#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on September 19, 2025; 18:34 GMT

TreeBASE (cc) 1994-2008

Study reference:
Thayer M.K. 1997. Proglypholoma aenigma gen. et sp. nov., Glypholoma spp. nov. and
new records, and a phylogenetic analysis of Glypholomatinae (Coleoptera: Staphylinidae).
Annales Zoologici, 47(1/2): 157-174.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S731]

[ The following blocks are input data for analysis step 1133 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep1133;
        DIMENSIONS NTAX=10;
        TAXLABELS
            Glypholoma_chepuense
            Glypholoma_germaini
            Glypholoma_pecki
            Glypholoma_pustuliferum
            Glypholoma_rotundulum
            Glypholoma_temporale
            Glypholoma_tenuicorne
            Microsilpha_spp
            Nitidotachinus
            Proglypholoma_aenigma
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2303]

	TITLE Table_2;
	LINK TAXA = TaxaForAnalysisStep1133;
	DIMENSIONS NCHAR=40;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7" MISSING=? GAP= -;
	CHARSTATELABELS
		1  'body outline, dorsal view' / elongate oblong oval,
		2  'dorsal contour (transverse section)' / mod._convex sl._convex very_convex,
		3  epistomal_suture_stem / present absent,
		4  anteoc._pits_or_grooves / 'broad- shallow' 'narrow-elongate' absent,
		5  antennal_form / filiform clavate,
		6  antenna_#_club_segments / 0 3 4 5 6,
		7  'antenna - # glabrous basal segments' / 2 4 5 6 7,
		8  pronotum_widest / near_back behind_middle middle before_middle,
		9  pronotal_lateral_margin_explanate / basally entire_length,
		10  #_elytral_striae / 0 11 '<11',
		11  elytral_striae / impressed_between_punctures not_impressed_between_punctures absent,
		12  elytral_apex_color_compared_to_disk / concolorous abruptly_paler,
		13  elytral_pustules / absent present,
		14  elytral_pale_spots / absent present,
		15  elytral_intervals_with_fine_punctures / absent present 'striae absent-no interv',
		16  elytral_epipl_keel_length_vs._elytral_length / _0.6 ' 0.64-0.69' _0.8 _1.0,
		17  elytra_cover_abdomen_through_tergite / 3 4 5 6,
		18  prosternal_carina / absent present,
		19  prosternal_surface / punctate rugose smooth,
		20  mesosternal_punctation / present absent,
		21  metasternal_antecoxal_suture / short long absent,
		22  procoxal_mesal_groove / absent present,
		23  metacoxal_posterior_face / oblique vertical slightly_excavate distinctly_excavate,
		24  metafemoral_length_vs._metacoxal_length / much_greater_than greater_than less_than,
		25  tibial_external_spines / present absent,
		26  metatarsal_length_vs._metatibial_length / 0.56 'sl < 0.6' 0.6 0.61 0.67 0.7 0.71 0.76,
		27  'metatarsus, length of tarsomere 5 vs 1-4 together' / '>= 0.5' '=< 0.4',
		28  length_of_empodial_setae_vs._length_of_claws / greater_than equal_to less_than,
		29  'first well-sclerotized abdominal tergite' / 3 4 5 6,
		30  abdominal_spiracles_in_tergites_on_segments / '3-8' '4-8' '5-8',
		31  sternite_8_projections_lateral_to_median_defensive_gland_projection / absent narrow transversely_elongate,
		32  male_tergite_9_mediodorsal_connection / absent narrow wide,
		33  male_tergite_9_shortest_dorsal_length_vs._lateral_length / '0 (tergite 9 divided)' '0.03-0.08' 0.1 0.15 0.27,
		34  'male sternite internal anterior "button"' / absent present,
		35  anterior_part_of_male_stenite_9 / not_between_anterior_projections forming_wide__connection_between_projections forming_narrow_connection_between_projections,
		36  aedeagus_with_bellows_action / 'dorsal-ventral' 'lateral-lateral' 'anterior-posterior',
		37  parameres_with_subapical_setae / present absent,
		38  aedeagus_with_apex_of_median_lobe / not_digitiform digitiform,
		39  aedeagal_basal_piece / absent narrow wide,
		40  teeth_on_aedeagal_internal_sac / 'even-sized' 'mixed-sized' absent,
		;

MATRIX
[                                   10        20        30        40]
[                                    .         .         .         .]
Glypholoma_chepuense        1002142111001000200001310202322?????????
Glypholoma_germaini         1011132111001011?010?12206011??111200120
Glypholoma_pecki            1012114211010011310101320401221111201110
Glypholoma_pustuliferum     1001133211011111201001320201112111201010
Glypholoma_rotundulum       2202114001010111310101310100121121200020
Glypholoma_temporale        1012121111011011201111320500221131200110
Glypholoma_tenuicorne       011200100111000020010132151111?111200110
Microsilpha_spp             2202124000200021200010110702211000011022
Nitidotachinus              0001000000100013000020000310000000021011
Proglypholoma_aenigma       0100001302101002100000201002201240110101
;
END;

[ The following blocks are output data for analysis step 1133 ]
BEGIN TREES;
      TITLE Tb5909;
      LINK TAXA = TaxaForAnalysisStep1133;
      TREE Fig._32 = [&R] (Nitidotachinus,(Microsilpha_spp,((((((Glypholoma_pustuliferum,Glypholoma_germaini),Glypholoma_chepuense),Glypholoma_temporale),(Glypholoma_pecki,Glypholoma_rotundulum)),Glypholoma_tenuicorne)Glypholoma,Proglypholoma_aenigma)));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr3802]


END;
BEGIN TREES;
      TITLE Tb5909;
      LINK TAXA = TaxaForAnalysisStep1133;
      TREE Fig._32_inset = [&R] (Nitidotachinus,(Microsilpha_spp,(((((Glypholoma_pustuliferum,(Glypholoma_chepuense,Glypholoma_germaini)),Glypholoma_temporale),(Glypholoma_pecki,Glypholoma_rotundulum)),Glypholoma_tenuicorne)Glypholoma,Proglypholoma_aenigma)Glypholomatinae));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr3801]


END;