#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 20, 2025; 23:46 GMT TreeBASE (cc) 1994-2008 Study reference: Davalos L.M., Velazco P.M., Warsi O., Smits P.D., & Simmons N. 2014. Integrating Incomplete Fossils by Isolating Conflicting Signal in Saturated and Non-Independent Morphological Characters. Systematic Biology, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S13970] [ The following blocks are input data for analysis step 10991 ] BEGIN TAXA; TITLE TaxaForAnalysisStep10991; DIMENSIONS NTAX=113; TAXLABELS Anoura_caudifer Anoura_cultrata Anoura_geoffroyi Anoura_geoffroyi_peruana Anoura_latidens Artibeus_gnomus Artibeus_jamaicensis Artibeus_lituratus Artibeus_obscurus Brachyphylla_cavernarum Brachyphylla_nana Carollia_brevicauda Carollia_perspicillata Choeroniscus_godmani Choeroniscus_minor Choeroniscus_periosus Choeronycteris_mexicana Chrotopterus_auritus Desmodus_rotundus Diaemus_youngi Diphylla_ecaudata Enchisthenes_hartii Erophylla_bombifrons Erophylla_sezekorni Glossophaga_commissarisi Glossophaga_leachii Glossophaga_longirostris Glossophaga_morenoi Glossophaga_soricina Glyphonycteris_daviesi Glyphonycteris_sylvestris Hylonycteris_underwoodi Lampronycteris_brachyotis Leptonycteris_curasoae Leptonycteris_nivalis Leptonycteris_yerbabuenae Lichonycteris_obscura Lionycteris_spurrelli Lonchophylla_handleyi Lonchophylla_mordax Lonchophylla_robusta Lonchophylla_thomasi Lonchorhina_aurita Lonchorhina_inusitata Lonchorhina_orinocensis Lophostoma_brasiliense Lophostoma_brasiliense_minuta Lophostoma_carrikeri Lophostoma_evotis Lophostoma_kalkoae Lophostoma_occidentalis Lophostoma_schulzi Lophostoma_silvicolum Lophostoma_silvicolum_centralis Lophostoma_silvicolum_silvicolum Macrophyllum_macrophyllum Macrotus_californicus Macrotus_waterhousii Micronycteris_brosseti Micronycteris_hirsuta Micronycteris_homezi Micronycteris_matses Micronycteris_megalotis Micronycteris_microtis Micronycteris_microtis_mexicana Micronycteris_minuta Micronycteris_schmidtorum Mimon_bennettii Mimon_cozumelae Mimon_crenulatum Mimon_crenulatum_keenani Mimon_crenulatum_longifolium Monophyllus_plethodon Monophyllus_redmani Mormoops_blainvillei Musonycteris_harrisoni Neonycteris_pusilla Noctilio_albiventris Noctilio_leporinus Notonycteris_magdalenensis Notonycteris_sucharadeus Palynephyllum_antimaster_2 Phylloderma_stenops Phylloderma_stenops_septentrionalis Phyllonycteris_aphylla Phyllonycteris_major Phyllonycteris_poeyi Phyllostomus_discolor Phyllostomus_discolor_verrucosus Phyllostomus_elongatus Phyllostomus_hastatus Phyllostomus_hastatus_panamensis Platalina_genovensium Platyrrhinus_helleri Pteronotus_davyi Pteronotus_parnellii Pygoderma_bilabiatum Rhinophylla_alethina Rhinophylla_fischerae Rhinophylla_pumilio Sturnira_lilium Thyroptera_tricolor Tonatia_saurophila Tonatia_saurophila_bakeri Tonatia_saurophila_saurophila Trachops_cirrhosus Trachops_cirrhosus_coffini Trinycteris_nicefori Uroderma_bilobatum Vampyrodes_caraccioli Vampyrodes_major Vampyrum_spectrum Xeronycteris_vieirai ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M22191] TITLE Chiroptera_Morphological_data; LINK TAXA = TaxaForAnalysisStep10991; DIMENSIONS NCHAR=278; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5" MISSING=? GAP= -; CHARSTATELABELS 1 Upper_incisors_direction_of_projection_of_tooth_shaft / orthodont proodont, 2 Upper_incisors_orientation_of_tip_of_crown / orthoclivous retroclivous, 3 Upper_I1_presence / I1_present absent, 4 Upper_I1_crown_differentiation / crown_clearly_differentiated_from_root_shaft_ not_differentiated_from_root_shaft, 5 Upper_I1_shape_of_occlusal_margin / large_main_cusp_that_tapers_to_a_blunt_point_additional_lateral_cusp_may_be_present_ straight_or_slighlty_convex_and_rounded bilobed blade_like_with_concave_C_shaped_cutting_edge, 6 Upper_I1_form_of_lobes / mesial_lobe_smaller_than_distal_lobe lobes_of_equal_size mesial_lobe_larger_than_distal_lobe, 7 Upper_I1_direction_of_lobe_projection / ventrally mesially, 8 Upper_I1_position_of_pointed_main_cusp / main_cusp_offset_mesially_from_axis_of_tooth main_cusp_in_center_of_tooth_so_tooth_crown_is_roughly_bilaterally_symmetrical_ main_cusp_offset_distally_from_axis_of_tooth, 9 Upper_I1_distal_accessory_cusp / absent present, 10 Upper_I1_relative_size_of_pointed_mesial_cusp / mesial_cusp_2x_the_height_of_distal_cusp pointed_mesial_cusp_only_slightly_higher_than_distal_cusp, 11 Upper_I1_lingual_cingulum_presence / absent present, 12 Upper_I1_occlusion / I1_in_contact_with_i1_or_separated_by_a_small_gap_when_cheek_teeth_are_in_occlusion I1_separated_from_i1_by_a_large_gap_when_cheek_teeth_are_in_occlusion I1_occludes_with_mesial_side_of_i2 I1_occludes_with_anterior_cingulum_of_lower_canine. I1_occludes_with_cingular_shelf_on_lower_canine_well_posterior_to_i1 I1_occludes_posterior_to_i1_in_mandibular_fossa, 13 Upper_incisors_relative_crown_height / height_of_I1_less_than_height_of_I2 I1_and_I2_subequal_in_height height_of_I1_greater_than_height_of_I2_but_less_than_twice_the_height_of_I2 height_of_I1_at_least_twice_that_of_I2, 14 Upper_I2_occlusal_exposure / I2_in_normal_position_between_I1_and_C_not_excluded_from_occlusion_toothrow I2_moved_dorsally_and_excluded_from_occlusal_toothrow_by_close_apposition_of_I1_and_C I2_moved_ventrally_and_excluded_from_occlusal_toothrow_by_close_apposition_of_I1_and_C, 15 Upper_I2_presence / I2_present absent, 16 Upper_I1_and_I2_relative_position / I2_located_more_posterior_than_I1 I2_lateral_to_I1_not_displaced_posteriorly_or_anteriorly_incisors_form_transverse_line I2_displaced_anteriorly, 17 Upper_I2_crown_morphology / well_developed_crown_morphology simplified, 18 Upper_I2_lingual_cingulum_presence / absent cingulum_present, 19 Upper_I2_and_C_diastema_presence / diastema_present diastema_absent_between_I2_and_C_teeth_in_contact, 20 Upper_C_crown_height / upper_C_greater_than_or_equal_to_twice_the_height_of_I1 upper_C_much_less_than_twice_the_height_of_I1, 21 Upper_C_crown_proportions / not anteroposteriorly_elongated, 22 Upper_C_labially_swollen / not upper_C_labially_swollen, 23 Upper_C_anteromedial_accessory_cusp / no_raised_cingulum_or_cusp raised_cingulum_present_but_cusp_absent cusp_present_but_low cusp_tall_greater_than_half_height_of_main_cusp, 24 Upper_C_anteromedial_groove / groove_absent anteromedial_groove_present, 25 Upper_C_lingual_cingulum_ / lingual_cingulum_absent faint_lingual_cingulum_present narrow_well_developed_cingulum_present wide_well_developed_cingulum_present, 26 Upper_lingual_cingulum_indentation / absent weakly_developed strongly_developed, 27 Upper_C_lingual_cingulum_cusp / absent present, 28 Upper_C_lingual_cingulum_slope / not_steep steep, 29 Upper_C_labial_cingulum_development / cingulum_well_developed_and_complete cingulum_weakly_developed_but_complete cingulum_weakly_developed_and_incomplete cingulum_absent, 30 Upper_C_labial_cingulum_incompleteness / incomplete_anteriorly incomplete_in_the_middle incomplete_posteriorly, 31 Upper_C_distal_cingulum / absent cingulum_small large, 32 Upper_C_posteromedial_ridge / ridge_absent posteromedial_ridge_present two_posteromedial_ridges_present_forming_a_groove, 33 Upper_C_posterolingual_surface / rounded posterolingual_surface_flattened, 34 Upper_C_posterolateral_accessory_cusp / cusp_absent trace_of_cusp_present cusp_tall_and_conspicuous_but_less_than_half_the_height_of_the_main_cusp cusp_very_tall_equal_to_or_greater_than_half_the_height_of_the_main_cusp, 35 Upper_C_posterolateral_accessory_cusp_form / conical_and_well_separated_from_main_cusp bladelike, 36 Upper_C_posterolateral_accessory_cusp_position / cusp_located_on_posterior_cingulum_well_seperated_from_main_canine_cusp cusp_originates_on_posterior_face_of_main_canine_cusp_not_well_seperated_from_that_cusp, 37 Upper_C_posterolabial_surface / rounded posterolabial_surface_flattened, 38 Upper_C_curvature / less_recurved more_recurved, 39 Upper_C_angle_of_projection / does_not_project_anteriorly_long_axis_of_tooth_shaft_is_perpendicular_to_plane_of_occlusal_surface_of_molar_toothrow projects_anteriorly_long_axis_of_tooth_shaft_forms_an_oblique_angle_with_the_plane_of_the_occlusal_surface_of_the_molar_toothrow, 40 Upper_P1_development / P1_with_well_developed_crown_that_is_greater_in_diameter_than_the_root_crown_conical_triangular_or_lenticular_in_labial_view P1_peglike_crown_simplified_and_flattened_crown_diamenter_subequal_to_or_only_slightly_larger_than_root_diameter P1_present_as_a_tiny_spicule_with_crown_diameter_less_then_or_equal_to_root_diameter P1_absent, 41 Upper_P1_number_of_roots / 1_root 2_roots, 42 Upper_P1_labial_cingulum_presence / absent labial_cingulum_present, 43 Upper_P1_position / P1_labially_extruded_from_toothrow_P5_directly_contacts_tooth_anterior_to_it_usually_C_while_P1_is_located_labial_to_this_contact P1_partially_extruded_from_toothrow_on_labial_side_but_still_partially_separates_teeth_anterior_and_posterior_to_it. P1_lies_in_line_with_C_P5_and_M1_2 P1_partially_extruded_from_toothrow_on_lingueal_side_but_still_partially_separates_teeth_anterior_and_posterior_to_it P1_lingually_extruded_from_toothrow_P5_directly_contacts_tooth_anterior_to_it_usually_C_while_P1_is_located_lingual_to_this_contact, 44 Upper_P1_and_P4_relative_crown_length / P1_shorter_than_P4 P1_and_P4_subequal P1_longer_than_P4, 45 Upper_P4_presence / P4_present absent, 46 Upper_P4_number_of_roots / three_roots two_roots one_root, 47 Upper_P4_position / P4_lies_in_line_with_C_P5_and_M1_2 P4_partially_extruded_from_toothrow_but_still_partially_separates_teeth_anterior_and_posterior_to_it P4_entirely_extruded_from_toothrow_P5_directly_contacts_tooth_anterior_to_P4_usually_C_, 48 Upper_P4_labial_cingulum_development / labial_cingulum_well_developed weakly_developed absent, 49 Upper_P4_lingual_cingulum / cingulum_absent weakly_developed well_developed, 50 Upper_P4_lingual_cusp / absent_no_cusp_or_cingulum raised_cingulum_present_but_no_cusp cusp_present, 51 Upper_P4_postparacrista / absent single_crest pair_of_roughly_parallel_crests_with_a_groove_in_between, 52 Upper_P4_postparacrista_extent / postparacrista_decreases_sharply_in_height_near_paracone_and_does_not_extend_to_distal_edge_of_tooth postparacrista_extends_as_a_continuous_crest_to_the_distal_aspect_of_the_tooth, 53 Upper_P4_distal_accessory_cusps / distal_accessory_cusps_absent one_distal_accessory_cusp_present two_distal_accessory_cusps, 54 Upper_P4_and_P5_diastema_presence / no_diastema_present_between_P4_and_P5_teeth_in_contact_or_nearly_so diastema_present, 55 Upper_P4_and_P5_relative_crown_length / P4_shorter_than_P5 P4_and_P5_subequal P4_longer_than_P5, 56 Upper_P4_and_P5_relative_crown_height / P4_shorter_than_P5 P4_and_P5_subequal_in_height P4_taller_than_P5, 57 Upper_P5_lingual_cingulum_development / cingulum_absent lingual_cingulum_narrow_no_lobe_present lingual_cingulum_large_forms_distinct_lobe_does_not_extend_as_far_lingually_as_protocone_of_M1. lingual_cingulum_very_large_forms_distinct_lobe_extends_as_far_lingually_or_farther_than_protocone_of_M1., 58 Upper_P5_anterior_margin_ / narrow wide, 59 Upper_P5_lingual_cingular_cusp / cingulum_with_raised_edge_but_no_cingular_cusp lingual_cusp_weakly_developed lingual_cusp_well_developed two_well_developed_lingual_cusps, 60 Upper_P5_labial_cingulum_development / labial_cingulum_well_developed weakly_developed absent, 61 Upper_P5_labial_cingular_cusp / cusp_absent weakly_developed strongly_developed, 62 Upper_P5_anterolabial_cusp / absent weakly_developed strongly_developed, 63 Upper_P5_postparacrista / absent single_crest pair_of_roughly_parallel_crests_with_a_groove_in_between, 64 Upper_P5_postparacrista_extent / postparacrista_decreases_sharply_in_height_near_paracone_and_does_not_extend_to_distal_edge_of_tooth postparacrista_extends_as_a_continuous_crest_to_the_distal_aspect_of_the_tooth, 65 Upper_P5_distal_accessory_cusps / distal_accessory_cusps_absent one_distal_accessory_cusp_present two_distal_accessory_cusps, 66 Upper_P5_proportions_in_occlusal_view / P5_length_less_than_width length_and_width_subequal P5_length_greater_than_width, 67 Upper_P5_and_M1_diastema_presence / no_diastema_between_P5_and_M1_teeth_in_contact diastema_present, 68 Upper_P5_and_M1_M2_alignment / not_aligned_longitudinal_axis_through_M1_and_M2_paracone_runs_markedly_labial_to_P5_paracone_ aligned_longitudinal_axis_through_M1_and_M2_paracones_intersects_P5_paracone_ not_aligned_longitudinal_axis_through_M1_and_M2_paracone_runs_markedly_lingual_to_P5_paracone_, 69 Upper_P5_and_M1_relative_length / P5_length_conspicuously_greather_than_M1_length P5_and_M1_subequal_in_length P5_length_conspicuously_less_than_M1_length, 70 Upper_molars_ectocingulum_development / strongly_developed present_but_weak absent, 71 Upper_molars_pericone_presence / absent pericone_present, 72 Upper_M1_and_M2_diastema_presence / no_diastema_present_between_M1_and_M2_teeth_in_contact diastema_present, 73 Upper_M1_and_M2_lingual_cingulum / absent present, 74 Upper_M1_and_M2_stylar_shelf_presence / stylar_shelf_present_on_M1_and_M2 absent, 75 Upper_M1_and_M2_protocone_presence / present absent, 76 Upper_M1_and_M2_protocone_and_paracone_relative_height / protocone_shorter_than_paracone protocone_and_paracone_subequal_in_height, 77 Upper_M1_and_M2_metacone_and_paracone_relative_height / metacone_taller_than_paracone metacone_and_paracone_subequal_in_height metacone_shorter_than_paracone, 78 Upper_M1_and_M2_protocone_sharpness_and_inclination / less_sharp_and_less_inclined_anteriorly more_sharp_and_more_inclined_anteriorly, 79 Upper_M1_and_M2_metacone_and_paracone_inclination / not paracone_and_metacone_lingually_inclined, 80 Upper_M1_and_M2_angle_between_postparacrista_and_premetacrista / very_acute_cristae_nearly_or_completely_parallel_angle_is_20_degrees_or_less_ acute_cristae_lie_at_acute_angle_to_one_another_angle_21_90_degrees_ oblique_cristae_meet_at_an_angle_somewhat_more_than_a_right_angle_angle_91_120_degrees_ very_oblique_cristae_nearly_or_completely_form_straight_line_angle_well_over_120_degrees_, 81 Upper_M1_and_M2_postparacrista_and_premetacrisita_contact / postparactista_contacts_premetacrista_on_labial_aspect_of_tooth_trigon_closed_labially postparacrista_does_not_contact_premetacrista_trigon_open_labially, 82 Upper_M1_and_M2_trigon_width_and_depth / wide_and_shallow narrow_and_deep, 83 Upper_M1_and_M2_mesostyle_presence_and_location / mesostyle_absent mesostyle_formed_at_labial_terminus_of_postparacrista mesostyle_formed_at_point_of_contact_of_postparacrista_and_premetacrista mesostyle_formed_at_labial_terminus_of_premetacrista, 84 Upper_M1_and_M2_mesostyle_size_and_labial_extent / large_and_extended_labially small_and_less_labially_extended, 85 Upper_M1_and_M2_lingual_displacement_of_contact_between_postparacrista_and_premetacrista_ / not_lingually_displaced_lies_in_line_with_parastyle_and_metastyle somewhat_lingually_displaced strongly_lingually_displaced, 86 Upper_M1_and_M2_mesostylar_crest_presence / absent present, 87 Upper_M1_and_M2_mesostylar_crest_direction / projects_mesiolabially projects_distolabially, 88 Upper_M1_and_M2_cuspule_on_premetacrista_presence / cuspule_absent cuspule_present_on_premetacrista, 89 Upper_M1_and_M2_paraconule_presence / paraconule_present absent, 90 Upper_M1_and_M2_metaloph_presence / Metaloph_present absent, 91 Upper_M1_and_M2_secondary_cusp_on_postprotocrista / absent present, 92 Upper_M1_parastyle_presence / present absent, 93 Upper_M1_parastyle_form / conical crest, 94 Upper_M1_parastyle_separation_from_preparacrista / not_separated separated, 95 Upper_M1_parastyle_length / short long, 96 Upper_M1_parastyle_curvature / straight curved, 97 Upper_M1_ectoflexus / ectoflexus_absent single_ectoflexus_present_labial_to_mesostyle single_ectoflexus_present_between_mesostyle_and_parastyle_pre_ectoflexus_ single_ectoflexus_present_between_mesostyle_and_metastyle_postectoflexus_ pair_of_ectoflexi_present_one_between_mesostyle_and_parastyle_one_between_mesostyle_and_metastyle, 98 Upper_M1_single_ectoflexus_development / shallow_ectoflexus_present deep_ectoflexus_present, 99 Upper_M1_double_ectoflexus_development / both_shallow anterior_deep_posterior_shallow anterior_shallow_posterior_deep both_deep, 100 Upper_M1_preparacrista_presence / present absent, 101 Upper_M1_relative_length_of_preparacrista_and_postparacrista / preparacrista_longer_than_postparacrista preparacrista_subequal_in_length_to_postparacrista preparacrista_shorter_than_postparacrista, 102 Upper_M1_relative_length_of_postmetacrista_and_premetacrista / postmetacrista_length_subequal_to_premetacrista_length postmetacrista_longer_than_premetacrista_but_not_twice_as_long postmetacrista_2x_as_long_as_premetacrista_or_longer, 103 Upper_M1_trigon_basin_surface_texture / enamel_smooth surface_with_striations_or_small_enamel_folds, 104 Upper_M1_postprotocrista_orientation / oriented_distolabially_toward_metacone_metaconule_or_metastylar_region oriented_toward_midpoint_between_metastyle_metacone_and_hypocone_region oriented_anteroposteriorly_toward_hypocone_or_hypocone_region, 105 Upper_M1_postprotocrista_extent_ / postprotocrista_extends_only_to_metaconule_region_regardless_of_presence_absence_of_metaconule_does_not_reach_metacone_or_metacingulum joins_lingual_base_of_metacone postprotocrista_extends_to_hypoconal_region extends_to_become_confluent_with_metacingulum_or_edge_of_hypocone_shelf, 106 Upper_M1_metaconule_presence / metaconule_present absent, 107 Upper_M1_endoloph_presence / absent present_approaches_postprotocrista_at_an_angle present_but_continuous_with_postprotocrista_as_a_single_straight_crest_along_lingual_side_of_tooth present_does_not_approach_postprotocrista_runs_lingually, 108 Upper_M1_endoloph_contact_with_postprotocrista / endloloph_contacts_postprotocrista_ no_contact, 109 Upper_M1_hypocone_shelf_development / hypocone_shelf_very_large_maximum_mesiodistal_lenght_equal_to_or_greater_than_distance_between_paracone_and_metacone hypocone_shelf_large_maximum_mesiodistal_length_greater_than_half_the_distance_between_paracone_and_metacone_but_not_100%_of_that_distance hypocone_shelf_small_well_developed_and_clearly_distinct_from_trigon_maximum_mesiodistal_length_less_than_half_the_distance_between_the_paracone_and_metacone hypocone_shelf_very_small_present_as_a_cingulum_but_may_not_be_clearly_distinguished_from_the_trigon hypocone_shelf_absent, 110 Upper_M1_hypocone_shelf_width / narrower wider, 111 Upper_M1_hypocone_development / hypocone_absent hypocone_incipient_or_poorly_developed_but_not_entirely_absent hypocone_well_developed, 112 Upper_M2_presence / M2_present absent, 113 Upper_M2_parastyle_presence / present absent, 114 Upper_M2_parastyle_form / conical crest, 115 Upper_M2_parastyle_length / short long, 116 Upper_M2_parastlye_curvature / straight curved, 117 Upper_M2_ectoflexus / ectoflexus_absent single_ectoflexus_present_labial_to_mesostyle single_ectoflexus_present_between_mesostyle_and_parastyle_pre_ectoflexus_ single_ectoflexus_present_between_mesostyle_and_metastyle_postectoflexus_ pair_of_ectoflexi_present_one_between_mesostyle_and_parastyle_one_between_mesostyle_and_metastyle, 118 Upper_M2_single_ectoflexus_development / shallow_ectoflexus deep_ectoflexus, 119 Upper_M2_double_ectoflexus_development / both_shallow anterior_ectoflexus_deep_posterior_extoflexus_shallow. both_deep, 120 Upper_M2_position_of_metacone / metacone_in_normal_position metacone_shifted_lingually, 121 Upper_M2_relative_length_of_postmetacrista_and_premetacrista / postmetacrista_absent postmetacrista_shorter_that_premetacrista postmetacrista_length_subequal_to_premetacrista_length postmetacrista_much_longer_than_premetacrista_but_not_twice_as_long postmetacrista_2x_longer_than_premetacrista_or_longer, 122 Upper_M2_orientation_of_postprotocrista / oriented_distolabially_toward_metacone_metaconule_or_metastylar_region oriented_toward_midpoint_between_metastyle_metacone_and_hypocone_region oriented_anteroposteriorly_toward_hypocone_or_hypocone_region, 123 Upper_M2_endoloph_presence / absent present_approaches_postprotocrista_at_an_angle present_but_continuous_with_postprotocrista_as_a_single_straight_crest_along_lingual_side_of_tooth present_does_not_approach_postprotocrista_runs_lingualy, 124 Upper_M2_endoloph_contact_with_postprotocrista / postprotocrista_and_endoloph_at_an_angle postprotocrista_and_endoloph_at_an_angle_but_not_in_contact, 125 Upper_M2_metaconule_presence / metaconule_present absent, 126 Upper_M2_hypocone_shelf_development / hypocone_shelf_very_large_maximum_mesiodistal_lenght_equal_to_or_greater_than_distance_between_paracone_and_metacone hypocone_shelf_large_maximum_mesiodistal_length_greater_than_half_the_distance_between_paracone_and_metacone_but_not_100%_of_that_distance hypocone_shelf_small_well_developed_and_clearly_distinct_from_tricon_maximum_mesiodistal_length_less_than_half_the_distance_between_the_paracone_and_metacone hypocone_shelf_very_small_present_as_a_cingulum_but_may_not_be_clearly_distinguished_from_the_trigon hypocone_shelf_absent, 127 Upper_M2_hypocone_shelf_width / narrower wider, 128 Upper_M2_hypocone_development / hypocone_absent hypocone_incipient_or_poorly_developed_but_not_entirely_absent hypocone_well_developed, 129 Upper_M3_presence / M3_present absent, 130 Upper_M3_reduction / little_reduced_M3_occlusal_area_75%_of_M2_area_ moderately_reduced_in_length_and_or_width_M3_area_50%_75%_of_M2_area_ very_reduced_in_length_and_or_width_M3_area_is_25%_but_50%_of_M2_area_ extremely_reduced_10%_M3_area_25%_M2_area_ peglike_10%_area_, 131 Upper_M3_anterior_segment_size / M3_anterior_segment_nearly_as_large_as_that_of_M2 much_smaller_than_that_of_M2, 132 Upper_M3_parastyle_presence / present absent, 133 Upper_M3_parastyle_form / conical crest, 134 Upper_M3_parastyle_length / short long, 135 Upper_M3_parastyle_curvature / straight curved, 136 Upper_M3_paraconule_presence / paraconule_present absent, 137 Upper_M3_postmetacrista / postmetacrista_present_W_shaped_ectoloph_complete_ postmetacrista_absent_ectoloph_N_shaped_or_V_shaped_, 138 Upper_M3_premetacrista_development / premetacrista_longer_than_postparacrista premetacrista_and_postparacrista_length_subequal premetacrista_one_half_to_three_quarters_the_length_of_the_postparacrista premetacrista_one_third_to_slightly_less_than_one_half_the_length_of_the_postparacrista premetacrista_present_but_less_than_one_third_the_length_of_the_postparacrista premetacrista_absent_, 139 Upper_M3_metacone / metacone_present_as_a_disinct_cusp metacone_indistinct_incorperated_into_premetacrista metacone_entirely_absent, 140 Upper_M3_metastylar_fovea_presence / fovea_present absent, 141 Upper_M3_protocone_presence / present absent, 142 Upper_M3_hypocone_shelf_presence / absent present, 143 Lower_i1_presence / i1_present absent, 144 Lower_i1_morphology_of_cutting_edge / lobed_bilobed_or_trilobed_or_with_four_lobes_ undulate_lobe_structure cutting_edge_roughly_straight_without_notches_or_grooves pointed peglike_no_real_cutting_edge, 145 Lower_i1_number_of_lobes / bilobed trilobed four_lobes, 146 Lower_i1_trilobed_morphology / three_lobes_of_equal_size mesial_lobe_very_small_middle_lobe_large_distal_lobe_intermediate_in_size mesial_lobe_large_middle_and_distal_small two_mesial_lobes_small_and_twinned_well_separated_from_a_larger_distal_lobe, 147 Lower_i1_crown_proporitions / width_greater_than_length length_and_width_subequal length_greater_than_width, 148 Lower_i1_diastema_between_right_and_left_teeth / absent diastema_present, 149 Lower_i2_presence / i2_present absent, 150 Lower_i2_morphology_of_cutting_edge / lobed_bilobed_or_trilobed_or_with_more_lobes_ cutting_edge_roughly_straight_without_notches_or_lobes pointed peglike_no_real_cutting_edge, 151 Lower_i2_number_of_lobes / bilobed trilobed four_lobes more_than_four, 152 Lower_i1_and_i2_relative_size / i2_conspicuously_smaller_than_i1 i1_and_i2_subequal_in_size i2_conspicuously_larger_than_i1, 153 Lower_i1_and_i2_position / i2_lateral_to_i1 i2_posterior_to_i1, 154 Lower_i3_presence / i3_present absent, 155 Lower_i3_crown_height / i3_height_noticably_less_than_i2_height i3_height_subequal_to_i2_height i3_height_noticably_greater_than_i2_height, 156 Lower_incisor_alveoli_spacing / alveoli_of_i1_and_i2_nearly_coalesced_small_gap_between_i2_and_i3 alveoli_of_i1_i2_and_i3_evenly_spaced, 157 Lower_incisors_procumbent / not_procumbent somewhat_procumbent more_procumbent, 158 Lower_c_anterior_diastema / absent diastema_present_between_canine_and_last_lower_incisor_i1_i2_or_i3_, 159 Lower_c_tip_divergence / lower_canines_not_laterally_divergent canines_laterally_divergent_shafts_slanted_outward_so_that_the_tip_of_each_tooth_lies_dorsal_or_dorsal_and_lateral_to_the_lateral_edge_of_the_root_where_the_tooth_enters_the_alveolus, 160 Lower_c_space_between_base_of_left_and_right_canines / lower_canines_well_separated_from_one_another_at_the_base bases_of_right_and_left_canines_adjacent_or_nearly_so_but_not_in_contact bases_of_right_and_left_canines_adjacent_and_in_contact, 161 Lower_c_shape / extremely_short_premolariform_ short_and_stocky tall_and_lancoelate exceptionally_slender_with_a_subterete_shaft bladelike, 162 Lower_c_anterior_cuspule_presence / absent anterior_cuspule_present, 163 Lower_c_posterior_cingulid / cingulid_absent cingulid_present_but_small cingulid_broad, 164 Lower_c_posterobasal_cusp / absent weakly_developed strongly_developed, 165 Lower_c_labially_swollen / not lower_c_labially_swollen, 166 Lower_c_labial_cingulum / labial_cingulum_absent weakly_developed well_developed, 167 Lower_c_labial_cingular_structures / present absent, 168 Lower_p1_presence_ / p1_present absent, 169 Lower_p1_number_of_roots / one_root two_roots, 170 Lower_p1_shape / large_and_cuspidate peg_like_with_flattened_crown tiny_spicule_with_undifferentiated_crown, 171 Lower_p1_distal_cuspule / absent cuspule_present, 172 Lower_p1_and_p4_relative_crown_length / p1_longer_than_p4 subequal p1_shorter_than_p4, 173 Lower_p4_presence / p4_present absent, 174 Lower_p4_number_of_roots / three_roots two_roots one_root, 175 Lower_p4_orientation_of_roots_when_two_are_present / two_roots_oriented_longitudinally_with_resepct_to_the_long_axis_of_the_tooth_row two_roots_oriented_slightly_obliquely two_roots_oriented_stongly_obliquely_to_the_long_axis_of_the_toothrow, 176 Lower_p4_alignment_within_toothrow / p4_aligned_in_row_with_surrounding_teeth_when_seen_in_occlusal_view p4_lingually_displaced_from_toothrow_not_in_line_with_other_teeth p4_labially_displaced_from_toothrow_not_in_line_with_other_teeth, 177 Lower_p4_anterior_diastema / no_diastema_between_p4_and_next_adjacent_anterior_tooth_p1_or_C_ conspicuous_diastema_present, 178 Lower_p4_overall_morphology / p4_tribosphenic_with_a_distinct_trigonid_and_talonid p4_premolariform_with_cusps_but_not_arranged_in_tribosphenic_pattern p4_peglike_or_nearly_featureless, 179 Lower_p4_anterolingual_cuspule / absent cuspule_present, 180 Lower_p4_lingual_cuspule / absent cuspule_present, 181 Lower_p4_distal_cuspule / absent cuspule_present, 182 Lower_p4_posterior_crests / two_posterior_crests_separated_by_a_concavity only_one_posterior_crest_present_, 183 Lower_p4_occlusal_outline / long_and_strongly_laterally_compressed_length_width_or_1.7_ short_and_broad_length_width_1.7_, 184 Lower_p4_labial_cingulid_development / labial_cingulid_well_developed weakly_developed absent, 185 Lower_p4_and_p5_relative_crown_height / p4_much_taller_than_p5 p4_slightly_taller_than_p5 p4_and_p5_subequal_in_height p4_slightly_shorter_than_p5 p4_very_small_much_shorter_than_p5, 186 Lower_p4_and_p5_relative_crown_length / crown_length_of_p4_greater_than_length_of_p5 p4_and_p5_subequal_in_length p4_length_less_than_p5_length_but_more_than_one_half_the_length_of_p5 p4_less_than_or_equal_to_one_half_the_length_of_p5, 187 Lower_p4_and_p5_crown_morphology / crown_morphology_of_p4_and_p5_similar p4_and_p5_with_different_crown_morphologies, 188 Lower_p5_number_of_roots / three_roots two_roots one_root, 189 Lower_p5_orientation_of_roots_when_two_are_present / two_roots_oriented_longitudinally_with_resepct_to_the_long_axis_of_the_tooth_row two_roots_oriented_slightly_obliquely_ two_roots_oriented_stongly_obliquely_to_the_long_axis_of_the_toothrow, 190 Lower_p5_paraconid / paraconid_large_and_high small_and_low very_low cusp_absent, 191 Lower_p5_paracristid_presence / paracristid_present absent, 192 Lower_p5_lingual_cingulid / absent partial_lingual_cingulum_present complete_lingual_cingulum_present, 193 Lower_p5_labial_cingulid_development / labial_cingulid_strongly_developed weakly_developed absent, 194 Lower_p5_labial_cingulid_form / straight undulate, 195 Lower_p5_labial_cingulum_thickness / cingulum_of_constant_thickness_throughout_length thicker_toward_distal_end, 196 Lower_p5_presence_of_lingual_lobe / absent present, 197 Lower_p5_extension_of_lingual_lobe / lingual posterior, 198 Lower_p5_lingual_lobe_cusp / absent present, 199 Lower_p5_lingual_cuspule / absent Cuspule_present, 200 Lower_p5_metaconid / absent one_metaconid two_metaconids, 201 Lower_p5_metaconid_development / metaconid_large metaconid_present_but_small, 202 Lower_p5_protocristid / absent one_protocristid two_protocristids, 203 Lower_p5_postmetacristid / absent short long, 204 Lower_p5_talonid_length / talonid_long talonid_short absent, 205 Lower_p5_protoconid_development / less_robust_taller_sharper more_robust_shorter_more_blunt extremely_short, 206 Lower_p5_mesial_cuspule / absent cuspule_present, 207 Lower_p5_overall_morphology / p5_triosphenic_with_distinct_trigonid_and_talonid p5_premolariform_with_cusps_but_not_arranged_in_tribosphenic_pattern p5_without_distinct_cusps_may_be_nearly_featureless_or_peglike, 208 Lower_p5_and_m1_relative_length / p5_and_m1_subequal_in_length p5_length_less_than_m1_length_but_more_than_three_quarters_the_length_of_m1 p5_length_more_than_one_half_but_less_than_three_quarters_the_length_of_m1 p5_length_less_than_or_equal_to_one_half_the_length_of_m1, 209 Lower_molar_cusp_morphology / cusps_well_individualized_concial_or_comprised_of_crests_that_meet_to_form_a_sharp_point cusps_not_well_individualized_not_very_conical_or_pointed, 210 Lower_molars_lingual_cingulid_presence / absent lingual_cingulid_present, 211 Lower_molars_trigonid_fovea / trigonid_fovea_closed trigonid_fovea_open_lingually, 212 Lower_molars_position_of_hypoconulid / hypoconuiid_located_on_distal_border_of_talonid_midway_between_hyponid_and_entoconid_ hypoconulid_located_on_distolabial_border_of_talonid_well_separated_from_entoconid_closer_to_midway_point_than_to_entoconid_ hypoconulid_located_on_distolabial_border_of_talonid_well_separated_from_entoconid_but_closer_to_entoconid_than_to_midway_point hypoconulid_located_on_labial_border_of_talonid_immediately_posterior_to_and_directly_adacent_to_the_entoconid_hypoconulid_and_entoconid_may_be_partially_fused_at_their_bases_twinning_of_hypoconuilid_and_entoconid_., 213 Lower_molars_nyctalodont_or_myotodont / nyctalodont myotodont, 214 Lower_molars_with_cristid_between_entoconid_and_midline_hypoconulid / cristid_present absent, 215 Lower_molars_talonid_basin_depth / basin_absent shallow deep, 216 Lower_molars_talonid_anteroposterior_compression / not talonid_anteroposteriorly_compressed, 217 Lower_molars_lingual_cusp_relative_height / relatively_low_ relatively_high_lingual_cusps_at_least_75%_height_of_labial_cusps_, 218 Lower_molars_notches_in_cristid_obliqua_and_postcristid / notches_absent notches_weakly_developed notches_strongly_developed_carnassia_like, 219 Lower_molars_notches_in_paracristid_and_protocristid / Notches_absent Notches_weakly_developed Notches_strongly_developed, 220 Lower_molars_cristid_obliqua_curature_or_flexion / straight gently_curved flexed, 221 Lower_molars_orientation_of_cristid_obliqua_ / towards_protoconid directed_toward_a_point_in_between_protoconid_and_metaconid towards_metaconid, 222 Lower_molars_cristid_obliqua_termination_on_metaconid / cristid_obliqua_terminates_at_base_of_metaconid cristid_obliqua_extends_up_flank_of_metaconid, 223 Lower_molars_groove_paralleling_cristid_obliqua / absent groove_present, 224 Lower_molars_entocristid_flexion / not labially_flexed, 225 Lower_molars_entocristid_distrupted / entocristid_contacts_postvalid_thus_closing_off_lingual_side_of_talonid_basin entocristid_short_disrupted_so_that_talonid_basin_drains_lingually entocristid_absent, 226 Lower_molars_height_of_protoconid_and_hypoconid / not unusually_tall_and_robust, 227 Lower_molars_protoconid_and_hypoconid_anteroposteriorly_compressed / not_compressed protoconid_and_hypoconid_anteroposteriorly_compressed, 228 Lower_molars_protoconid_and_hypoconid_ventral_extension_of_base / not protoconid_and_hypoconid_with_ventral_extension_of_labial_cusp_base, 229 Lower_molars_protoconid_and_hypoconid_buccal_bulging / absent prononounced_buccal_bulging_present, 230 Lower_m1_and_m2_extent_of_cristid_obliqua / does_not_reach_trigonid reaches_trigonid, 231 Lower_m1_and_m2_hypoconulid_size / large small_and_low absent, 232 Lower_m1_and_m2_entocristid_height / low_crest high_crest, 233 Lower_m1_and_m2_relative_length / m1_longer_than_m2 m1_and_m2_subequal m1_shorter_than_m2, 234 Lower_m1_relative_height_of_protoconid_and_hypoconid / protoconid_markedly_higher_than_hypoconid protoconid_slightly_higher_than_hypoconid protoconid_and_hypoconid_subequal_in_height protoconid_slightly_smaller_than_hypoconid, 235 Lower_m1_protoconid_height_compared_to_subsequent_molars / m1_protoconid_higher_than_m2_or_m3_protoconid m1_protoconid_subequal_in_height_to_protoconids_on_m2_and_m3 m1_protoconid_noticably_shorter_than_protoconids_on_m2_and_m3, 236 Lower_m1_paraconid_presence / paraconid_present_on_m1 absent, 237 Lower_m1_metaconid_presence / metaconid_present_on_m1 absent, 238 Lower_m1_relative_height_of_paraconid_and_metaconid / paraconid_markedly_taller_than_metaconid cusps_subequal_in_height paraconid_markedly_shorter_than_metaconid, 239 Lower_m1_presence_of_metacristid / present absent, 240 Lower_m1_paracristid_and_metacristid_relative_length / paracristid_markedly_shorter_that_metacristid paracristid_and_metacristid_subequal_in_length paracristid_markedly_longer_than_metacristid, 241 Lower_m1_entoconid_development / strongly_developed_tall weakly_developed_low entoconid_absent, 242 Lower_m1_entocristid_accessory_cusp / absent present, 243 Lower_m1_labial_cingulid_development / strongly_developed weakly_developed absent, 244 Lower_m1_labial_cingulid_completeness / labial_cingulid_complete incomplete_posteriorly incomplete_anteriorly, 245 Lower_m1_precingulum / absent present, 246 Lower_m1_precingulid_and_labial_cingulid_connection / not_connected cingulids_confluent_around_mesiolabial_corner_of_tooth, 247 Lower_m1_alignment_of_lingual_cusps / paraconid_metaconid_and_entoconid_aligned_in_a_straight_line_ paraconid_situated_labial_to_a_line_drawn_through_the_metaconid_and_entoconid, 248 Lower_m1_angle_between_protoconid_and_metaconid / large_angle intermediate_angle small_angle, 249 Lower_m1_talonid_width / talonid_wider_than_trigonid talonid_and_trigonid_subequal talonid_narrower_than_trigonid, 250 Lower_m1_hypoflexid_depth / deep shallow absent, 251 lower_m1_length_width_ratio / 1.10_1.44 1.45_1.74 1.75_2.04 2.05_2.19 2.20_2.70 3.00_4.60, 252 Lower_m2_and_m3_relative_height_of_paraconid_and_metaconid / subequal_in_height paraconid_slightly_shorter_than_metaconid paraconid_markedly_shorter_than_metaconid, 253 Lower_m2_and_m3_precingulum / absent present, 254 Lower_m2_and_m3_precingulid_and_labial_cingulid_connected. / not_connected cingulids_confluent_around_mesiolabial_corner_of_tooth, 255 Lower_m2_presence / m2_present absent, 256 Lower_m2_crown_morphology / m2_with_crown_structure_resembling_m1 simplified_cuspidate_structure_compared_to_m1 m2_peg_like_with_flattened_crown, 257 Lower_m2_height_of_trigonid_relative_to_talonid_cusps / protoconid_markedly_higher_than_hypoconid protoconid_slightly_higher_than_hypoconid protoconid_and_hypoconid_subequal_in_height, 258 Lower_m2_labial_cingulid_development / strongly_developed weakly_developed absent, 259 Lower_m2_labial_cingulid_completeness / labial_cingulid_complete incomplete_posteriorly incomplete_anteriorly, 260 Lower_m2_entoconid_development / strongly_developed_tall weakly_developed_low very_weak_small absent, 261 Lower_m2_talonid_width / talonid_wider_than_trigonid talonid_and_trigonid_subequal talonid_narrower_than_trigonid, 262 Lower_m2_hypoflexid_depth / deep shallow absent, 263 Lower_m3_presence / m3_present absent, 264 Lower_m3_crown_morphology / m3_crown_morphology_similar_to_that_of_m2 m3_markedly_different_from_m2, 265 Lower_m3_length / m3_length_subequal_to_that_of_m2 m3_slightly_shorter_75%_than_m2 m3_length_much_less_than_m2, 266 Lower_m3_number_of_roots / 2_roots 1_root, 267 Lower_m3_labial_cingulid_development / strongly_developed weakly_developed absent, 268 Lower_m3_labial_cingulid_completeness / labial_cingulid_complete incomplete_posteriorly incomplete_anteriorly, 269 Lower_m3_extent_of_cristid_obliqua / reaches_trigon present_but_does_not_reach_trigon cristid_obliqua_absent, 270 Lower_m3_hypoconid_presence / present absent, 271 Lower_m3_height_of_protoconid_relative_to_hypoconid / protoconid_notably_higher_than_hypoconid protoconid_slightly_higher_than_hypoconid protoconid_and_hypoconid_subequal_in_height protoconid_smaller_than_hypoconid, 272 Lower_m3_hypoflexid_depth / absent shallow deep, 273 Lower_m3_entocristid_orientation / transverse_to_long_axis_of_tooth oblique_to_long_axis_of_tooth parallel_to_long_axis_of_tooth, 274 Lower_m3_presence_of_postcristid / present absent, 275 Lower_m3_postcristid_orientation / directed_posterolabially_at_an_oblique_angle_to_the_long_axis_of_the_tooth perpendicular_to_the_long_axis_of_the_tooth, 276 Lower_m3_hypoconulid_presence / present absent, 277 Lower_m3_hypoconulid_size / large small_and_low, 278 Lower_m3_talonid_width / talonid_wider_than_trigonid talonid_and_trigonid_subequal talonid_narrower_than_trigonid, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Anoura_caudifer 00010--1--0-0001000010010---3-0110--000010200102001111001002-1101200220000001001103000-0110011114-202002110-300000--4-00200-14--00000--10100001-----1----1----10201000-01011010011101102320100002--0--020100001100130-2001201-00000001100010021-002-0-1101220-0012-00100002-0001201010 Anoura_cultrata 00001-----0-1000000010210---3-111200000000200102001111002002-2101201120000001001100-0110110010004-202102110-30001---4-00200-14--00000--10100001-----1----1----10200100-01000010011101102320100002--0--020100001100130-2001100-00000001100010021-002-0-1101220-0012-00100002-0001201010 Anoura_geoffroyi 00001-----0-0001000010110---3-111201000000200102001011002022-2101201120000001001100-0110110010104-002102312-200001004-00222-120000000--10100001-----1----1----10201000-01012010011101102210101002--0--020100001100130-2001101-00000001100110021-002-0-1101320-0012-00100002-0001201010 Anoura_geoffroyi_peruana 00001-----0-1001000010010---3-1111-1000010200102001011102012-210120112000000100110300110110010004-002102310-20001---4-10200-120000000--10200001-----1----1----10201000-01012011011101102210100002--0--020100001000130-2001101-00000001100110021-012-0-1101320-0012-00100002-0001201011 Anoura_latidens 00001-----0-0001000010110---3-1111-0000000200102001011002012-2101201120000001001100-0110110010004-002002312-20001---4-10202-120000000--10200001-----1----1----10201000-01010010011101102220100002--0--010100001100130-2001100-00000001100110021-012-0-1101320-0012-00100002-0001201011 Artibeus_gnomus 0000200---0120000010001030001-0210--1003----0201211010002102-0110000220001002003000-00-01101----0--01011003-11201---10-1013-02021-------------000-10001101--0000302000-0000-1--------------103002--0--010122001300130-1010010-001000010-00010-1-012-0--002020-0002-01201210021-0-1-1-2 Artibeus_jamaicensis 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00001-----103001011000003100210100--1003----0201201000002102-0110000220010001001010-0110110010004-2021013110100001004-203110111002001011152-01000-201----1--000020220210001-1--------------103121000---10102001200130-2000101-01000001101010021-000011010002110000000000101010021011-2 Mimon_crenulatum_keenani 00001-----003001011000003200211111--1003----021121110000310100110000220000001001010-0110110010014-3021013110101001004-203110101002001011152-010010101----1--000020220110000-1--------------103120000---11102001200130-2001101-01000011101010021-000011010002110000000000101000021011-2 Mimon_crenulatum_longifolium 10001-----0030010110000132002{12}1110--1003----021021110000210100110000220010001001012100-0110010004-202101010-101001104-203110110002001011152-010010201----1--000020220210000-1--------------103120000---11102001200130-2001101-01000001111010021-000011010002110000000000100000021011-2 Monophyllus_plethodon 00001-----011001000010210---3-1111--1003----0102001011102002-2111201120000001001010-00-0110010014-002100110-4--0010120-0200-14--0000101101000104--110????1--0110202100-00011010001101102310111002--0--011101001100131-2010101-00000001101010021-002-0-1201220-0002-00100102-0001201011 Monophyllus_redmani 00001-----011001000010210---3-1111--1003----010200101111{23}002-2110201120000001001{01}10-00-0110010014-002000110-4--001004-00200-14--0000100101000004--1103-111--0110202010-00012010001101102210100102--0--011101001000131-2010101-00000000101010021-002-0-1201320-0002-00100102-0001201010 Mormoops_blainvillei 0000220---003001000010212000211111--1003----000020100100310100110200120010001001000-00-0110010014-302101110-100001114-202110101001001111110-00001020001101--1010202202100001010001001100310113121100---11102011100130-2001101-00000000101010021-00100-00002211000100000000101002201001 Musonycteris_harrisoni 00001-----0-10011-0010000---3-1112000003----01020010111100-2-1110210120101001003100-00-01101----0--02100110-4--01---0--0200-14--0001---00101001-----1----1----10200000-01011011011101102310111102--0--01010210100010-01010001-001000000-1010021-102-0-0201520-0002-20100102-1001200001 Neonycteris_pusilla 10001-----1030010011001030000-2211--0103----010021110102213001110200200000000001010-10-01101----4-002202312-111001004-00322-111102001011152-01001020001101--0000102200-00011010011000110120103120010---11102101200130-1002201-00000001002020011-00001111002211000000000010000002101012 Noctilio_albiventris 10000--0101030000100002120000-1110--1003----1-----------311110110001220000000001012000-0010010014-0020001010112001014-003010011201001011120-0001--101----1--001120200210001-1--------------0-102101111-0-1-2101200131-1010012100000011111210020100100-000002110021100000101000021001-2 Noctilio_leporinus 10000--010103000010000{12}120000-1110--1003----1-----------311100110001220000000001012000-0110010014-0020001111002001014-002011110201001011120-0001--101----1--001120200210001-1--------------0-102100111-0-1-2101200131-10000121000000111110100201001011000002110011100000101000021001-2 Notonycteris_magdalenensis ??????????????????????????????????????????????????????????????????????0?00000001010--0-0110????????????????????000--11-0400-1012???????????????????????????????????????00???02-0???????????10???????????????????00130-2002201-0000000101?0?0021-1000111?001211000000100??0???????????? Notonycteris_sucharadeus ????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????00130-2002201-0000000101?0?0021-0000111?00121100000000???????????????? Palynephyllum_antimaster_2 ?????????????????????????????????????????????????????????????????????2??000010010?0-0???1?0????????????????????01---30-022??14--?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Phylloderma_stenops 0100220---1030010111000020003-0110--1003----0?0111110000100100110201220000001001010-00-0110010014-002102310-110001014-20320-111002000--1152-0102--1001-011--000010200110000002-002000-10431103120100---11102001200130-2002201-01000001111010021-00001111001211000000000010000002101012 Phylloderma_stenops_septentrionalis 0000210---103000011100002100210100--0103----000110101001200100110201220000001001010-0110110010014-202102310-111001014-10320-111002001011152-0102--0001-001--000010200110000002-002000-11431103120100---11102101100130-2002201-01000001101010021-000011020022110000000000100000022011-2 Phyllonycteris_aphylla 0000220---012001000010112100210110--1003----02020011010000-2-01102112200011-1-03002100-01101----10-0110--10-11001---4-002-0-10100101---1101-1-04--1103-211--1110202000-0000-1--------------103102--0--00-2-11012101---1010000-000000012101010-1-2-2-0--1022-0-0022-21200102-01-02001-1 Phyllonycteris_major ??0???????????00??0?101?2?00??0110--1003----02020011010000-2-01102012201011-1-03002100-01101----10-0110--10-01001---4-002-0-10100101---1101-1-0????10???11--??10202100-00??-1--------------1?3102--0--00-2-0101?101---1010000-0000000121???????????????????-0-0?12-21200102-01-02001-1 Phyllonycteris_poeyi 0100220---0030010000101111-0210200--1003----0202001101001102-01102112200011-1-03002100-01101----20-0100--10-01001---4-002-0-10100101---1101-1-04--1103-211--1110202100-0000-1--------------103101100--00-2-01012101---1010000-00000001210000021-2-2-0--0022-0-0022-20200102-01-01001-1 Phyllostomus_discolor 11001-----1030010010000122003-0100--1003----0000111100003112-0110201220000001001010-00-0110010014-002100110-200001004-00300-110102001001152-0102--0001-101--101020220110000-1--------------103111100---11102001000130-2001201-00000001101010021-002-1-110022110001000000101110021011-2 Phyllostomus_discolor_verrucosus 10001-----0030000010000121003-0100--0103----0001100-00002112-0110200220000001001010-00-0110010014-202101010-200001004-10300-110002000--1152-0102--0001-101--1010202200-0000-1--------------103101100--011102001100130-2001101-00000001101010021-000011010012110001000000100000021011-2 Phyllostomus_elongatus 11001-----1030010110000130002{01}0100--0003----000110100000212100110001220000001001010-00-0110010014-2021013110111001004-20310-111102001011152-0100100001-101--001020220110000-1--------------113120000---11102001100130-2001101-00000001101010021-000011010002110000000000100000021011-2 Phyllostomus_hastatus 11001-----1030010010000120001-0101--1003----000110100000212000111101210000001001010-00-0110010004-202101010-112001014-00310-111202001011152-01001010000101--001020210210000-1--------------113100100--011102001100130-2001101-00000001101010021-000011010002110000000000100000021001-2 Phyllostomus_hastatus_panamensis 11001-----103001001000012000210101--0003----000110100000211100110100210000001001010-00-0110010004-2021010111112001004-203111111202001011152-01001000000101--001020210110000-1--------------103110100---11112001200130-2001101-00000001101010021-000011010002110000000000100000021011-2 Platalina_genovensium 10001-----0010011-0010000---3-1111--0013----01021111010000-2-1101211120101001003000-00-01101----0--02200110-4--01---0--0200-14--0201---101010002--2001-101--1110201100-01011010011000102310113102--0--0111021010001---1010011-0--000002-1010021-2-2-0--222520-0012-32200102-1001-1-1-2 Platyrrhinus_helleri 00001-----0130000000101020001-1211--1003----020021110100310000202100210011002003000-00-0110010010--01002310-4--01---0--1220-14--04-1---1152-1-000-11000101--0000202000-0001-1--------------11212000100-2010200120011-01010000-0-2000011-10001---0-020---02020-0000001200210000302011-1 Pteronotus_davyi 0000220---103001010000112100211110--1003----010020110000200101110000220010001001000-00-0110010114-001001310-0120011120-0210-101201001111100-00001000001001--001020220210000002-002000-10431113110100---11102001100130-1000001-01000001101210021-001011000011110001000000000110022011-2 Pteronotus_parnellii 0000220---103000011000012100211110--1003----010020100000200101110101220010011001000-00-01100101020-01001310-1120011120-0310-111201101111110-00001000000001--001120200210000002-102000-10431101101100--011101011200131-1001001-01000001101110021-00001100001211001000000010000001201012 Pygoderma_bilabiatum 00000--010113001001000103010210111--1103----0202211100003122-0210100120001002003000-10-01101----0--02011110-20201---0--1003-12001-------------000-00000201--000010200110000-1--------------2-100000100-11212111100130-1010000-0-2000011-010002000-2-1-10020-0-002000221--------------- Rhinophylla_alethina 11001-----103001000000110---3-0211--1103----02020011010000-1011112112201001-0-03000-20-011-000--10-0020--10-4--000--10-14-0-14--0311---1152-1-001020000001--001010110111----02-011000112310103102--0--00-1-21012001---0000120-0-2000012-10001---2-2-0---122-0-0002-31200102-0000-1-1-2 Rhinophylla_fischerae 1100221---103001000000110---3-0211--1103----02020011010000-1011112112201001-0-03000-20-011-1----10-0020--10-4--000--10-14-0-14--0311---1152-1-001020000001--001010110111----02-011000112310103102--0--0111021012001---0000120-0-2000012-10101---2-2-0---122-0-0002-31200102-0000-1-1-2 Rhinophylla_pumilio 11000--0101030010000001020003-1211--1103----02010011000000-1011112102201001-0-03000-20-011-1----10-0020--10-4--01---10-04-0-14--0311---1152-1-001020000011--001010110211----02-0010001123102-3102--0--0111021012001---0000120-0-2000012-10101---2-2-0---112-0-0002-31200102-0000-1-1-2 Sturnira_lilium 00001-----003001001000013100211110--0003----0202211010003102-0101000220001000003000-00-01101----10-1--02310-4--01---1--0020-14--0211---1120-0002--0001-101--0000102000-0001-1--------------2-1020000---102021012001---1010010-001000012-0000011-002-0-00021-0-0012-01201212-01-01001-2 Thyroptera_tricolor 10000--1--1020000100101030001-1100--10000120010021100000312101110100220010001001012100-0100010114-302002012-201001004-20312-120201001011100-00001020001110211010112002100012010001000100220103120100---11102101100131-2000201-010000000-121001020000111101100-001001000000001002-011-1 Tonatia_saurophila 10011-----1521010010000132001-0110--1003----02011011000021110011020120000000100100310110110010004-0020010111110001014-202110111002001011152-0102--101----1--000120210110000002-001000111431113121100---0-1-2001200130-2002201-00000001101110021-00001101002211000000000000000002201012 Tonatia_saurophila_bakeri 00001-----1020010010000132001-0111--1003----02011011000021110011020120000000100101210110110010004-0021013111110001014-103111111002001011152-0102--101----1--000120210110000002-001000111431113120100---0-1-2001200130-2001101-00000001101010021-00001101001211000000000010010002201012 Tonatia_saurophila_saurophila ???????????????????????????????????????3----020???????0?2001001102?????????????????????????????????????????????0????????????????0????????????????????????1--???????????00??002-0?????????3?113121100---0-1-2001200130-2002201-00000001101010021-00001101001211000000000??0???????????? Trachops_cirrhosus 1000220---1031020010000122001-0111--1003----02011010100010000011120121000000000101201110110010004-3022013110102001014-204110111202001011151-00001010001101--000020220110000002-102001-12431101020100---10102001100130-2000202000000001102220021-00001101001211000000000000101002100012 Trachops_cirrhosus_coffini 1000200---1031020010000122001-0111--1003----02001010100021000011120121000000000101201110110010004-2022001111102001004-204111111202001011152-00001010001101--001020220110000002-102000-12431111020100---11102001100130-2002201-00000001101220021-00001101000211000000100010101002101012 Trinycteris_nicefori 10001-----1030010011101020003-0111--1103----010111110110202102210201200010001001010-011011000---30-021000111111001004-003111111202001011152-01001010001101--0000102200-00010010001101100420100020000---21202001100130-2001201-01000001101120021-000011010111110000000000101000021011-2 Uroderma_bilobatum 0100210---112000010010112000211210--1003----020021200100310000202100120001002103000-00-01101----0--00011203-11201---0--1023-02010311---0152-0002--1101-101--0000212001-0001-1--------------110010000---10102001100130-1010000-001000011-10001---00100---02120-0000101100200001-22011-2 Vampyrodes_caraccioli 00001-----013001001010112000212210--1003----020021201000210000202001100001002003002100-00101----0--01011010-200001000--1020-12001-------------000-10000001--0000202000-0000-1--------------110100000--0202020011001---1010000-011000012-20001---00000---02020-0000101201210021-01001-2 Vampyrodes_major 01001-----013001000010112000211210--1003----020021101000210000202101200001002003001100-00101----0--00211010-200000--0--1020-12001-------------02--11000001--0100202000-0000-1--------------100100000--0202020011001---1010000-001000012-20001---00010---02010-0000102201210021-01001-2 Vampyrum_spectrum 00001-----142102011000012100211111--1003----010210101000300100112201220000000001012120-01100100111-01200110-010000--11-0400-101003101011152-0002--1001-101--0001302100-0000102-001000112431103122--0---11102001100130-2000201-00000001112020021-00101112013211000000210010100001100012 Xeronycteris_vieirai 10001-----0020001-0010010---3-121200001{03}102001020010111100-2-2101211020100001002000-10-0110010010--01100110-4--000--10-0400-14--01000--1020100001020001101--0110201000-01012010011101102210101002--0--0101221010001---1010001-0--000002-1110021-2-2-0--?21520-0012-32100102-1001-1-1-2 ; END; [ The following blocks are output data for analysis step 10991 ] BEGIN TREES; TITLE Chiroptera_Morphological_data; LINK TAXA = TaxaForAnalysisStep10991; TREE Supp._Fig._4B_MLT = [&R] (Thyroptera_tricolor:0.13573457579097195,(Mormoops_blainvillei:0.12182706843998141,(Pteronotus_davyi:0.03634095545656937,(Pteronotus_parnellii:0.08341586935012575,((Phylloderma_stenops:0.04779659158560122,Phylloderma_stenops_septentrionalis:0.08087708097585289)0.82:0.03702676500811124,(((Macrophyllum_macrophyllum:0.10378860857258379,(Lonchorhina_orinocensis:0.0682584878562408,(Lonchorhina_aurita:0.11358346767330509,Lonchorhina_inusitata:0.056737561458192184)0.73:0.042808180426192916)0.53:0.030702600700274547)0.51:0.0263421826136414,((Phyllostomus_elongatus:0.021433772563751237,((Phyllostomus_discolor:0.08539065035269301,Phyllostomus_discolor_verrucosus:0.04980967089660967)0.93:0.07968213612647361,(Phyllostomus_hastatus:0.054595803196291925,Phyllostomus_hastatus_panamensis:0.02164758759388403)0.43:0.030734769262975664)0.25:0.028761645323284762)0.65:0.05256663956969628,((Mimon_crenulatum:0.031885176485860194,(Mimon_bennettii:0.06286408701090246,Mimon_cozumelae:0.03767088071008071)0.91:0.05768438703934938)0.52:0.024129152116765862,(Mimon_crenulatum_longifolium:0.04429458708624136,(Mimon_crenulatum_keenani:0.024679127200726972,(Noctilio_albiventris:0.05223147792294931,Noctilio_leporinus:0.01705405128137011)1.00:0.3696694803770624)0.3:0.020535769040843307)0.4:0.027100106417385365)0.20:0.03693194588994489)0.8:0.037188548088746595)0.0:0.025347709171721185,(((Trachops_cirrhosus:0.03713201672047136,Trachops_cirrhosus_coffini:0.052549326238605705)0.94:0.07739923326066568,(Chrotopterus_auritus:0.09315487183036636,(Vampyrum_spectrum:0.08658568279468082,(Notonycteris_magdalenensis:0.03087344232657442,Notonycteris_sucharadeus:0.004647447962178754)0.44:0.02374358434456695)0.45:0.05395707884366255)0.47:0.09486927109456886)0.31:0.064091414923952,(((Tonatia_saurophila_saurophila:8.3333375002424E-7,(Tonatia_saurophila:0.04692662993489393,Tonatia_saurophila_bakeri:0.03305516916810309)0.70:0.0265915165783152)0.71:0.02857450851044572,((Lophostoma_evotis:0.0596732576437547,(Lophostoma_carrikeri:0.029108471213821218,(Lophostoma_kalkoae:0.104324401040163,(Lophostoma_brasiliense:0.05078021049463138,Lophostoma_brasiliense_minuta:0.049191234966420486)0.47:0.010342081701107028)0.18:0.01942072874227378)0.15:0.014782320028757933)0.14:0.013011458609703621,(Lophostoma_silvicolum:0.05600181022396788,(Lophostoma_schulzi:0.04061905307010716,(Lophostoma_silvicolum_silvicolum:0.013808828130393382,(Lophostoma_occidentalis:0.01902325106083884,Lophostoma_silvicolum_centralis:0.01781828482469542)0.74:0.02117798669873175)0.89:0.035227349502156294)0.41:0.01666370766907988)0.11:0.0058022296225213755)0.59:0.032332122231040386)0.28:0.018583836008432872,((Micronycteris_minuta:0.04158714496559577,((Micronycteris_microtis:0.07119494111129057,Micronycteris_microtis_mexicana:0.01162499082429022)0.39:0.02088695463919241,((Micronycteris_matses:0.0730764985906352,Micronycteris_schmidtorum:0.06239335325431199)0.8:0.01397862469081627,(Micronycteris_megalotis:0.05835111072265749,((Micronycteris_brosseti:0.05596046400739946,Micronycteris_hirsuta:0.05179465826254238)0.31:0.01766892817160335,(Micronycteris_homezi:0.0370792886403234,(Neonycteris_pusilla:0.12043607922408112,(Trinycteris_nicefori:0.12782983768920075,(Glyphonycteris_daviesi:0.17497961567073425,Glyphonycteris_sylvestris:0.08319637122590667)0.97:0.10311081444849218)0.45:0.03646175120793642)0.35:0.058857815524081473)0.14:0.015877911329388348)0.7:0.017986311949778906)0.10:0.016341180813216984)0.4:0.0035495798489613)0.19:0.021495272834504777)0.48:0.03737825487931003,(Lampronycteris_brachyotis:0.10992091165040083,((Macrotus_californicus:0.03298350941080783,Macrotus_waterhousii:0.09648322795723321)1.00:0.060052114380095944,((((Carollia_brevicauda:0.05613909648864387,Carollia_perspicillata:0.08763216911790722)0.94:0.06793980995953898,(Rhinophylla_pumilio:0.032043091426895216,(Rhinophylla_alethina:0.02060686245715419,Rhinophylla_fischerae:8.3333375002424E-7)0.95:0.04331919362064558)0.95:0.21764267682026026)0.48:0.10334659638013355,(((Erophylla_bombifrons:0.03126591495784339,Erophylla_sezekorni:8.3333375002424E-7)0.86:0.06746450686831611,(Phyllonycteris_major:0.01352036502754473,(Phyllonycteris_aphylla:0.02193806572758536,Phyllonycteris_poeyi:0.0758325065092939)0.79:0.010975982192707593)0.99:0.152776288876354)0.66:0.08567165068237846,((Pygoderma_bilabiatum:0.23144129998114724,(Brachyphylla_cavernarum:0.0625353659617773,Brachyphylla_nana:0.02376071852032125)1.00:0.19736879001930718)0.24:0.047930012971938184,(Sturnira_lilium:0.17387535405472646,(((Platyrrhinus_helleri:0.12664876987801482,Uroderma_bilobatum:0.1690441583869879)0.55:0.037241998478948914,(Vampyrodes_caraccioli:0.034805031645331136,Vampyrodes_major:0.05583676162402534)0.95:0.09724019939505019)0.51:0.07044636964793975,(Enchisthenes_hartii:0.07633507788668646,(Artibeus_gnomus:0.07185254908778567,(Artibeus_obscurus:0.04697470910517307,(Artibeus_jamaicensis:0.08617784581433187,Artibeus_lituratus:0.038410098301970036)0.53:0.030718858357641342)0.63:0.033574843582934034)0.82:0.08912951396325498)0.22:0.014152895313404343)0.23:0.06447262388085803)0.30:0.06759869300665133)0.83:0.08187310298943541)0.36:0.09687186638627139)0.22:0.009996105929922904,((Glossophaga_soricina:0.032089135588981986,(Glossophaga_morenoi:0.041337791292420725,(Glossophaga_longirostris:0.028257287681187907,(Glossophaga_commissarisi:0.073430334450104,Glossophaga_leachii:0.017401403547261093)0.59:0.025056270919890537)0.77:0.05920638258972578)0.60:0.027060852385151105)0.67:0.03895459969888022,((Monophyllus_plethodon:0.031039412505188452,Monophyllus_redmani:0.05064529439466355)0.83:0.08151321964965029,((Lonchophylla_thomasi:0.03837708503582683,(Lonchophylla_mordax:0.09409205701550914,(Lonchophylla_robusta:0.06943745521099445,(Lonchophylla_handleyi:0.025400598402318543,(Lionycteris_spurrelli:8.3333375002424E-7,Palynephyllum_antimaster_2:0.17339360052111238)0.25:0.1442256633301256)0.13:0.027470125803018004)0.24:0.0516722594673425)0.14:0.00862443706445891)0.26:0.06423057864757749,((Leptonycteris_nivalis:0.05316533008006431,(Leptonycteris_curasoae:0.084744146997587,Leptonycteris_yerbabuenae:0.12520901602983972)0.34:0.020038029235063166)0.88:0.06019958867505376,((Anoura_caudifer:0.06632126627000264,(Anoura_cultrata:0.03552160166854759,(Anoura_geoffroyi:0.023400288636459786,(Anoura_geoffroyi_peruana:0.04256130933988702,Anoura_latidens:0.023800206384751988)0.60:0.027795290326220766)0.87:0.061694807207141586)0.35:0.024623876987314417)0.87:0.0793748322188093,(Hylonycteris_underwoodi:0.09275867888679244,(Lichonycteris_obscura:0.16130163335424783,((Choeronycteris_mexicana:0.12443469076518858,(Choeroniscus_godmani:0.058609403956628885,(Choeroniscus_minor:0.051130192511521934,Choeroniscus_periosus:0.04782506854398366)0.78:0.034961088579000175)0.44:0.028958357111866458)0.75:0.0361822989413476,(Musonycteris_harrisoni:0.06628167464331493,(Xeronycteris_vieirai:0.11310705757690603,(Platalina_genovensium:0.04703662515740182,(Diphylla_ecaudata:0.010280198524009103,(Desmodus_rotundus:0.09868443796853658,Diaemus_youngi:0.038481318804575435)0.92:0.05022074964408515)1.00:0.33877711913629904)0.35:0.07014201713513407)0.31:0.06334115271818175)0.31:0.06094069936970891)0.25:0.0549756186580058)0.17:0.02362466108245458)0.27:0.05540755489276695)0.15:0.03618000399517157)0.8:0.01723763728032458)0.4:0.032037837475938795)0.19:0.06409339277508969)0.36:0.14725913441257002)0.91:0.1849656555937533)0.24:0.0703361361802053)0.5:0.02202046921268951)0.13:0.0504136761625699)0.3:0.024712322436632562)0.3:0.00790133031323199)0.6:0.03713916807532705)0.31:0.11822269015737684)0.18:0.05441388731944402)0.41:0.09948273153101989):0.13573457579097195); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr75067] END;