#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on November 08, 2025; 13:53 GMT TreeBASE (cc) 1994-2008 Study reference: Whitmore D., Pape T., & Cerretti P. 2013. Phylogeny of Heteronychia: the largest lineage of Sarcophaga (Diptera: Sarcophagidae). Zoological Journal of the Linnean Society, 169(3): 604-639. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S14253] [ The following blocks are input data for analysis step 11465 ] BEGIN TAXA; TITLE TaxaForAnalysisStep11465; DIMENSIONS NTAX=88; TAXLABELS Sarcophaga_abramovi Sarcophaga_agnata Sarcophaga_amita Sarcophaga_amputata Sarcophaga_anatolica Sarcophaga_ancilla Sarcophaga_arcipes Sarcophaga_armeniaca Sarcophaga_balanina Sarcophaga_belanovskyi Sarcophaga_benaci Sarcophaga_bezziana Sarcophaga_boettcheri Sarcophaga_bulgarica Sarcophaga_chaetoneura Sarcophaga_chiquita Sarcophaga_clarahenae Sarcophaga_consanguinea Sarcophaga_croca Sarcophaga_cucullans Sarcophaga_curvifemoralis Sarcophaga_depressifrons Sarcophaga_desertorum Sarcophaga_dissimilis Sarcophaga_enderleini Sarcophaga_esthera Sarcophaga_ferox Sarcophaga_filia Sarcophaga_filiola Sarcophaga_gabrielei Sarcophaga_gallica Sarcophaga_giganta Sarcophaga_haemorrhoa Sarcophaga_haemorrhoides Sarcophaga_helenae Sarcophaga_hellenica Sarcophaga_heptapotamica Sarcophaga_infantilis Sarcophaga_infixa Sarcophaga_javita Sarcophaga_kataphygionis Sarcophaga_kerteszi Sarcophaga_kozlovi Sarcophaga_kunonis Sarcophaga_lacrymans Sarcophaga_lederbergi Sarcophaga_lejlekensis Sarcophaga_longestylata Sarcophaga_mediterranea Sarcophaga_metopina Sarcophaga_minima Sarcophaga_monspellensia Sarcophaga_mutila Sarcophaga_nanula Sarcophaga_nipponensis Sarcophaga_obvia Sarcophaga_pandellei Sarcophaga_pauciseta Sarcophaga_penicillata Sarcophaga_plotnikovi Sarcophaga_pontica Sarcophaga_porrecta Sarcophaga_proxima Sarcophaga_pseudobenaci Sarcophaga_pumila Sarcophaga_recta Sarcophaga_rondaniana Sarcophaga_rosellensis Sarcophaga_santospintosi Sarcophaga_schineri Sarcophaga_setinervis Sarcophaga_shnitnikovi Sarcophaga_sicilia Sarcophaga_siciliana Sarcophaga_siciliensis Sarcophaga_slovaca Sarcophaga_tangerensis Sarcophaga_tetrix Sarcophaga_thirionae Sarcophaga_tricolor Sarcophaga_tunisiae Sarcophaga_turana Sarcophaga_uncicurva Sarcophaga_vagans Sarcophaga_variegata Sarcophaga_vicina Sarcophaga_villeneuveana Sarcophaga_violovitshi ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M17099] TITLE Genitalia_matrix; LINK TAXA = TaxaForAnalysisStep11465; DIMENSIONS NCHAR=70; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7 8 9 A B C D" MISSING=? GAP= -; CHARSTATELABELS 1 'Sternite 5, setae', 2 'Syntergosternite 7+8, microtrichosity', 3 'Epandrium, ventral elongation', 4 'Cercus, dorsal excavation', 5 'Cercus, median keel', 6 'Cercus, subapical hump', 7 'Cercus, dorsal saddle', 8 'Distal part of cercus, direction', 9 'Distal part of cercus, direction', 10 'Cercus tip, shape', 11 'Cercus, subapical ventral indentation', 12 'Cercus, subapical lateral keel', 13 'Cercus, distribution of setae', 14 'Cercus, setulae distal ventral surface', 15 'Cercus, microtrichia', 16 'Surstylus, microtrichia', 17 'Pregonite, orientation compared to postgonite', 18 'Pregonite, preapical twist', 19 'Pregonite tip, orientation', 20 'Pregonite tip, shape', 21 'Pregonite, type and distribution of setae', 22 'Pregonite, middle section', 23 'Pregonite, sclerotization', 24 'Postgonite, tip', 25 'Phallic tube, basal section', 26 'Paraphallus, basal part', 27 Membrane, 28 'Vesica, number of lobes', 29 'Vesica, shape', 30 'Vesica, microtrichia', 31 'Harpes, structure', 32 'Harpes, proximal part (lateral view)', 33 'Harpes, proximal part - sclerotization', 34 'Harpes, proximal part - inner surface', 35 'Harpes, apical process', 36 'Harpes, apical process - position', 37 'Harpes, apical process - shape', 38 'Harpes, apical process - direction', 39 'Harpes, apical process - median expansion', 40 'Harpes, apical process - denticles', 41 'Harpes, microtrichia', 42 'Lateral styli, volume', 43 'Lateral styli, spines - number and arrangement', 44 'Lateral styli, spines - type', 45 'Lateral styli, length', 46 'Lateral styli, position with respect to juxta', 47 'Lateral styli, tip', 48 'Juxta, general aspect', 49 'Juxta, length', 50 'Juxta, orientation of main axis', 51 'Juxta, dorsal surface lateral view', 52 'Juxta, structure', 53 'Juxta, median process - preapical part', 54 'Juxta, median process - sclerotization', 55 'Juxta, median process - median suture', 56 'Juxta, median process - median keel', 57 'Juxta, median process - general shape and orientation', 58 'Juxta, median process - lateral margins', 59 'Juxta, median process - direction of tip', 60 'Juxta, median process - shape of tip in lateral view', 61 'Juxta, median process - preapical lateral indentation', 62 'Juxta, median process - apical tooth-like protuberances', 63 'Juxta, median process - median depression on tip', 64 'Juxta, median process - preapical widening', 65 'Juxta, median process - wide constriction near base', 66 'Juxta, median process - longitudinal furrows', 67 'Juxta, median process - tip in dorsal/apical view', 68 'Juxta, lateral process', 69 'Juxta, basal process', 70 'Juxta, connective part', ; MATRIX [ 10 20 30 40 50 60 70 80 ] [ . . . . . . . . ] Sarcophaga_abramovi 1?10000102000011101110002011302200200000012111211201110000200100001?11 Sarcophaga_agnata 24102000210000101?2351011051101201??????001311310020011000220200000??? Sarcophaga_amita 2210010011001010001100001011402200300500012111211211110012300010001?21 Sarcophaga_amputata 2?11121110100010111100000011203001305600212?20012311010031400000107?40 Sarcophaga_anatolica 1010010011002010111110002011402200200000012111210221010000200000001?02 Sarcophaga_ancilla 1010010012000{01}10111110002011402200200000012111210211010010200000001?01 Sarcophaga_arcipes 2211000102000010001100002211413001305600212110212311010021430000005?30 Sarcophaga_armeniaca 2?10010012000010011110002011402200200000012111{23}10221010000200000001?01 Sarcophaga_balanina 2111020012002010112200000011A04101311400012120012210021033510000004??? Sarcophaga_belanovskyi 1110010012000010011110002011402200200000012111210211010000200000001?01 Sarcophaga_benaci 22100101120020101111100020215022012000000121112101220100001000000030?2 Sarcophaga_bezziana 2210010101001010001110001011602200300500012111311211110012300010001?21 Sarcophaga_boettcheri 1210110112003010101421002141805310322500012110210210040013200000000??? Sarcophaga_bulgarica 2210010102000010011110002011602200200200012111210201010000200010001?21 Sarcophaga_chaetoneura 10101001120{01}00101121310020214022003000000121112102220100000000000032?{234} Sarcophaga_chiquita 2010010102002010000110002011402200200000012111210221010000200000001?03 Sarcophaga_clarahenae 1010010011000010001110002011402200200000012111310221010000200000001?03 Sarcophaga_consanguinea 22100100120100100011100020214022003003000121102102220100002110000031?4 Sarcophaga_croca 1210010102000010112101002011402200200000012111310211010000200000001?03 Sarcophaga_cucullans 2111120011000010001100002211313201100600212120012311010021430000005?30 Sarcophaga_curvifemoralis 2110010010002010111341002011402200340700012110110211010000000000000?11 Sarcophaga_depressifrons {12}210010002000010001110002011602200200200012111210221010000200000001?21 Sarcophaga_desertorum ??11?2001100????1121?0002011A?420?2??????12?200122100???3??000??002??? Sarcophaga_dissimilis 101010011200001011213100202140220030000001211{01}2102220100002000000032?{234} Sarcophaga_enderleini 21101001120020100023300020217022003003000121102102120100000100000031?4 Sarcophaga_esthera 2111120011000010001100002211313201100600212120012311010021430000005?30 Sarcophaga_ferox 22111201{12}2000010002200002011A04101311400012120012110020133400000002??? Sarcophaga_filia 2110010111003010101421102141805310322800012100211220030003400000000??? Sarcophaga_filiola ??1001002000?0????142??0214????31????????12?002112100??010200?0?000??? Sarcophaga_gabrielei 21101100210020101010310020117022003000002121100101121100002000000031?1 Sarcophaga_gallica 2210010002010010112110002011402200200000012111311221110000201000001?10 Sarcophaga_giganta 21100000220020101023310030214022002003000121100101121100403000000011?4 Sarcophaga_haemorrhoa 2210010{01}0{23}000010001110002021602200200000012111211211111010310010008?20 Sarcophaga_haemorrhoides 2210010{01}02000010001110002011602200200000012111{23}11{12}{02}11100{01}0301110001?10 Sarcophaga_helenae 2210010{01}02000010001110001021602200100000{01}12111211211110010310000000?21 Sarcophaga_hellenica 22101000201000100112200020214022003002001121102101220100001000000092?2 Sarcophaga_heptapotamica 221001001200001000{01}110002011602200200000012111{23}10201010000200000001?0{23} Sarcophaga_infantilis 12100100110000100011100020115022003000000121112102020100002000000030?2 Sarcophaga_infixa 22100100{01}2010010112110002011402201200000012111311201110000200020001?10 Sarcophaga_javita 22100100{01}20001100021200020112022002003000121110102120100103100000191?{12} Sarcophaga_kataphygionis 22100100020000100011100020116022002000000121112112{02}1{01}10000200010001?21 Sarcophaga_kerteszi 2211120102001010001330002211B022013004003121100101121100003100000013?1 Sarcophaga_kozlovi 2?10?000200?20???02330002021??220030030001211??101120100002000000031?? Sarcophaga_kunonis 20100200001020101102010100112032010?????2121200123110100?1430000100?30 Sarcophaga_lacrymans 10100100120100101005010020211022002003000121112102120100002000000030?1 Sarcophaga_lederbergi 22100100{01}2000010001110001011602200200000012111310201010000201000001?23 Sarcophaga_lejlekensis 2210010020100110002330002021702200300200012110?102020100002100000031?4 Sarcophaga_longestylata 1010000011003010101421101141D053112220000121102102100?0003200000000??? Sarcophaga_mediterranea 2110031120000011101400101011206201233100012220012213050014310000008??? Sarcophaga_metopina 1010010012000010001110001011402200200000012111210211010000200000001??5 Sarcophaga_minima 2011000{01}12000010001410101011206101233101012212012213010013200000004??? Sarcophaga_monspellensia 201013112000?010112201001011107201236500012111011211010000200000004?10 Sarcophaga_mutila 22111001110000100011210010214082010?????012110010311110010301000004?21 Sarcophaga_nanula 12100100020000100011100020116022002002000121112102{02}1010000200000001?21 Sarcophaga_nipponensis 2010100020010{01}100011000020217022002003000121112102220100002100000031?4 Sarcophaga_obvia 2210010002010110112100002021C022002003000121112102120100000100010092?4 Sarcophaga_pandellei 2210010002000110112100002021C022002003000121112102120100001100010092?4 Sarcophaga_pauciseta 21102100020120100020100020214022002003000121110102120100002010000031?4 Sarcophaga_penicillata 2110031120000011101401101011206101230500012210012213050034510000006??? Sarcophaga_plotnikovi 2210010002002010011110002021602200200300012111211211011010201010001?10 Sarcophaga_pontica 1210010011000010001110002011402200200000012111210231010000200000001?01 Sarcophaga_porrecta 20101001{01}20020100023300020217022003003000121100102020100002100000031?4 Sarcophaga_proxima 22102100120022100002000020212022002003000121112102120100002100000031?4 Sarcophaga_pseudobenaci 1210110021000{01}100011200020215022013000000121112102220100002000000030?2 Sarcophaga_pumila 211110?012000010001100001211613001305600212120012311010021430000005?30 Sarcophaga_recta 22102101120000101000010020212022002003000121102102120100102000000131?4 Sarcophaga_rondaniana {12}{01}100000120100100005000020211022002003000121112102121100002000000030?2 Sarcophaga_rosellensis 22102100120120100010000020212022002003000121112102220100002100000031?4 Sarcophaga_santospintosi 2110011112000010101401100011306201230110012210012113050014210000006??? Sarcophaga_schineri 11100000220001100010300020216022003003000121112102220100002000000031?4 Sarcophaga_setinervis 2211110{01}1{12}000010{01}0112{01}0010214082010?????012110010311110010301000004?21 Sarcophaga_shnitnikovi 21100000110020100023300020217022002003000121100102120100102100000031?4 Sarcophaga_sicilia 1010000111003010101421102141805310322500012100210220030003{23}00000000??? Sarcophaga_siciliana 22101000001000100020200020214022003002000121110102020100002000000092?2 Sarcophaga_siciliensis 2311110020100021101401101011206101233101012212211213010013300000001??? Sarcophaga_slovaca 1210010002000010001110002011602200200200012111310221010000200000001?21 Sarcophaga_tangerensis 2?100100120001101020000020112022002003000121110101220100002010010031?4 Sarcophaga_tetrix 2110031100000011001410101011906201234110012210111213050050500000006??? Sarcophaga_thirionae 2110031120000011101400101011206201233100012120012213050030510000004??? Sarcophaga_tricolor 2010010120000010101400101011206201230100012110211213050014210000000??? Sarcophaga_tunisiae 21100100210020100022200020214022003004000121111102120100002000000090?2 Sarcophaga_turana 1{13}10010111003010101421102141805310322800012100210220030003301000000??? Sarcophaga_uncicurva 2{01}10031120000011101401101011206201233110012110011113050054??011000A??? Sarcophaga_vagans 21102000110120100021100020314022002003000121112102020100002100000031?4 Sarcophaga_variegata 0400000020100000000150012300000?????????000011300020000100220000000??? Sarcophaga_vicina 2110100{01}220020100023300030217022003003000121100102120100102000000031?4 Sarcophaga_villeneuveana 2{01}10031120000011001400{01}01011906201234110012?10112113050054510000006??? Sarcophaga_violovitshi 1210010002000010112310002011402200200000012111211201110000201110001?10 ; END; [ The following blocks are output data for analysis step 11465 ] BEGIN TREES; TITLE Consensus_genitalia_matrix; LINK TAXA = TaxaForAnalysisStep11465; TREE Imported_tree_1 = [&R] (Sarcophaga_variegata,(Sarcophaga_agnata,(((Sarcophaga_filiola,(Sarcophaga_longestylata,(Sarcophaga_boettcheri,(Sarcophaga_sicilia,(Sarcophaga_filia,Sarcophaga_turana))))),((Sarcophaga_tricolor,(Sarcophaga_santospintosi,(Sarcophaga_penicillata,((Sarcophaga_mediterranea,Sarcophaga_thirionae),(Sarcophaga_uncicurva,(Sarcophaga_tetrix,Sarcophaga_villeneuveana)))))),(Sarcophaga_minima,Sarcophaga_siciliensis))),(Sarcophaga_monspellensia,((Sarcophaga_croca,(Sarcophaga_anatolica,Sarcophaga_ancilla,Sarcophaga_armeniaca,Sarcophaga_belanovskyi,Sarcophaga_chiquita,Sarcophaga_clarahenae,Sarcophaga_metopina,Sarcophaga_pontica,(Sarcophaga_bulgarica,Sarcophaga_heptapotamica,Sarcophaga_lederbergi,(Sarcophaga_abramovi,Sarcophaga_haemorrhoa,Sarcophaga_haemorrhoides,Sarcophaga_helenae,Sarcophaga_kataphygionis,Sarcophaga_plotnikovi,(Sarcophaga_amita,Sarcophaga_bezziana),(Sarcophaga_violovitshi,(Sarcophaga_gallica,Sarcophaga_infixa))),(Sarcophaga_depressifrons,Sarcophaga_nanula,Sarcophaga_slovaca)),(Sarcophaga_infantilis,(Sarcophaga_benaci,(Sarcophaga_chaetoneura,Sarcophaga_dissimilis)),(Sarcophaga_consanguinea,Sarcophaga_javita,Sarcophaga_nipponensis,Sarcophaga_tangerensis,Sarcophaga_proxima,Sarcophaga_recta,Sarcophaga_rosellensis,Sarcophaga_schineri,(Sarcophaga_lejlekensis,(Sarcophaga_shnitnikovi,(Sarcophaga_enderleini,Sarcophaga_porrecta),(Sarcophaga_vicina,(Sarcophaga_kozlovi,(Sarcophaga_gabrielei,Sarcophaga_giganta))))),(Sarcophaga_lacrymans,Sarcophaga_rondaniana),(Sarcophaga_obvia,Sarcophaga_pandellei),(Sarcophaga_pauciseta,Sarcophaga_vagans)),(Sarcophaga_pseudobenaci,(Sarcophaga_tunisiae,(Sarcophaga_hellenica,Sarcophaga_siciliana)))))),(Sarcophaga_curvifemoralis,((((Sarcophaga_amputata,Sarcophaga_kunonis),((Sarcophaga_arcipes,Sarcophaga_pumila),(Sarcophaga_cucullans,Sarcophaga_esthera))),(Sarcophaga_desertorum,(Sarcophaga_balanina,Sarcophaga_ferox))),(Sarcophaga_kerteszi,(Sarcophaga_mutila,Sarcophaga_setinervis))))))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr63602] END;