#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on November 15, 2026; 5:25 GMT

TreeBASE (cc) 1994-2008

Study reference:
Krueger D., Binder M., Fischer M., & Kreisel H. 2001. The Lycoperdales. A molecular
approach to the systematics of some gasteroid mushrooms. Mycologia, 93(5): 947-957.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S750]

[ The following blocks are input data for analysis step 1167 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep1167;
        DIMENSIONS NTAX=12;
        TAXLABELS
            Bovista_paludosa
            Bovista_plumbea
            Bovista_polymorpha
            Bovista_pusilla
            Bovistella_radicata
            Lycoperdon_echinatum
            Lycoperdon_foetidum
            Lycoperdon_mammiforme
            Lycoperdon_perlatum
            Lycoperdon_pyriforme_Bavaria
            Morganella_subincarnata
            Vascellum_pratense
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2345]

	TITLE ITS_no_Lepiota;
	LINK TAXA = TaxaForAnalysisStep1167;
	DIMENSIONS NCHAR=703;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                        10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510       520       530       540       550       560       570       580       590       600       610       620       630       640       650       660       670       680       690       700   ]
[                                         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .   ]
Bovista_paludosa                 TTAGGA--CTTATTGAATATTCTTGATGGGTTGTAGCTGGCTCTTTGGAGCATGTGCACGCCTGTCTTGACTTTATTC-ATCCACCTGTGCACCTTTTGTAGTCTTGGAGGTTGAGAGCGG-TCAAC--TATCGGATGGTTTCCGGATGTGAGGAATTGCTGAGTGC-AAAA-GCATACAGCTCTTCTCAAAACGACTTGCCTCTCCTTCAAGTACTATGTATT---CATATACCACATAGTATGTTGTAGAATGT-GATCAATGGGCTTATGTGCCTATAATAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCTAGCTTTTGCGAGCTGTGATGGGGCTTGGATGTT--GGGTTTGCG-GGCC-TTTATTAA--TAGAGGTCGGCTCTCCTGAAATGCATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTGTCGGACAATATTTAATGAACTT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Bovista_plumbea                  TTAGGA--CTTATTGAATATTCTTGATGGGTTGTAGCTGGCTCTCCGGAGCATGTGCACGCCTGTCTTGACTTAATTC-ATCCACCTGTGCACTTTTTGTAGTCTTGGAGGTTGAGAGCGG-TCGAC--TATCGGATGGTTTCCGGATGTGAGGAATTGCTGAGTGC-AAAA-GCATACAGCTCTTCTCAAAATGACTCGCCTCTCCTTCAAGTACTATGTATT---CATATACCACATAGTATGTTGTAGAATGT-GATCAATGGGCTTATG-GCCTATAATAATGAAATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCTAGCTTTTGCGAGCTACGATGGGGCTTGGATGTT--GGGTTTGCG-GGCC-TTTATTAA--TAGAGGTCGGCTCTCCTGAAATGCATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTATCGGACAATACTTAATGAACTT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Bovista_polymorpha               TTAGGA--CTTATTGAATATTCTTGATGGGTTGTAGCTGGCTCTTCGGGGCATGTGCACGCCTGTCTTGACTTTATTC-GTCCACCTGTGCACCTTTTGTAGTCTTGTGGGTTGAGAGCAG-TCAAC--TATCGGACAGGCTCCGGATGTGAGGAAT-GCTGAGTGC-GAAA-GCATACAGCTCTTCTCAA--CGA-TTGCCTCTCCCTCGAGTACTATGTATT---CATATACCAC-TTGTATGTTGTCGAATGT-GATCAATGGGCTTATATGCCTATAATAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCC-TCCAGCTTTTGCGAGTTGTGATGGG-CTTGGATAT-GGGGGTTTGCG-GGCC-TTTATTAA----GAGGTCGGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATAGTCCCTTGTTGGACAACATT-AATGAAATT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Bovista_pusilla                  TTAGGA--CTTATTGAATACTCTTGATGGGTTGTAGCTGGCTCCTTGGGGCATGTGCACGCCTGTCTTGACTTTATTC-ATCCACCTGTGCACCTTTTGTAGTCTTGGGGGTTGAGAGCAG-TCAAC--TGTCGGATGGGCTCCGGATGTGAGGGAT-GCTGAGTGC-GAAA-GCATACAGCTCTTCTCAAT-CGACTTGCCTCTCCCTCGAGTACTATGTATT---CATATACCAC-TAGTATGTTGTAGAATGT-GATCAATGGGCTTTTATGCCTATAATAATCA-ATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCCAGCTTTTGCGAGCTGCGATGGGGCTTGGATGTTGGGGGTTTGCG-GGCC-TTTATTAA----GAGGTCGGCTCTCCTGAAATGCATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCATTC--GGACAATGTTTAATGAACTT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Bovistella_radicata              TTAGGA--CTTATTGAATACTCTTGATGGGTTGTAGCTGGCTCTTCGGGGCATGTGCACACTTGTCTTGACTTTATTC-ATCCACCTGTGCACCTTTTGTAGTCTTGTGGGTTGAGAGCAG-TCGAC--TATCGGATGGTTTCCGGACGTGAGGAAT-GCTGAGTGC-GAAA-GCATACAGCTCTTCTCAAA-TGACTTGCCTCTTCCTCAAGTACTATGTATT---CATATACCACATAGTATGTTGTCGAATGT-GATCAATGGGCCTATGTGCCTATAAAAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCCAGCTTTTGCGAGTTGTGATGGGGCTTGGATAT-GGGAGTTTGCG-GGTC-TTTATCAA--TAAAGGTCAGCTCTCCTGAAATGCATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTAT-GGACAATACTTAATGAACAT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Lycoperdon_echinatum             TTAGGATCATTATTGAATATTCTTGATGGGTTGTAGCTGGCTCTCCGGGGCATGTGCACACTTGTCTTGACTTTATTC-GTCCACCTGTGCACCTTTTGTAGTCTTGGGGGTTGAGAACAAATCGAC--TATCGGATGGTTTCCGGACGTGAGGATT-GCTGAGTGT-GAAA-GCATACAGCTCTTCTCAAA-TGACTTG-CTCTCCCTCGAGTACTATGTATT---CATATACCACATAGTATGTTGTAGAATGT-GATCAATGGGCCTACGTGCCTATAATAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCTTCCAGCTTTTGCGAGTTGTGATGGG-CTTGGATTG-GGGAGTTTGCG-GGCC-TTTATCAA--TAAAGGTCAGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTAT-GGACAACACTTAATGAACTT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Lycoperdon_foetidum              TTAGGATCATTATTGAATATTCTTGATGGGTTGTAGCTGGCTCTTAGGGGCATGTGCACACTTGTCTTGACTTTATTC-ATCCACCTGTGCACCCTTTGTAGTCTTGGGGGTTGAGAGCAG-TCGAC--TATCGGATGGTTTCCGGACATGAGGATT-GCTGAGTGC-GCAA-GCATACAGCTCTTCTCAAA-TGGCTTGCCTCTCCCTCGAGTACTATGTATT---CATATACCACATAGTATGTTGAAGAATGT-GATCAATGGGCCTATGTGCCTATAACAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCCACCTTTTGCGAGTTGTGATGGGGCTTGGATCT-GGGAGTTTGCG-GGTC-TTTATCAA--TGAAGGTCAGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACGCTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTAT-GGACAACACTTAATGAACTT-CTTGACCTCAA-TCAGGTAGGACTACCCGA
Lycoperdon_mammiforme            TTAGGATCATTATTGAATATTCTTGATGGGTTGTAGCTGGCTCTCAGGAGCATGTGCACACTTGTCTTGACTTTATTC-GTCCACCTGTGCACCTTTTGTAGTCTTGGGGGTTACGAGCAG-TCGAC--TATCGGATGGTTTCCGGACGTGAGGATT-GCTGAGTGC-GAAA-GCATACAGCTCTTCTCTAA-TGACTT----CTCCCTCGAGTACTATGTATT---CATATACCACATAGTATGTTGTAGAATGT-GATCAATGGGCCTATGTGCCTATAATAATGAAATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAATCCCTCCAGCTTTTGCGAGTTGTGATGGGGCTTGGATCT-GGGAGTTTGCG-GGTC-TTTATCAA--TAAAGGTCAGCTCTCCTGAAATGCATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTAC-GGACAACACTTAATGAACTTACTTGACCTCAAATCAGGTAGGACTACCCGC
Lycoperdon_perlatum              TTAGGATCATTATTGAATATGCTTGATAGGTTGTAGCTGGCTCTTCGGAGTATGTGCACGCTTATCTTGACTTAATTC-ATCCACCTGTGCACCTTTTGTAGTCTTGGGGGTTGAGATCGG-TCGAC--TATCGGATGGATTCCGGATGTGAGGAAT-GCTGAGTGC-GAAA-GCATACAGCTCTTCTCAAA-TGACTTG--TCTTCCTCGAGTACTATGTATT---CATATACCACATAGTATGTTGTAGAATGT-GATCAATGGGTCTATGTACCTATAATAATTATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCCAGCTTTTGCGAGTTGTGATGGGGCTTGGATAT-GGGGGCTTGCG-GGTC-TTTACTGATGTAAAGGTCAGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA--TGGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCGTTAT-GGACAATGTTTAATGAACTT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Lycoperdon_pyriforme_Bavaria     TTAGGATCATTATTGAATATTCCTGATGAGTTGTAGCTGGCTCTTTGGAGCATGTGCACGCTTGTCTTGATCTTATTC-ATCCACCTGTGCACATTTTGTAGTCTTTTGGGCTGAG------TCGAG--TATCGGATGGTATCCGGATGTGAGGACT-GCTGAGTGC-GAAA-GCATACGGCTCTTCTCTGTTTGACCT----CACCCTTGAGTACTATGTATT---CATATACCACATAGTATGTTGTAGAATGT-GATCATTGGGCCTTTGTGCCTATAATAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCACTAGCTTTTGTGAGCTATGCTGGGGCTTGGATGTTGGGAGTTTGCG-GGCC-TTTACT-------AGGTCGGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGACTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCACTGT-GGACAATCATTAATGAACTT-CTTGACCTCAAATCAGGTAGGACTACCCGC
Morganella_subincarnata          TTAGGATCATTATTGAATATACTTGATGGGTTGTAGCTGGCTCTACGGGGCATGTGCACGCTTGTCTTGATGTTATTC-ATCCACCTGTGCACCTTTTGTAGTCTTGGGGAGTCTGAGCAAGTTGAC--TATCGG----ATTCCGGATGTGAGGATT-GCTG-----------GCATACAGCTCTTCTCTAT-TGACTT---TCTCCCTCAAGTGCTATGTATT---CATATACCACATAGTATGTTGTAGAATGTTGATCAATGGGTCTATGTACCTATAGTAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCTAGTTCT-CT-AGTTG-----GGGCTTGGATGT-GGGAGTTTGCG-GGTCCTTTATCAC---AAAGGTCAGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCCCGTCACTAGTGTGATAATTATCTACACTGT-GAA-TAGATTGCTCTCTGACTAGTTCAGCTGCTAATAGTCCACTAT-GGACAATGTTGAATGAACTT-GTTGACCTCAA-TCAGGTAGGACTACCCGC
Vascellum_pratense               TTAGGA--CTTATTGAATATTCTTGATGAGTTGTAGCTGGCTCTCCGGGGCATGTGCACGCTTGTCTTGACTCTATTC-ATCCACCTGTGCACCTTTTGTAGTCTTGGGGGTTGAAAGCAG-TCGAC--TATCGGATGGACTCCGGATGTGAGAAAT-GCTGAGCGCAGAGACGCATACAGCTCTTCTCAAA-TGACTTG-----CCCTTGAGTACTATGTATT---CATATACCATGTCGTATGTTGCAGAATGT-GATCAATGGGTCTATGTACCTATAATAATCATATACAACTTTCAGCAACGGATCTCTTGGCTCTCGCATCGATGAAGAACGCAGCGAAATGCGATAAGTAATGTGAATTGCAGAATTCAGTGAATCATCGAATCTTTGAACGCACCTTGCGCTCCTTGGTATTCCGAGGAGCATGCCTGTTTGAGTGTCATTAAATTCTCAACCCCTCCAGCTTTTGCGAGTTGTGATGGGGCTTGGATAT-GGGAGTTTGCG-GGTCCTTTATCAG--TAAAGGTCGGCTCTCCTGAAATACATTAGCGGAACCGTTTGCAGTCCC-GTCACTAGTGTGATAATTATCTACACTGT-GA---TGATTGCTCTCTGACTAGTTCAGCTGCTAATCGTCCGGTAT-GGACAACATTTAATGAACTTACTTGACCTCAAATCAGGTAGGACTACCCGC
;
END;

BEGIN CODONS;
CODONPOSSET * CodonPositions (CHARACTERS = ITS_no_Lepiota) = 
		N: 1-703;
CODONPOSSET  CodonPositions (CHARACTERS = ITS_no_Lepiota) = 
		N: 1-703;
END;
[ The following blocks are output data for analysis step 1167 ]
BEGIN TREES;
      TITLE Tb5943;
      LINK TAXA = TaxaForAnalysisStep1167;
      TREE Fig._5 = [&R] (Bovista_plumbea,(Bovista_paludosa,((Bovista_pusilla,Bovista_polymorpha),((Morganella_subincarnata,((Lycoperdon_perlatum,Vascellum_pratense),(((Lycoperdon_mammiforme,Lycoperdon_echinatum),Lycoperdon_foetidum),Bovistella_radicata))),Lycoperdon_pyriforme_Bavaria))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5576]


END;