevington J., Meier R., Yeates D., Blagoderov V., & Wiegmann B.M. 2013. The phylogenetic relationships among infraorders and superfamilies of Diptera based on morphological evidence. Systematic Entomology, 38: 164-179. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S13373] [ The following blocks are input data for analysis step 10449 ] BEGIN TAXA; TITLE TaxaForAnalysisStep10449; DIMENSIONS NTAX=46; TAXLABELS Acrosathe_novella Anopheles_gambiae Arachnocampa_flava Asilus_crabroniformis Bibio_longipes Bittacus_mastrillii Bombylius_major Bradysia_tilicola Ceratitis_capitata Chamaepsila_hennigi Chironomus_tepperi Clogmia_albipunctata Coboldia_fuscipes Cochliomyia_macellaria Ctenocephalides_felis Delia_radicum Drosophila_melanogaster Edwardsina_gigantea Episyrphus_balteatus Exeretonevra_angustifrons Exorista_larvarum Glossina_morsitans Haematopota_pluvialis Hermetia_illucens Lonchoptera_lutea Mayetiola_destructor Megaselia_scalaris Micropteryx_calthella Minettia_flaveola Musca_domestica Nannochorista_philpotti Nymphomyia_dolichopeza Ogcodes_basalis Oreogeton_obscurus Physocephala_marginata Phytomyza_ilicicola Protoplasa_fitchii Sarcophaga_bullata Scathophaga_stercoraria Sepsis_cynipsea Spelobia_bifrons Sylvicola_fenestralis Teleopsis_dalmanni Thaumatomyia_glabra Tipula_abdominalis Trichocera_brevicornis ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M15322] TITLE FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names; LINK TAXA = TaxaForAnalysisStep10449; DIMENSIONS NCHAR=371; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7" MISSING=? GAP= -; CHARSTATELABELS 1 Head_capsule / absent present, 2 head_capsule / not_cone_shaped_ Cone_shaped_, 3 Metacephalic_rods / absent present, 4 paired_lateral_Metacephalic_rods / absent present, 5 single_medial_Metacephalic_rod / absent present, 6 Metacephalic_rods / continuous_with_head_capsule hinged, 7 Metacephalic_rods / straight spatulate_at_tip, 8 Posterodorsal_cranial_margin / incapable_of_intermolt_cuticle_deposition capable_of_intermolt_cuticle_deposition, 9 Postgenae / forming_a_postgenal_bridge narrowly_in_contact_or_separated_by_a_narrow_membranous_area widely_separated, 10 Instar_I_frontal_sclerite / contiguous_with_surrounding_cranial_sclerites separated_from_surrounding_cranial_sclerites_by_a_band_of_membranous_cuticle, 11 Head / not_retracted_into_thorax retracted_into_thorax, 12 Head / not_invaginated_into_thorax 'partly_invaginated_into_thorax_(not_closed_dorsally)_' invaginated_into_thorax_forming_complete_atrium, 13 Instar_3_Intermediate_sclerite / fused_with_vertical_plate not_fused_with_vertical_plate, 14 intermediate_sclerite / V_shaped H_shaped 'elongated,_with_the_longitudinal_bars_almost_completely_fused_due_to_an_expanded_ventral_bridge', 15 tentorial_bridge / present absent, 16 Antennal_segments / none one two three_or_more, 17 Length_of_first_instar_larval_antenna / short_about_as_long_as_maximum_diameter long_several_times_longer_than_maximum_diameter, 18 Antenna / not_prehensile prehensile, 19 'antennal structure, ring and peg' / absent present, 20 Larval_stemmata / more_than_five five four three two one none, 21 'Bolwigs organ, internalized in optical depression' / absent present, 22 Pigment_of_adult_eye / appearing_in_the_pupa_after_the_larval_stage developing_precociously_becoming_conspicuous_as_early_as_the_second_instar_and_always_well_developed_by_the_last_instar, 23 Frontoclypeal_apotome / present absent, 24 Frontoclypeal_apotome / triangularly_V_shaped broadly_U_shaped, 25 '''Labrum-like'' blade in first instar larva' / present _absent, 26 '''Labrum-like'' blade in final instar larva' / extending_beyond_base_of_mandible 'reduced,_ending_at_base_of_mandible^n', 27 Labrum / dorsoventrally_flattened conical, 28 Brushes_on_ventral_surface_of_labrum / absent well_developed complex, 29 Premandibles / absent present, 30 Premandible / a_simple_unadorned_sclerite with_a_comb_like_row_of_evenly_spaced_anteriorly_directed_teeth, 31 Invaginated_premandible_apodeme / absent present, 32 Premandible / mainly_external_with_small_invaginated_apodeme_for_insertion_of_labral_retractor_muscles with_external_portion_and_invaginated_internal_portion_bearing_separate_insertions_of_two_labral_retractor_muscles_and_capable_of_forceps_like_action mainly_internal_serving_almost_entirely_as_a_point_of_insertion_of_labral_retractor_muscles_with_small_remnant_of_external_sclerite, 33 Premandible_with_tongs_like_or_spike_like_external_portion / absent present, 34 Torma / present absent, 35 Torma / fused_with_the_dorsal_labral_sclerite articulated, 36 Mandibular_rotation / horizontal oblique_or_vertical, 37 Mandible / dicondylous Cyclorrhaphan_monocondylous, 38 Mandibular_epicondyle / near_junction_of_genae_and_frontoclypeal_apotome on_tentorial_phragma, 39 Mandible / uniformly_sclerotized_with_apex_and_base_fused_solidly apex_and_base_separated_by_a_zone_of_membranous_cuticle separated_into_two_distinct_components subdivided_into_more_than_two_articles, 40 Basal_mandibular_sclerite_from_posterior_view / broadly_inverted_U_shaped laterally_compressed, 41 Subapical_adoral_surface_of_mandible / bare with_comb_like_row_of_long_curved_setae, 42 Apex_of_mandible / without_comb with_multi_toothed_comb, 43 Mandibular_brush / absent present, 44 Mandibular_brush_position / on_membrane_along_lateral_arm_of_basal_sclerite on_horizontal_rod on_vertical_rod, 45 Mandibular_chela / absent present, 46 Mandible / without_poison_canal with_poison_canal_a_groove_or_canal_in_anterodorsal_position, 47 Inner_face_of_mandibular_hook / without_groove with_a_groove_that_forms_canal_when_mandibular_hooks_adpressed_, 48 Egg_burster_on_frontal_sclerite_of_first_instar / present absent, 49 Mandiblular_prostheca / present absent, 50 Mandiblular_prostheca / on_articulated_lobe arising_directly_from_adoral_surface_of_mandible, 51 Cardo / free_from_anterior_margin_of_head_capsule fused_with_or_closely_appressed_to_anterior_margin_of_head_capsule, 52 Antennal_and_maxillary_lobes / separate fused, 53 Maxillary_palpus / palpiform flush_with_the_surface_of_the_maxilla_and_recognizable_only_by_a_ring_surrounding_the_sensilla, 54 Maxillary_palpus / separate_from_mandible weakly_fused_to_basal_mandibular_sclerite_dorsal_hook_absent completely_fused_to_basal_mandibular_sclerite, 55 Salivary_pump / small_slightly_larger_than_duct greatly_enlarged_sac_like_chamber, 56 Cibarial_filter / present absent, 57 Cibarium / simple 'comb-like,_transverse_filaments_' 'series_of_transverse_folds_with_"comb"-like_projection_used_to_filter_particles_from_waters' 'T-shaped_ridges' 'cibarial_filter_in_basal_sclerite_without_t-ridges', 58 Pharyngeal_grinding_mill / absent present, 59 Prementohypopharyngeal_apparatus / absent present, 60 Prementohypopharyngeal_apparatus / not_connected_to_paraclypeal_phragma dorsolateral_corner_connected_via_cibarial_bar_to_paraclypeal_phragma, 61 Sclerotised_postmentum / present absent, 62 Postmentum / a_simple_unadorned_sclerite with_serrate_anterior_margin, 63 Postmentum / a_separate_plate fused_to_postgenae, 64 Parastomal_bars_in_last_instar / present absent, 65 Thoracic_segments / not_enlarged swollen_distinctly_broader_than_head_and_abdomen, 66 Thoracic_legs / articulated_appendages_in_at_least_one_larval_instar 'vestigial,_never_articulated,_represented_in_at_least_one_larval_instar_by_Keilin?s_organ' absent_in_all_larval_instars, 67 First_instar_prothoracic_prolegs / absent present, 68 Cuticle_encrusted_with_warts_of_calcium_carbonate / absent present, 69 Free_abdominal_segments_excluding_anal_division / 9_segments 8_segments 7_segments 6_segments 5_or_fewer_segments, 70 Body_segments / without_secondary_divisions_into_annuli_or_intercalary_rings with_complete_secondary_divisions, 71 Elongate_eversible_crochet_tipped_prolegs_on_abdominal_segments_one_to_seven / absent one_pair two_or_more_pairs, 72 Abdomen_with_single_proleg_on_segment_one / absent present, 73 'First instar larva with terminal pair of sac-like vesicles on anal division' / absent present, 74 Crotchet_tipped_prolegs_on_anal_division / absent present, 75 'Anal division with fan-like row of long midventral setae' / absent present, 76 Base_of_midventral_setae_on_anal_division / simple transverse_T_shaped, 77 Anal_division_with_sclerotised_dorsal_plates_surrounding_the_spiracles_and_ending_in_a_pair_of_hook_like_processes / absent present, 78 Anal_papillae / present absent, 79 Number_of_Malpighian_tubules / six five four three_or_less, 80 Respiratory_system_of_full_grown_larva / polypneustic_ie_holo_peri_or_hemipneustic oligopneustic_ie_amphi_or_metapneustic apneustic, 81 Anterior_spiracles / single_opening multiple_openings, 82 Arrangement_of_multiple_papilla_in_anterior_spiracles / in_a_row_ arranged_along_a_stalk arranged_on_bulb, 83 Spiracles_on_anal_division / type_I type_II type_III, 84 Position_of_posterior_abdominal_spiracle / on_anal_division on_abdominal_segment_8, 85 Position_of_posterior_abdominal_spiracle / lateral dorsal_or_terminal, 86 Posterior_abdominal_spiracles_in_a_cavity / absent present, 87 'Stigmatophore (siphon bearing posterior abdominal spiracles) longer than diameter of spiracle' / absent Present__longer_than_diameter_of_spiracle_, 88 Each_posterior_spiracle_surrounded_by_four_tufts_of_flattened_cuticular_outgrowths_101 / absent present, 89 Openings_of_each_posterior_tracheal_trunk_of_last_instar / single two_openings three_or_more_openings, 90 Reproduction / eggs_not_incubated_within_female eggs_incubated_within_female, 91 Reproduction_with_incubated_eggs / several_larvae 'macrolarviparous,_with_single_larva', 92 Reproduction_with_incubated_eggs / eggs_hatch_during_deposition 'eggs_hatch_in_female,_larvae_nourished_by_special_secretion.', 93 Number_of_instars / more_than_three three two, 94 Mode_of_pupation / pupa_free pupation_within_puparium_formed_from_last_larval_cuticle, 95 Cyclorrhaphan_puparium__cleavage_lines / mid_dorsal_longitudinal transverse_dorsal horizontal_cleavage_line_on_each_side H_shaped_restricted_to_abdominal_tergites_1_to_3 H_shaped_through_mesothoracic_and_abdominal_tergites_1 'T-shaped_through_meosthoracic_and_abdominal_tergites', 96 Cyclorrhaphan_puparium_anterior_spiracles / anterior_spiracles_penetrating_puparium_as_puparial_horns no_puparial_horns, 97 Body / tergites_and_sternites_darkly_sclerotized tergites_darkly_sclerotized_and_sternites__lightly_sclerotised tergites_and_sternites_lightly_sclerotised, 98 Mandibles / decticous adecticous, 99 Ovoid_Ventrolateral_adhesive_discs_on_abdomenal_segments / absent present, 100 'plastron-bearing gill' / absent present, 101 'plastron-bearing gill' / expands_and_hardens_after_the_larval_pupal_ecdysis acquires_its_final_volume_and_shape_before_the_larval_pupal_ecdysis, 102 Developing_adult_wing / an_unexpanded_version_of_its_final_form_expanded_only_after_emergence reaching_its_final_size_and_shape_before_emergence_and_unfolding_and_functional_immediately_at_emergence, 103 Metathoracic_leg_sheath / extended_beyond_wing_sheath_parallel_to_sheaths_of_other_two_legs_or_superposed bent_in_an_S_shape_concealed_beneath_wing_sheath, 104 Terminus_of_abdomen_with_pair_of_articulated_membranous_paddles_bearing_supporting_midrib / absent present, 105 Abdominal_ganglia / more_than_six six five four three two one abdominal_and_thoracic_ganglia_fused, 106 Abdominal_ganglia / T2_fused_with_thorax T2_fused_with_abdomen, 107 Head_and_thorax_setae / some_setae_feather_like all_setae_simple, 108 Eyes_configuration_male / dichoptic holoptic_separated_by_no_more_than_width_of_median_ocellus 'intermediate,_clearly_narrower_than_in_female_but_separated_by_more_than_width_of_median_ocellus?', 109 Eyes_configuration_female / dichoptic holoptic_separated_by_no_more_than_width_of_medial_ocellus, 110 Ocelli / three two one vestigial_or_absent, 111 Ocelli_on_raised_plate / absent present, 112 Vertex / flat_not_modified concave, 113 Ocellar_setae_or_bristles / absent present, 114 Inner_vertical_bristles / absent present, 115 Outer_vertical_bristles / absent present, 116 Postocellar_bristles / absent present, 117 Postocellar_bristles / divergent parallel convergent, 118 Ptilinal_fissure / absent present, 119 Frons_differentiated_into_frontal_vitta_and_lateral_fronto_orbital_plates / absent present, 120 Orbital_bristles / absent present, 121 Orbital_bristles / reclinate proclinate medioclinate lateroclinate, 122 Anterior_frontal_bristles / absent present, 123 Anterior_frontal_bristles / reclinate proclinate medioclinate lateroclinate, 124 Interfrontal_bristles / absent present, 125 Face_with_mystax / absent present, 126 Vibrissa / absent present, 127 Adult_clypeus / concave_or_flattened bulbous_and_projecting, 128 Adult_clypeus / connected_to_face widely_separated_from_lower_margin_of_face_by_membrane, 129 Clypeus / not_divided divided_into_postclypeus_and_distal_anteclypeus, 130 Clypeal_ridge_position / articulated_with_labrofulcral_articulation_point distinctly_removed_from_labrofulcral_articulation_point, 131 Occipital_sclerite / with_well_developed_median_occipital_sclerite without, 132 Median_occipital_sclerite_OR_Median_occipital_plate / without_reinforcement_ridges with_reinforcement_ridges, 133 'Occipital internal thickenings - dorsal lateral apodemes arising above occipital condyles' / absent present, 134 Occipital_pockets / absent present_partial_or_complete, 135 Hypostomal_bridge / absent present, 136 Well_defined_row_of_postocular_setae / absent present, 137 Postocular_setae / equal_size_or_gradually_decreasing_in_size_from_the_vertex alternating_size, 138 Antennal_flagellomere_1 / not_decumbent decumbent, 139 Pedicel / short_length_equal_to_or_less_than_twice_scape_length elongate_length_more_than_twice_scape_length, 140 Pedicel / without_seam with_dorsal_seam, 141 Pedicel_flagellum_articulation / simple tip_of_pedicel_with_process_into_flagellum base_of_flagellum_with_process_into_pedicel, 142 Antennal_pedicel_of_male / not_enlarged enlarged_globular_and_much_wider_than_scape_because_of_greatly_enlarged_Johnstons_organ, 143 Flagellum_segments_not_including_arista_stylus / more_than_eight eight_to_five four_to_two one, 144 Antennal_flagellum_of_male / simple plumose setae_in_encircling_whorls_, 145 Antennae_consisting_of_enlarged_first_flagellomere_with_apically_elonged_1_to_3_segment_arista / absent present, 146 Arista_insertion / dorso_apically apically, 147 Arista / not_plumose plumose, 148 Mouthparts_and_digestive_track / well_developed atrophied, 149 Labrum_Adult_mouthparts / simple 'modified_for_piercing,_with_a_pair_of_apical_epipharyngeal_blades_', 150 Hyoid_sclerite / absent present, 151 Adult_hypopharynx / a_simple_blade_like_projection greatly_elongate_and_thread_like robust_bearing_rows_of_barb_like_hairs, 152 Galea / present absent, 153 Lacinia / blunt_apically 'blade-like_or_reduced', 154 Maxillary_palpus_segments / more_than_two_segments two_segments one_segment, 155 Anterior_mandibular_condyle / present absent, 156 Glossae_and_paraglossae_of_labium / present strongly_reduced, 157 Segments_of_labial_palps / 3_or_more 2_or_fewer, 158 Labellum / absent present, 159 Labellum / separate_from_prementum reduced_fused_with_prementum, 160 Pseudotracheae / absent present, 161 Pseudotracheae / opening_directly_into_mouth opening_into_one_or_two_main_trunks, 162 Prestomal_teeth / absent present, 163 Precervicale / absent present, 164 Prothorax / not_elongate elongate, 165 Postpronotal_callus / absent present, 166 Postpronotal_ie_humeral_setae / absent present, 167 Proepisternal_depression / absent present, 168 Proepisternum_ie_propleuron / flat with_an_anterior_carina, 169 Proepisternum_dorsally / bare setose, 170 Episternal_lobe / absent present, 171 Precoxal_bridge / absent present, 172 Scutum / _with_transverse_suture without_transverse_suture, 173 Scutum_with_transverse_suture / incomplete complete, 174 Scutal_transverse_suture / straight v_shaped, 175 Scutum_acrostichal_bristles / absent present, 176 Supra_alar_bristles / absent present, 177 Post_alar_bristles / absent present, 178 Dorsocentral_bristles / absent present, 179 Pleural_suture_of_mesothorax_between_episternum_and_epimeron / straight_or_slightly_sinuous bent_twice_at_almost_a_right_angle, 180 Posterior_anepisternal_bristles / absent present, 181 Katepisternal_bristles / present absent, 182 Katepisternal_bristles / 1_and_1_and_1_pattern 2_and_1_and_1_pattern 2_and_1_pattern 1_and_1_pattern 1_and_2_pattern '0_+_1_pattern', 183 Anepimeral_bristle / absent present, 184 Anepimeron / without_tubercle 'with_a_very_prominent_tubercle_from_which_a_bristle_arises_(Diopsid)', 185 Notopleuron / absent present, 186 Notopleural_bristles / absent two_or_more one_anterior one_posterior, 187 Posterior_notopleural_bristle / on_flat_surface on_notopleural_callus, 188 Postalar_wall / absent present, 189 Postalar_wall / bare setose, 190 Scutellum / not_modified greatly_enlarged, 191 Paired_scutellar_outgrowths / absent present, 192 Scutellar_bristles / absent present, 193 Scutellar_bristles / one_pair two_or_more_pairs, 194 Lower_surface_of_scutellum / setose_medially bare_medially, 195 Suprasquamal_ridge / absent present, 196 Suprasquamal_ridge / bare setose, 197 Dorsal_Anatergite_setosity / bare setose_ie_infrasquamal_setulae, 198 Subscutellum / not_convex_or_weakly_convex strongly_convex 'very_strongly_convex,_bulging', 199 Greater_ampulla / absent present, 200 Meron / a_small_triangular_sclerite large_round_as_fully_developed_behind_midcoxa_as_katepisternum_is_in_front_of_midcoxa, 201 Meral_setae / absent present, 202 Coxopleural_streak / present absent, 203 Metepisternum / broad narrowed_ventrally_intersegmental_suture_between_meso_and_metapleuron_converging_towards_metacoxal_condyle dorsal_and_ventral_sclerites_of_metepisternum_widely_separated_by_coming_together_of_intersegmental_suture_and_metapleural_suture_for_some_distance, 204 Metathoracic_spiracle_with_fringe_composed_of_dense_hairs / absent present, 205 Metathoracic_spiracle_with_opercular_lappet / posterior_fringe_unmodified 'posterior_fringe_operculum-shaped', 206 Postspiracular_area / unmodified a_membranous_fold a_distinct_scale, 207 Metasternal_area / bare setose, 208 Meron_of_metacoxa / present absent, 209 Postmetacoxal_bridge / absent metapleural_epimera_extended_behind_metacoxal_cavities, 210 Metasternal_furca / furca_with_anteriorly_directed_projection_and_occasionally_with_dorsoposterior_tip_also_developed furca_rod_shaped_lacking_apical_projections, 211 Wings_on_mesothorax / present absent, 212 Wings_on_metathorax / present absent, 213 Wings_of_female / attached_throughout_life deciduous, 214 Upper_calypter / not_developed developed, 215 Upper_calypter / moderately_developed_posteriorly_narrow_inner_edge_diverging_from_body_axis strongly_developed_broad_posteriorly_median_edge_converging_with_body_axis, 216 'Hair-like setae well developed in the fold between upper and lower calypter or in that region of the axillary wing membrane' / absent present, 217 Lower_calypter / not_developed developed, 218 Lower_calypter / moderately_developed_posteriorly_narrow_inner_edge_diverging_from_body_axis strongly_developed_broad_posteriorly_median_edge_converging_with_body_axis, 219 Stem_vein_ie_basal_section_of_R_upper_surface / bare with_row_of_setae, 220 Costa / circumambient_continuous_around_the_wing_but_weakened_along_the_posterior_margin abbreviated_ending_at_or_before_M2, 221 Basal_costal_bristles / absent present, 222 Basal_costal_bristles / one more_than_one, 223 Costal_setae / about_same_size alternating_thick_and_thin, 224 Humeral_crossvein / present absent, 225 Humeral_break / absent present, 226 Costal_break / absent present, 227 Wing_vein_Sc_course / 'straight_at_level_of_humeral_break/_crossvein_or_proximal_quarter_of_length' 'with_a_convexity_at_level_of_humeral_break/_crossvein_or_proximal_quarter_of_length', 228 Subcosta_termination / complete_reaching_costa 'subcosta_incomplete,_ending_just_short_of_costa' reduced_never_longer_than_half_length_of_basal_cells_or_ending_opposite_branching_of_Rs fused_apically_to_R1, 229 Subcosta / gradually_curved_or_faded with_90_degree_bend, 230 Crossvein_sc_to_r / present absent, 231 Crossvein_sc_to_r / near_humeral_crossvein not_near_humeral_crossvein, 232 Wing_vein_R1_configuration_opposite_end_of_subcosta / without_a_knob with_a_knob, 233 Rs / present absent, 234 Rs_origin / opposite_or_very_near_humeral_crossvein distal_distant_from_humeral_crossvein, 235 Rs / not_branched two_branched three_branched four_branched, 236 Radial_vein_distribution / evenly_distributed crowded_towards_costal_margin_R5_ending_before_wing_apex, 237 Wing_vein_R2plus3 / ending_near_wing_apex ending_near_R1, 238 Node_at_base_of_vein_R4plus5 / bare setose_dorsally, 239 R_4_plus_5 / branched unbranched_or_fused, 240 Fork_of_wing_vein_R4plus5 / not_strongly_divergent_usually_ending_either_above_or_below_wing_tip strongly_divergent_encompassing_wing_tip_such_that_R5_is_much_closer_to_M1_than_R4, 241 Crossvein_r_to_r_between_the_first_and_second_radial_veins_usually_named_R2 / absent present, 242 'R4+5 or R5 and M1' / simple strongly_curved_at_tip_of_wing_ending_anterior_to_wing_apex, 243 'R4+5 or R5 and M' / not_convergent convergent joined, 244 M / 'not_upcurved_at_fork_of_discal_cell_or_at_junction_with_crossvein_r-m' UPCURVED_at_fork_of_discal_cell_or_at_junction_with_crossvein_r_to_m, 245 Vein_M / a_straight_continuation_i.e._distal_part_not_bent_and_swung_forward distally_with_an_abrupt_bend_and_distal_part_swung_forward, 246 M2 / present absent, 247 Wing_vein_M3 / present absent, 248 Discal_or_discal_medial_cell / present absent, 249 Discal_cell / large small_short_and_broad, 250 Basal_or_basal_medial_cell / present absent, 251 Crossvein_bm_to_cu / present 'incomplete_or_absent_(cells_bm_and_dm_partially_united_or_fused)', 252 Relative_lengths_of_basal_and_discal_cells / basal_cells_greater_than_half_length_of_cells_d_or_dm basal_cells_shortened_less_than_half_length_of_d_or_dm_cells, 253 Spurious_vein / absent present, 254 Wing_venation / diagonal_vein_from_br_to_posterior_margin_of_wing_absent diagonal_vein_present, 255 CuA1 / without_kink 'with_a_baso-medial_kink_', 256 Vein_CuA2 / present absent, 257 Shape_of_distal_side_of_cup / tapering truncate rounded distinctly_convex with_angular_extension, 258 Veins_CuA2_and_A1 / apices_far_apart apices_close_together apices_fused_length_of_cell_cup_longer_than_apical_portion_of_A1 apices_fused_length_of_cell_cup_equal_to_or_shorter_than_apical_portion_of_A1, 259 First_anal_vein_ie_A1_plus_CuA2 / present absent, 260 First_anal_vein_ie_A1_plus_CuA2 / reaching_wing_margin_even_as_a_fold not_reaching_wing_margin, 261 'Vein A2 sensu Stary (2008)' / present absent_or_extremely_shortened, 262 'Alular fold sensu Stary (2008) (=A2 MND)' / not_converging_with_first_anal_vein converging_with_first_anal_vein, 263 Vein_A3 / present absent, 264 Veins_of_wing / 'Posterior_veins_well_developed,_as_distinct_as_the_veins_of_radial,_subcostal_and_costal_sectors' 'costalized,_posterior_veins_indistinct', 265 Halteres_on_metathorax / absent present, 266 Halteres_on_mesothorax / absent present, 267 Mid_coxal_prong / absent present, 268 Hind_coxa / simple well_developed_rounded_projection_on_anterior_face, 269 Hind_surface_of_hind_coxa / bare setose, 270 Femora / uniformly_sclerotized_undivided subdivided_by_apparently_membranous_cuticle_into_short_basal_zone_and_elongate_distal_zone, 271 Hind_femur / no_ventral_rows_of_stout_setae one_ventral_row_of_stout_setae two_ventral_rows_of_stout_setae, 272 Fore_tibial_spurs / present absent, 273 Mid_tibial_spur / present absent, 274 Hind_tibial_spurs / present absent, 275 Preapical_dorsal_tibial_bristle / absent present, 276 Male_fore_tarsus_or_fore_and_hind_tarsus / without_process with_thumb_nail_like_process, 277 Hind_tarsus / tarsomere_1_longer_than_tarsomere_2 tarsomere_1_shorter_than_tarsomere_2, 278 Last_tarsomere_of_male / not_capable_of_being_folded_proximally capable_of_being_folded_forward_against_penultimate_tarsomere, 279 Arolium / present absent, 280 Mediolobus / absent present, 281 Setiform_empodium / absent present, 282 Abdominal_spiracles / some_present all_absent, 283 Abdominal_spiracles_in_segments_1_to_5 / all_or_some_in_membrane all_in_tergite_margin, 284 Female_spiracles / present_on_segments_1to7 absent_on_segments_6_and_7_5_pairs absent_on_segments_5to7_only_4_pairs_present, 285 T1_to_2 / separate fused_into_syntergum, 286 First_abdominal_tergite / without_adventitious_suture with_adventitious_suture, 287 Abdominal_tergite_1 / simple divided_longtitudinally_in_midline, 288 Abdominal_tergite_2 / simple with_an_area_of_modified_setae, 289 Sternite_2_degree_of_exposition / freely_exposed_but_only_separated_from_margins_of_T1_to_2_by_a_narrow_strip_of_unfolded_membrane contiguous_with_or_slightly_overlapped_by_margins_of_T1_to_2 completely_overlapped_by_margins_of_T1_to_2 overlapping_margins_of_T1_to_2 freely_exposed_and_widely_separated_from_tergal_margins_by_more_or_less_folded_membrane, 290 Abdominal_plaques / present absent, 291 Abdominal_spiracle_8_in_males / present absent, 292 Rectal_papillae / three four five six 12_to_80, 293 Female_abdominal_segments_5_to_8_directed / posteriorly anteroventrally, 294 Apical_female_abdominal_segments / all_segments_exposed_gradually_telescopic segment_8to10_retracted segments_7to10_retracted segments_6to10_retracted segments_5to10_retracted, 295 Female_T6_and_ST6 / fused free, 296 Female_T7_ST7 / free fused, 297 Female_segment_7 / complete divided_into_broad_thickened_anterior_and_narrowed_posterior_regions, 298 Ovipositor_shape / unmodified lance_like sickle_shaped_, 299 Alpha_setae_sensilla_trichodea_on_ovipositor_sternites / present absent, 300 T8 / not_divided_longitudinally divided_longitudinally_, 301 ST8 / undivided 'plate_like_,_partly_divided_posteriorly,_divided_forming_hypogynial_valve' completely_divided_longitudinally_, 302 Sclerites_of_segment_8_of_female / separated articulated_anteriorly_or_fused, 303 Female_abdominal_tergite_9 / present absent, 304 Sternite_9 / present absent, 305 Sternite_9 / external partially_or_fully_invaginated, 306 Female_T10 / present_as_separate_sclerite not_a_separate_sclerite, 307 T10 / simple completely_divided_medially, 308 T10 / spines_absent 'bearing_large,_stout_spines_(acanthophorite_spines)', 309 Sternite_10 / present absent, 310 Segment_10 / tergite_and_sternite_10_separate T10_articulated_anteriorly_with_S10, 311 hypoproct / without_lingulae with_lingulae_lobe_like_lateral_extensions, 312 Female_Cerci / absent present, 313 Female_Cerci / two_segmented one_segmented_free_from_each_other, 314 Female_cercus / first_segment_not_expanded_ventrolaterally first_segment_expanded_ventrolaterally, 315 Cerci / not_distinctly_flattened distinctly_flattened_dorsal_and_ventral_faces_flat_in_cross_section, 316 Female_cerci / simple separated_by_abdominal_tergite_10, 317 Cercus / lightly_sclerotized heavily_sclerotized, 318 Spermathecae / absent one two three four, 319 Spermathecal_ducts / of_moderate_length_at_most_less_than_the_length_of_the_abdomen very_long_at_least_much_longer_than_the_abdomen, 320 Branching_of_spermathecal_ducts / 1_1_1_with_or_without_a_common_duct '2_plus_one,_2__with_common_duct', 321 Sperm_pump / absent present_well_developed_musculature_and_sclerotised_collars, 322 Female_accessory_glands / pair one absent, 323 Ventral_receptacle / absent present, 324 Male_tergite_6 / 'at_least_half_length_tergite_5_and_grading_into_the_terminalia,_usually_exposed' 'at_most_half_length_tergite_5_and_part_of_a_well-defined_unit_"terminalia",_rarely_exposed', 325 Syntergosternite_comprised_of_at_least_fusion_of_sternite_and_tergite_of_segment_8 / absent present_, 326 Syntergosternite / comprised_of_fusion_of_sternite_and_tergite_of_segment_8 comprised_of_fusion_of_sternites_and_tergites_of_segment_7_and_8_, 327 Vasa_deferentia_configuration / extended_caudally_not_arching_back_anteriorly U_shaped_apically_approximated, 328 Accessory_gland / separate_or_not_continuous_with_vasa_deferentia continuous_with_vasa_deferentia, 329 Epididymis / absent present, 330 Male_terminalia_resting_position / unflexed_projecting_posteriorly deflexed_ventrally dorsoflexed, 331 Hypopygium / unrotated_or_not_permanently_rotated rotated_permanently_between_45and90 'rotated_permanently_between_90and180;' rotated_permanently_almost_360_degrees circumversion_rotated_permanently_through_360 'rotated_180?,_with_rotation_occurring_between_segments_7_and_8', 332 Position_of_hypopygium / symmetrical_in_medial_line_of_abdomen asymmetrical_on_right_side_of_abdomen, 333 Epandrium_articulation / fused_to_or_lying_flat_on_hypandrium articulated_on_gonocoxites_or_hypandrium, 334 Epandrium / shallowly_notched_or_posterior_margin_straight posterior_margin_deeply_emarginate_forming_U_shape_with_basal_connection completely_divided_separated_along_midline, 335 Epandrium_and_hypandrium / fused_ring separate, 336 Hypandrium_and_gonocoxites / separate 'completely_fused,_no_seams_', 337 Gonocoxal_apodemes / confined_to_anterior_margin_of_hypandrium_not_extended_as_narrow_process project_little_beyond_margin_of_hypandrium elongate_extending_well_beyond_anterior_margin_of_hypandrium, 338 Postgonites / absent present, 339 Postgonal_apodeme / absent present, 340 Gonostyli / present absent, 341 Gonostyli / apical subapical, 342 Gonostyli_articulation / transverse_or_oblique dorsoventral, 343 Aedeagal_complex / surrounded_by_parameres_ 'fused_to_parameres_at_apex,_forming_phallus', 344 Parameres / paired_and_separate fused, 345 Aedeagus_or_phallus / at_least_partly_sclerotised entirely_membranous, 346 Aedeagus_or_phallus / single_tube_terminal_opening_apically two_tubes_at_the_apex with_three_slender_filaments_each_with_separate_opening, 347 Phallus / undivided subdivided_into_a_dorsal_guide_and_a_ventral_needle_like_component, 348 Phallus_dorsal_wall_of_distiphallic_tube / not_modified sclerotized_and_forming_the_dorsal_plate, 349 Phallus_dorsal_plate / divided_middorsally not_divided_middorsally, 350 Phallus_apical_part_of_dorsolateral_processes / tips_confluent_with_part_of_the_wall_of_the_phallic_tube free_tips, 351 Phallus_armature_small_sclerotized_teeth_on_distalmost_ventral_and_ventrolateral_surfaces / absent present, 352 Phallus / no_division divided_into_basi_phallus_and_distiphallus, 353 Phallus / without_epiphallus with_epiphallus, 354 Mesohypophallic_sclerotisation_mid_ventrally_along_distiphallus / absent_or_not_developed present, 355 Phallapodeme / absent present, 356 Phallapodeme / present_as_ingrowth_of_the_hypandrium lever_like, 357 Ejaculatory_apodeme / present absent, 358 Ejaculatory_apodeme / simple_without_posterior_projection with_a_posterior_extension_into_the_sperm_sac_endoaedeagal_guide inapplicable, 359 Ejaculatory_apodeme / fused_medially_to_basiphallus articulating_with_phallus loosely_attached_to_basiphallus, 360 Lateral_ejaculatory_processes / present absent, 361 Lateral_ejaculatory_processes / large_external small_retracted_within_base_of_phallus small_not_retracted_within_base_of_phallus_closely_associated_with_base_of_aedeagal_tines, 362 Ventral_plate / absent present, 363 Aedeagal_tines / absent present, 364 Male_tergite_10 / present absent, 365 Male_sternite_10 / present absent, 366 Surstyli_composed_of_epandrium_dorsally_and_bacilliform_sclerites_ventrally / absent present, 367 Surstylar_position / apical subapical_on_inner_margin_of_epandrium, 368 Subepandrial_sclerite / absent_membranous_at_least_in_part_of_length sclerotised_along_its_length_from_base_of_phallus_to_hypoproct_forming_subepandrial_sclerite sclerotised_divided_into_bacilliform_sclerites_processus_longi_laterally, 369 Male_cerci / present absent, 370 Cerci / not_modified_as_clasping_structure modified_as_clasping_structure, 371 Cerci / separate_i.e._not_fused fused, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 370 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Acrosathe_novella 101011102000??11?015001???100????1?10120000?001?1??00001000?1??101002100000?01211?11000020??00??0100?0005?1100100000?000?0?0000000000011000000200??0000112111101101010000110000111101?0011011001111100000100?001000101000?000?000000000001200001000100000000000002001010100110111100001010000000401?001000001000101100011000120?00000?000200101010?0111?0000??00000?0000100110?0000 Anopheles_gambiae 100????00000??1110010101??02101200110000100?0000010000001011011111001000001100110?00101000??00??0100?0111?1003?0?000?000?0?00010101?0001001001010??0001110111100?0000?000010010000001?000??0?000?11000000100?001000101000?000?00000001?00120001?00000011?00?0000?0001?101000000000000010100000004011001000001000000000011000010?0??00?00005000000??0000010????????0?1??1?10000?0000 Arachnocampa_flava 100????01000??10?0040000??0010???0000000000?000001101001000?000101001000000?00210?100010?0??00??0100?0001?1000000000?000?0?00010101?0000?00000000??0000110111100?0100?000100010110001?000??0?001?11000000100?0010?0101000?010?0000000010010??01?00010011?00?0000?0001010100000000000001100000000401?001000001000100000010000020?0?000?110000001010?0000100????????0?00?1?00000?0000 Asilus_crabroniformis 101011002010??110016001???100????1?10120000?001?1??00001000?1??101002000000?01210?11000020??00??0100?0001?0000110000?000?0?0100000010011000000200??0002112111110?01010000100000111101?0011011001011000000100?001100101100?000?000000000001201000000100000000000002001?101000002111000010100000004012001000001000100000011000130000000?000000111000?0111?0200??00000?0100000100?0000 Bibio_longipes 100????01000??00?00?0000??000????0000000000?000001000001000?000101001000000?01200??0001000??00??0100?0001?1100100000?000?0?00000001?0010?00000000??0000110111100?0100?000110010110001?000??0?001?11000000100?001010101000?010?0000010010010??01?00010011?00?0000?0011010100000000000001100000000401?00100000000011??0001100003000??00?110000000110?0000100????????0?00?1?00000?0000 Bittacus_mastrillii 100????0??00??02?0000000??000????1?00000000?000001000001000?1??100000000?00?01000????00010??00??0000?0002?1000000000?000?0?00000001?0000?00000000??00000001110?0?0010?000001??0??0001?000??0?001010?00000100?000000000??0?000?0000000000013000001000000000000000?0000?000000000000000100000000004003001000000000100010?11000010?00000?1110000100?0?0000000????????0????1?00000?0000 Bombylius_major 101010002010??11?016001???100????1?10120000?000?1??00001000?1??101002000000?00210?11000020??10??0100?0001?1100100000?000?0?0000001011110?00000200??0000112111101001010000100000010101?0011011000?11100000100?001000101100?000?00000001?001200001002000100000000011001?10100000111100001010000010401?011000?02000101100011000030010000?000120101010?0111?0000??00000?0000000100?0000 Bradysia_tilicola 100????01000??00?00?0000??0010???0000000000?000001101001000?000101001000000?01200?100010?0??00??0100?0001?1000000000?000?0?00010001?0000?00000000??0000110111100?0100?000110010110101?000??0?001?11000000100?001000101000?000?00000101?0010??01?00010011?11?0000?0011?11100000000000001100000000401?0010000010001000000100000???0??00?110000000110?0000100????????0?00?1?00100?0000 Ceratitis_capitata 0??????????2111??016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211020101120??11112100?0007?100010111101010120000101001011010110301000000112111101101011101100000011110?0011110001111000100000?0010001010010010?00111011?10010011?000001100001000043011?10101000010000001010001000411?011111102011?1??1?01100012??0011110?014011110101??1?0000??0100110021?001?102001 Chamaepsila_hennigi 0??????????2111??016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211020101120??11212100?000??10001011110101?0?0000100001010?00120301000000112111101101011101100000111101?0013110001011000000000?001010101000?010?00010101?00010001?000001100001000012011?10101000011100001010001000411?021100100011?1??1?011000020?0??01100014011110101??1?0000??0100110021?001?102001 Chironomus_tepperi 100????00000??13100?0000??00101110110000?00?0000010000010010011101101000010?0022??????00?0??00??01010010??1003?0?000?000?0?00010001?0000?00001010??0000110111100?0000?000010010000001?000??0?000?11000000100?001000101000?010?00100101?00111001?00000111?00?0000?01?1?11100000000000001010000000401?001000000000101010011000020?01000?110000000010?0000010????????0?1??1?00100?0000 Clogmia_albipunctata 100????00000??00?00?0000??10110000110110100?100001000001?00?011101002100000?00110??0101000??00??0100?000??1003?0?000?000?0?00010001?0000?00000000??0000110111100?0000?000010010000001?000??0?000?11000000100?001000101000?000?01000200?00030000000000001?00?0000?0001?10100000000000001100000000401?00100000000011??1?0110001??????00?010020000000?0010100????????0?00?1?00100?0000 Coboldia_fuscipes 100????01000??11000?0000??10110000110100100?100001001000100?000101001000000?01200??0101000??00??0100?000??1000000000?000?0?00000?11?0000?00000100??0000112111100?0100?000100010000101?000??0?000?11000000100?001000100??0?010?00000201?0010??01?00010011?11?0000?0011?11100000000000001100000000401?011?0000100011??0?011000010?0?000?000000000110?1??0100????????0?00?1?00100?0000 Cochliomyia_macellaria 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000300?1??001002000000?01211020101120??11102100?0007?1100101111011111200101010010111101103010100101121111011110111011001011111103101101100111101010100110110001010111111110111001?10010011?0010111000010000130110101010100111000010101010003111031000000011?1??1?111000030100011100014011110111??1?0001011111110021?0011102010 Ctenocephalides_felis 100????00000??0110060000??000????1?00000000?00001?000001000?000101000000?00?01200????000?0??00??0100???00?1??1000000?000?0?000?00?1?0000?00000000??0000110?100?0?0000?000001??0??0?01?0?0??0??????????0??10??0010111????????????????????????????????????????????????????000000011100001000000000010300100000000101??00011000010?02000?1010000000?0?0110?00????????0????1?00000?0000 Delia_radicum 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000?00?1??001002000000?01211020101120??11112100?000??1100101111011111210101010010111101103010000101121111011110111001001011111103001101000110100010000100010001010110011110110001?10010001?0000011000010000130010101010000111000010101010000111031000002011?1??1?111000030?00011100014011110101??1?0000??0110110021?0011102011 Drosophila_melanogaster 0??????????2111?0016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211120101120??11112100?0006010001011112101{01}0?0010101001011010120301010000112111101101011100100000111100?0011110001111000000?00?0010?0101000?01100011?101?00010001?00000110011?000012011?101000000101100010100010004111011000002011?1??1?00?????20?00101100014011110101??1?0000??0100110021?001?102000 Edwardsina_gigantea 100????10100??1110050000??000????0110100000?000001001000100?1?0101004010000?0012??????00?0??00??11111100??1000000000?000?0?00010101?00?0?00000000??0000110111100?0000?000000110000001?000??0?000?11000010100?0010?0101000?000?00000101?00120000000010011?00?0000?0001?10100000000000001??0000000401?001000001000100000011000030000000?000200001110?0000102????????0?00?1?00100?0000 Episyrphus_balteatus 0??????????2001??016101?10??0????1?11?0?000?000?1??11000000?1??001002000000?01211?20101020??11402100?0005?0100100000?000?0?00001001?1010?00020301000000112111101101010100100000000101?0010?10000?01000000100?0110001010010010?00000001?00110001?0010001000001000020010101000000111000010000011004111031000000011?00000011000030?0?011000014111110101??1?0000??0100110021?0010102000 Exeretonevra_angustifrons 111100000000??110006001???100????1?10120000?001?1??00001000?1??101002000000?11210?10100020??00??0100?0001?1000100000?000?0?0001010000011000000100??0000111111101101010000100000000101?0010?11000?11101000100?011000101000?000?000000000001200000010100000000010011001010100000000000001100000000401?001000001000101000010000130000000???0200001010?0000100????????0?0100000100?0000 Exorista_larvarum 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000?00?1??101002000000?01211020101020??11102100?0007?1000101111011111200101010010111101103010000101121111011010111001001011111102101101000111100210100110010001010111011110111001?10010011?0010111000010000130110101010000111000010101010002111031000000011?1??1?011000030?00011100014011110111??1?00011?1111110021?0011102011 Glossina_morsitans 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000?00?1??10?002000000?0121???01010211111112100?000??1000101100?11111200101010010110101103010100101121111011110111011001011111104001101000111100010010100110001010110011100111001?00010001?001001100001000013011010101000011100001010001000411101100010?011?1??1?011000020?00011100014011110101??1?0000??1100111??1?0011102000 Haematopota_pluvialis 100????02010??131006001???100????1?101210012010?1??00011000?1??101002000000?01210?00101020??00??0100?000{3456}01103?0?000?000?0?00010001?0001000000100??0000112111101101010000100000000101?0010?11000?11001000100?2111001010010000?000000000001200001001100000000000002001?101010000111000011000000104013001000001000101000011010030010000?110000121120?0000100????????0?0100201100?1000 Hermetia_illucens 100????02000??121005001???100????1?10120000?000?1??00100210?1??101012000000?01210?10110020??01??0100?000{456}01000100000?000?0?0001000010010?00000100??0000112111101101010000110000000101?0010?11000?11101000100?011000101000?010?000000000001211100000100001000000022001010100000011100001100000000401?021000000000100000010001130010000?000000001120?0101?0100??10000?0001?00000?0000 Lonchoptera_lutea 0??????????1001??016101??0??0????1?11?3?000?000?1??11000300?1??101002000000?01211?20101020??11502100?000??10001010112000?1300101001?101100002030110000011211110110100?100100001101101?0011010001011000000100?001000100?00?001000000001?00010011?000000110001000010001?101000000111100010101010004111001000000010100000011000020?0?001000004001110101??1?0000??0100110021?0010100000 Mayetiola_destructor 100????01000??00?00?0000??000????0000000000?000001101001000?000101001000000?01200??0001000??00??0100?000??1003?0?000?000?0?00000?01?0000?00000020??0000110111100?0100?000110010000001?000??0?000?11000000100?001000100??0?010?01000101?01?00?01?00000011?11?0000?01?1?11100000011100001100000000401?00100100000011??00011000020?0??00?110200001110?0000100????????0?00?1?00100?0000 Megaselia_scalaris 0??????????1001??016101?10??0????1?11?0?000?000?1??11000300?1??001002000000?01211?20101020??11402100?0007?0000100011100101110101011?101100002030110000011211110110100?100101??0111101?000??10001011100000100?001000100?00?011000000001?00011101?00000010011?000002001?111000000100000010101011004111021000000011?00000011000020?0?0011000040111100?1??1?0000??0100100021?0010102000 Micropteryx_calthella 100????0??00?????00000????000????1?00000000?000101000000000?000100000000?00?01000????00010??00??0000?0002??0010000000000?0?000100?010000?00000000??000?0000010?0?0000?000001??0??0001?000??0?000?10?00000100?000000000??0?000?000000001001300?1?00000001?00?0000?0000?10000000011000000000000000000000100000001001??1??0?????10?00000?0000000000?0?1??1?0000??00000?1??1?00000?01?? Minettia_flaveola 0??????????2111??016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211?20101?20??11112100?0006010001011112101?100000101001011000010301010000112111101101011100100001111110?0011110001111000000000?011000101000?011000001001?00010001?000001100001000013011?10101000000010001010001000011?031100102011?1??1?01100003010??1110?014011110101??1?0000??01?0110021?001?102000 Musca_domestica 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000300?1??101002000000?01211020101120??11102100?0007?1200101111011111200101010010110101103010100101121111011110111001001011111104001101000111100010000100010001010110011110111001?10010001?0010111000010000130110101010000111000010101110000111031000002011?1??1?111000030?00011101014011110101??1?0000??0110110021?0011102010 Nannochorista_philpotti 100????01?00???310060000??000????1?00000000?000000000001000?000100000000?00?00000??00000?0??00??0000?000??1000000000?000?0?00000101?0010?00000000??00001101110?0?0010?000001??0??0001?000??0?000?10?00000100?000000000??0?000?00000001?0012000001000000000000000?0000?000000000000000000000000004???001000000000000000010000010?0??0101110000000?0?0000000????????0?1??1?00000?0000 Nymphomyia_dolichopeza 100????00000??1110050000??100????0110000010?0000010000010010011101000010010?0022??????00?0??00??0100?0000?1003?0?000?000?0?000000?1?0010?00000000??1??????????????000?000000??0000001?000??0?000?11000000100?0010?0110??0?000?01000001?01?00????0??00111?11?0001??1?1?10100001011100001101000000401?001000000?0001??1?01100000??0??00???00000001?0?0100100????????0?1??1?00110?0000 Ogcodes_basalis 100????02010??12?016001?????0????1?10120000?000?1??00001000?1??101002000000?01200??1101020??10??0100?0002?1111100000?000?0?000??0?010010?00000300??1??????????????1010000111??0000101?0010?11000?11100000100?0010001010011010?01000000000100???????00111?11?0000?0011010100000011100001100100000401?02100000000011??00011000020?00000?000130101110?1??1?0000??00000?0000000100?0000 Oreogeton_obscurus 1010110020100011?016001???100????1?10130000?000?1??10001000?1??101002010010?01210?10101010??10??0100?0002?1100101110?000?0?00000001?0011000000300??0100112111101001011000100001111100?0011010001111010000120?001000101000?0010000000000001200000000000100000000023011?101000000111000010100000004011001000000010101000011000010?00000?00000011111101??1?0000??00000?0011?00100?2000 Physocephala_marginata 0??????????2001??016101?10??0????1?11?0?000?000?1??11000000?1??001002000000?01211220101020??11112100?000611003?00000?100?0?0000101001010?01010200??0000112111101101011100100000000101?0010?10000?11000000000?001100101000?010?00000000100010001?0010011000010000020010101000002111000010101011004111131000000011?1?01?011000030?0??01100014011110101??1?0000??0100110021?0011102000 Phytomyza_ilicicola 0??????????2021??016101?11??0????1?11?0?000?000?1??11000400?1??101002000000?01211020101?20??11112100?000??100010111101113120010101001011000010301000000112111101101011100100000111110?0011010001111000000000?001010101000?011100011101?00010001?00000110??0?000012011?11101000000000001010001000411?0101101?20?????????1000002??0011110?014011110101??1?0000??0100110021?001?102000 Protoplasa_fitchii 100????00000??12100?0000??01100000000000000?000001000000100?010101001000010?00110??100?000??00??0100?000??1003?0?000?000?0?000100?1?0000?00000000??0000110111100?0010?000000110000001?000??0?000?11000000100?001000101000?000?0000000010013000000000000000000000?0001?10100000000000011100000000401?0010000000001??????1000003000??00???0000001110?0000002????????0?00?1?00110?0000 Sarcophaga_bullata 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000300?1??001002000000?012110201111210011112100?0007?1200101111011111200101010010111101103010100101121111011110111001001011111102101101100111101010110110110001010111011110111001?10010011?0010111000010000130110101010100111000010101010003111031000100011?1??1?011000030100011100014011110111??1?00011?1101110021?0011102010 Scathophaga_stercoraria 0??????????2111?0016101?01??0????1?11?0?000?000?1??11000?00?1??101002000000?01211020101120??11112100?000??100010111101111120010101001011010110301010010112111101111011101100101111110500110100011110001000010001000100?10?011100111001?10010001?0000011000010000130010101010000111000010101010000111031000002011?1??1?011000030?00011101014011110101??1?0000??0110110021?0011102010 Sepsis_cynipsea 0??????????2111?0016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211120101120??11112100?000??10001011110101?0?0000100001010?00010301000000112111101001011101100000101111?0011110001011000000000?001000101000?011100010001?00010001?000001100001000013011?101010000100100010100010004111031000112011?1??1?011000020?0?111100014011110101??1?0000??0100110021?001?102000 Spelobia_bifrons 0??????????2111??016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211?20101?20??11112100?000??1000101111210130?1010101001011000010301000000112111101101011100100001111100?0011110001111000000000?001000100?00?01110011?001?00010101?00000110011?0001??011?11100000010000101010001000411?02?000100011?1????01100003010??1110?014011110101??1?0000??0100110021?001?10200? Sylvicola_fenestralis 100????02000??01000?0000??10110000110110100?10001?001000100?000101001100000?01210?10101000??00??0100?0001?1000100000?000?0?00000001?0001000000000??0000110111100?0100?000100010011001?000??10001011000000100?001000101000?010?00000000000110101?0000000000000000?0001010100000000000001100000000401?00100000100011??00011000010?0??00?010020020110?0010100????????0?00?1?00000?0000 Teleopsis_dalmanni 0??????????2111??016101?11??0????1?11?0?000?000?1??11000300?1??001002000000?01211020101120??11?12100?000??1000101010?111?0?0000101????11001010301000000112111101101110100110000000100?0110?10011011100100000?001100100?10?010??0000001?00010001?00000110011?000012011?10101000011100001010001000411?031100102011?0001?011000020?00101100014011110101??1?0000??0100110021?001?102000 Thaumatomyia_glabra 0??????????2111??016101?11??0????1?11?0?000?000?1??11000300?1???01002000000?01211020101?20??11112100?000611000101111011130?0010101001011000010301000000112111101101011110100000011101?0011010001111000000000?001000101000?01110001?201?00010001?00000110011?0011??1?1?10100000011100001010001000411?03??001???1????????11000020?00111100014011110101??1?0000??0100110021?001?10200? Tipula_abdominalis 100????10010??11?0010000??000????0110000000?00000000100??010011101002000000?00210?1010?000??00??0100?0001?1003?0?000?000?0?00000001?0000?00000000??000011011110110010?000000110000001?000??0?000?11000000100?001000101000?000?00000300100120001?1000000000000000?00000101000000000000000000000004011001001000000100000011000130000000?100000000110?000??00????????0?00?1?00110?0000 Trichocera_brevicornis 100????01000???1000?0000??10110000110110100?100001001000100?000101002100000?00210?1010?000??00??0100?0000?1000100000?000?0?00010001?0000?00000000??0000110111100?0010?000000110000001?000??0?000?11000000100?001000101000?000?0000000010012000001000000000000000?0000010100000000000000000000000401?00100100100010000001100013000?000?100000000010?0000100????????0?00?1?00100?0000 ; END; BEGIN SETS; CHARSET adult (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 105-371; CHARSET adultext (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 105-292; CHARSET extnonwing (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 105-210 265-292; CHARSET pupa (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 94-104; CHARSET male (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 324-371; CHARSET extwing (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 211-264; CHARSET larva (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 1-93; CHARSET adultint (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 293-371; CHARSET female (CHARACTERS = FLYTREE_Morphological_Matrix_with_Exemplar_Species_Names) = 293-323; END; [ The following blocks are output data for analysis step 10449 ] BEGIN TREES; TITLE Chosen_MPT_from_FLYTREE_morphology_matrix; LINK TAXA = TaxaForAnalysisStep10449; TREE FLYTREE = [&R] (((Micropteryx_calthella,(Bittacus_mastrillii,Nannochorista_philpotti)),Ctenocephalides_felis),(((Anopheles_gambiae,Chironomus_tepperi),((((((((((Arachnocampa_flava,Bibio_longipes),Bradysia_tilicola),Mayetiola_destructor),Coboldia_fuscipes),Sylvicola_fenestralis),Trichocera_brevicornis),Clogmia_albipunctata),(((((((Asilus_crabroniformis,Acrosathe_novella),Bombylius_major),(Oreogeton_obscurus,((Episyrphus_balteatus,(Physocephala_marginata,((Chamaepsila_hennigi,((((((Drosophila_melanogaster,Spelobia_bifrons),Thaumatomyia_glabra),Phytomyza_ilicicola),Minettia_flaveola),(Ceratitis_capitata,((((((Cochliomyia_macellaria,Sarcophaga_bullata),Exorista_larvarum),Musca_domestica),Delia_radicum),Scathophaga_stercoraria),Glossina_morsitans))),Sepsis_cynipsea)),Teleopsis_dalmanni))),(Megaselia_scalaris,Lonchoptera_lutea)))),Ogcodes_basalis),Haematopota_pluvialis),Hermetia_illucens),Exeretonevra_angustifrons)),(Edwardsina_gigantea,Protoplasa_fitchii)),Tipula_abdominalis)),Nymphomyia_dolichopeza)); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr59132] END;