#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on May 02, 2024; 0:46 GMT TreeBASE (cc) 1994-2008 Study reference: Klopfstein S., Vilhelmsen L., & Ronquist F. 2015. A non-stationary Markov model detects directional evolution in hymenopteran morphology. Systematic Biology, 64(6): 1089-1103. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S16060] [ The following blocks are input data for analysis step 14055 ] BEGIN TAXA; TITLE TaxaForAnalysisStep14055; DIMENSIONS NTAX=62; TAXLABELS Abia Acantholyda Acordulecera Aglaostigma Apoidea_A Arge Athalia Blasticotoma Calameuta Cephalcia Cephus Chrysopidae Cimbicinae Cladius Corynis Cynipoidea Decameria Diprion Dolerus Evanioidea Gilpinia Hartigia Hoplocampa Ichneumonidae Lophyrotoma Macroxyela Mecoptera Megalodontes_sp._c Megalodontes_sp._sk Megalyra Metallus Micropterigidae Monoctenus Monophadnoides Nematinus Nematus Neurotoma Onycholyda Orussus Pamphilius Paraneoptera Paremphytus Perga Phylacteophaga Polyphaga_sp. Raphidioptera Runaria Selandria Sirex Stephanidae_A Sterictiphora Strongylogaster Syntexis Taxonus Tenthredo Tremex Trigonalidae Urocerus Vespidae Xeris Xiphydria Xyela ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M32741] TITLE Hymenoptera_Morphological_62x354; LINK TAXA = TaxaForAnalysisStep14055; DIMENSIONS NCHAR=354; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6" MISSING=? GAP= -; CHARSTATELABELS 1 160._Anterior_branches_of_subcostae_of_forewings / absent present_at_least_as_cross_veins_in_costal_cells, 2 182._Cross_veins_cu_a_of_hindwings / absent present_and_closing_cells_Cu_distally, 3 242._Radial_cell_of_forewing / open distally_closed_distally, 4 164._Rs_of_forewings_apically / not_furcate furcate, 5 159._Subcostae_of_forewings / fused_with_R_at_most_short_transverse_anterior_branches_present not_fused_with_R, 6 168._Cross_veins_2r_m_of_forewings / absent present, 7 169._Cross_veins_3r_m_of_forewings / absent present, 8 179._Cross_vein_lr_m_of_hindwings / absent present, 9 180._Cross_vein_3r_m_of_hindwings / absent present, 10 181._Cross_veins_m_cu_of_hindwings / absent present, 11 171a.._Posterior_anal_vein_proximal_part / absent present, 12 171b.._Posterior_anal_vein_distal_part / absent present, 13 248._Cross_vein_2r_m / absent present, 14 166._Cross_veins_2r_of_forewings / absent present, 15 177._Costae_of_hindwings / absent present, 16 178._Subcostae_of_hindwings / absent present, 17 245._Cross_vein_in_anal_cell_of_forewing / absent present, 18 165._Cross_veins_1r_of_forewings / incomplete_or_absent present, 19 170._Cross_veins2m_cuofforewings / reduced_or_absent present, 20 183._1st_anal_veins_of_hindwings / reduced_or_absent present, 21 249._Second_anal_vein_of_hindwing / partly_reduced_or_completely_absent present_and_complete, 22 163._Abcissae_of_forewing_Rs / absent_as_tubular_veins_between_cells_1R_and_Rs present_between_cells_1R_and_Rs_at_least_partly_as_tubular_veins, 23 247._Veins_Cu_and_M_of_hindwing_at_the_base_of_the_wing / completely_fused separate, 24 208._LateralT9_2ndvalvifermuscles / absent present, 25 115._Transverse_metanoto_metapleural_muscles / absent present, 26 86a.._Anterior_mesolaterophragmal_mesofurcal_muscles / absent present, 27 86b.._Posterior_mesolaterophragmal_mesofurcal_muscles / absent present, 28 154._S2_metacoxal_muscles / absent present, 29 281a._Muscle_nb / absent present, 30 281b._Muscle_nd / absent present, 31 281c._Muscle_nl / absent present, 32 282._Muscles_connecting_the_two_volsellae / absent present, 33 286a._Muscle_s / absent present, 34 286b._Muscle_si / absent present, 35 287b._Muscle_t / absent present, 36 288a._Muscle_u / absent present, 37 291._Muscle_w / absent present, 38 292._Muscle_x / absent present, 39 293._Muscle_z / absent present, 40 267._Muscles_a_and_b / absent present, 41 290._Muscle_v_and_gonomacula / gonomacula_vestigial_or_absent_and_v_absent both_present_and_fully_developed, 42 269._Muscle_c / absent present, 43 271._Muscle_d / absent present, 44 119._2nd_phragmo_3rd_phragmal_muscles / absent present, 45 70._Dorsal_mesofurco_profurcal_muscles / absent present, 46 149._Anterior_metanoto_metacoxal_muscles / absent present, 47 150._Posterior_metanoto_metacoxal_muscles / absent present, 48 49._Pronoto_postoccipital_muscles / absent present, 49 50._Prophragmo_postoccipital_muscles / absent present, 50 67._Profurco_prospinal_muscles / absent present, 51 68._Mesofurco_prospinal_muscles / absent present, 52 69._Prophragmo_profurcal_muscles / absent present, 53 80._Mesonoto_mesotrochanteral_muscles / absent present, 54 85._Mesonoto_mesolaterophragmal_muscles / absent present, 55 103._Metafurco_mesospinal_muscles / absent present, 56 108._Mesonoto_mesocoxal_muscles / absent present, 57 109._Mesobasalar_mesocoxal_muscles / absent present, 58 120._Metanoto_metalaterophragmal_muscles / absent present, 59 124._Metanoto_metabasalar_muscles / absent present, 60 127._Metalaterophragmo_metafurcal_muscles / absent present, 61 141._Metapleural_S2_muscles / absent present, 62 151._Metanoto_trochanteral_muscles / absent present, 63 152._Metabasalar_metacoxal_muscles / absent present, 64 153._Metabasalar_metatrochanteral_muscles / absent present, 65 51._Prophragmo_latero_cervical / absent present, 66 12._Labral_compressor_muscles / absent present, 67 13._Anterior_labral_retractor_muscles / absent present, 68 31._Ventral_salivarial_dilatators / absent present, 69 272._Muscle_e / absent present, 70 273._Muscle_f / absent present, 71 276._Muscle_i / absent present, 72 279._Muscle_l / absent present, 73 280._Muscle_m / absent present, 74 283._Muscle_o / absent present, 75 284._Muscle_p / absent present, 76 285._Muscle_qr / absent present, 77 101._Mesofurco_metabasalar_muscles / absent present, 78 131._Muscles_Tl_S2 / absent present, 79 143._Apodemes_or_tendons_receiving_the_insertions_of_the_posterior_metapleuro_metafurcal_muscles / absent present, 80 16._Sclerotization_between_the_occipital_and_oral_foramina / absent_and_occipital_and_oral_foramina_confluent present_and_separating_occipital_and_oral_foramina, 81 205._Dorsal_flanges / continuous_with_median_bridge_and_sites_of_insertion_of_the_posterior_T9 '-' 2nd_valvifer_muscles separated_from_median_bridge_by_weakly_sclerotized_strips_and_posterior_T9 '-' 2nd_valvifer_muscles_insert_on_apodemes_continuous_with_3rd_valvulae, 82 64._Profurcal_bridge / absent present, 83 92._Postspiracular_sclerites / absent present, 84 241._Forewing_tegulae / absent present, 85 325._Volsella / absent present, 86 332._Basivolsellar_bridge / absent present, 87 336._Ventral_median_sclerotized_style / absent present, 88 337._Dorsal_rod / absent present, 89 43._Dorsal_cervical_sclerites / absent present_as_separate_sclerites, 90 287a._Harpe / absent_or_fused present_and_separate, 91 335._Fibula_ducti / absent_or_small large_and_on_surface_of_ductus, 92 207._Median_bridge / proximal_ends_of_3rd_valvulae_fused_medially_or_closely_associated widely_separated, 93 14._Mandibular_foramina / confluent_with_the_oral_foramen separated_from_the_oral_foramen_by_sclerotized_subgenal_bridges, 94 130._Relation_between_metacoxal_foramina_and_petiolar_foramen / continuous separate, 95 128._Metapostnotum / continuous_medially divided_medially, 96 113._Connection_between_2nd_phragma_and_anterior_margin_of_metanotum / totally_fused_or_at_most_connected_by_narrow_strip_of_unsclerotized_cuticle separated_by_membranous_area_of_considerable_length, 97 59._Configuration_of_the_katepisterna / undivided subdivided, 98 187._2nd_abdominal_tergum / undivided longitudinally_subdivided_by_narrow_membranous_strip, 99 99._Mesofurcal_bridge / fused_as_mesofurcal_bridge anterior_mesofurcal_arms_entirely_separate, 100 95._Occlusor_muscles_of_the_posterior_thoracic_spiracles / arise_from_the_posterior_margin_of_the_mesepimera_ventrally_of_the_spiracles arise_from_separate_sclerites_posteriorly_of_the_spiracles, 101 204._2nd_valvifers_and_3rd_valvulae / totally_fused at_least_partly_separated_by_membranous_areas, 102 317._Penisvalvae / fused_dorsally completely_separate, 103 300._Fusion_of_cupula / fused_or_closely_connected_to_gonostipites_cupula well_separate_from_gonostipites, 104 129._T1 / continuous_medially divided_medially, 105 65._Prospinasternum / closely_associated_or_fused_with_the_anterior_margin_of_the_mesothorax entirely_separate_from_the_anterior_margin_of_the_mesothorax, 106 48._Cervical_lines / laterocervicalia_and_propleura_proper_at_least_partly_fused laterocervicalia_and_propleura_proper_completely_separated, 107 62._Profurcal_arms / articulated_or_fused_with_the_propleura not_closely_associated_with_the_median_surfaces_of_the_propleura, 108 310._Median_fusion_of_parapenes / fused_basally_or_completely not_fused_basally, 109 18._Postgenal_bridge / postgenae_closely_abutting_or_fused_between_the_occipital_and_oral_foramina_postgenae not_in_contact_between_the_occipital_and_oral_foramina, 110 38._Postmentum / absent present, 111 58._Prothoracic_katepisterna / absent present, 112 60._Laterosternal_sclerites / absent present, 113 90._Prepecti / absent present, 114 104._Mesothoracic_trochantins / absent present, 115 123._Hindwing_tegulae / absent present, 116 135._Anapleural_sclerites / absent present, 117 158._Metatibial_preapical_spurs / absent present, 118 254._Mesotibial_apical_spurs / absent present, 119 255._Metatibial_apical_spurs / absent present, 120 299._Cupula / absent present, 121 353._Cerci_of_male / absent present, 122 132._Apodemal_parts_of_the_metabasalares / reduced_or_absent present, 123 148._Metathoracic_trochantins / absent present, 124 9b._Tormae_medially / fused separate, 125 134._Association_between_metapleural_arms_and_T1 / fused separate, 126 210._2nd_valvulae / fused_distally separate_distally, 127 157._Lateral_metafurcal_arms / fused_with_metapleural_apodemes separate_from_metapleural_apodemes, 128 352._Tenth_tergite / fused_with_T9 separate_from_T9, 129 61._Anterolateral_corners_of_prosternum / fused_with_the_propleura_at_the_propleuro '-' procoxal_articulations separate_from_the_propleura, 130 42._Subspiracular_articulations / articulposterolateral_margins_of_pronotum_closely_abut_or_articulate_with_the_anterior_margins_of_the_mesopleura_ventrally_of_the_anterior_thoracic_spiracles separated_from_the_anterior_margins_of_the_mesopleura, 131 328._Parossiculus / fused_with_gonostipes_or_gonoforceps free, 132 326._Digitus_and_basivolsella / fused separate, 133 34._Paraglossae / reduced_and_or_fused_with_glossa well_developed, 134 17._Postoccipital_bridge / insertion_points_connected_by_internal_sclerotized_bridge_ventrally_of_the_tentorial_bridge insertion_points_of_the_ventral_profurco '-' postoccipital_muscles_separate, 135 137._Metasubalares / absent_and_muscles_insert_on_membrane present_and_receiving_metapleuro '-' metasubalar_and_metasubalar_muscles, 136 71._Posterior_apical_protibial_spurs / reduced_or_absent present_and_well_developed, 137 19._Dorsal_tentorial_arms / absent_or_reduced present_and_well_developed_in_contact_with_head_capsule, 138 139._Association_between_T1_and_metepimera / partly_or_entirely_fused separate_and_might_abut, 139 344._Eighth_and_ninth_sternite / closely_connected_or_fused separate_only_connected_by_membrane, 140 351._Ninth_tergite / membranous_or_subdivided largely_sclerotized, 141 188._Pleural_region_of_abdominal_segments / pleural_regions_of_2nd_to_5th_abdominal_segments_membranous pleural_regions_around_the_spiracles_sclerotized, 142 196._1st_valvifers / posterior_margin_closely_appressed_or_fused_with_T9 posterior_margin_distinctly_separate_from_T9, 143 195._Basal_articulations_between_the_anteroventral_margins_of_the_2nd_valvifers_and_the_proximal_parts_of_the_2nd_valvulae / articulation_absent articulation_present, 144 79._Transscutal_fissure / articulation_absent_or_not_fully_developed straight_line_which_is_distinctly_less_sclerotized_than_the_rest_of_the_mesoscutum_present, 145 2._Subantennal_grooves / no_conspicuous_grooves_ventrally_of_the_eyes grooves_extending_posteriorly_from_the_anterior_tentorial_pits_ventrally_of_the_eyes_present, 146 53._Dorsal_parts_of_propleura / not_inflected_and_not_differing_in_sculpture_from_the_rest_of_the_propleura inflected_and_delimited_from_the_rest_of_the_propleura_and_smooth_and_articulating_with_the_lateral_margins_of_the_pronotum, 147 44._Cervical_prominences / fully_retracted_cervical_swellings_face_laterally not_retracted, 148 117._Shape_of_cenchri / about_as_broad_as_long_more_or_less_round about_twice_as_broad_as_long_oval about_three_times_as_broad_as_long_or_more_very_narrow, 149 304._Parapenis / antero '-' medial_corner_drawn_out not_set_off_from_the_rest_of_the_gonostipes set_off_but_without_constriction with_constriction_and_parapenisjugum_narrowed, 150 250._Length_of_anal_cell_of_hindwing / as_long_as_cell_1Cu_posteriorly_i_e_second_anal_vein_joins_first_anal_vein_almost_at_the_same_point_as_cu '-' a shorter_than_cell_1Cu_i_e_second_anal_vein_joins_first_anal_vein_further_proximal_than_cu '-' a_and_first_anal_vein_extends_beyond_cu '-', 151 142._Metapleural_apodemes / at_most_shallow_ridges_present_and_the_anterior_metapleuro '-' metafurcal_muscles_arising_above_the_metapleural_ridges distinct_apodemes_present_and_from_which_muscles_arise, 152 144._Metepisternal_depressions / at_most_shallow_and_weakly_demarcated_concavities_present well '-' developed_depressions_accommodating_the_mesocoxae_present_and_separated_medially_by_the_raised_metathoracic_discrimen_and_delimited_laterally_by_distinct_carinae, 153 20._Shape_of_tentorial_bridge / broad_and_not_arched narrow_and_not_arched narrow_and_arched, 154 167._Position_of_cross_veins_2r_of_forewings / connected_to_Rs_proximally_or_opposite_to_cross_veins_2r '-' m connected_to_Rs_distally_of_cross_veins_2r '-' m, 155 215._Olistheter_scaling / dense_and_with_large_scales weakly_developed_and_scales_sparse, 156 214._Olistheters / diverging_ventrally_in_cross '-' section approximately_vertical, 157 206._Distal_ends_of_dorsal_flanges / do_not_project_posteriorly_of_ventral_margins_of_2nd_valvifers_and_membranous_areas_separating_2nd_valvifers_and_3rd_valvulae_more_or_less_vertical distinctly_produced_posteriorly_of_ventral_margins_of_2nd_valvifers_and_membranous_areas_separating_2nd_valvifers_and_3rd_valvulae_oblique, 158 47._Insertions_of_the_profurco_laterocervical_muscles / double_and_both_on_the_posterior_ends_of_the_cervical_apodemes_and_on_the_ventral_internal_margins_of_the_cervical_swellings single_and_only_on_the_posterior_ends_of_the_cervical_apodemes, 159 156._Anterior_metafurcal_arms / elongate_and_at_most_only_slighter_shorter_than_the_lateral_metafurcal_arms reduced_and_considerably_shorter_than_the_lateral_metafurcal_arms, 160 225._Shape_of_larval_antennae / erect_and_segments_are_well_developed_and_situated_concentrically flattened_and_except_sometimes_distal_segment_segments_are_reduced_and_lie_next_to_each_other, 161 87._Mesopostnotum / exposed_and_without_opening_ventrally exposed_and_with_opening_ventrally_for_the_tendon_of_the_mesoscutello '-' metanotal_muscle covered_dorsally_by_anterior_parts_of_metanotum_and_without_opening, 162 192._Ovipositor_apparatus / fully_developed_and_lepismatoid_ovipositor_apparatus_present partly_reduced_and_1st_valvifer_fused_with_T9_and_1st_valvulae_fused_and_2nd_valvulae_not_discrete_structures highly_reduced_or_absent_and_nonfunctional, 163 228._Larval_thoracic_legs / fully_developed_and_segments_of_unequal_size reduced_and_segments_of_equal_size vestigial_and_unsegmented absent, 164 189._Position_of_abdominal_spiracles_41313 / in_the_pleural_regions_whether_these_are_sclerotized_or_unsclerotized in_the_lateral_parts_of_the_terga_as_indicated_by_unsclerotized_lines_between_the_terga_and_the_pleural_areas, 165 174._Distal_hamuli_placement / in_zig '-' zag_lines in_straight_lines, 166 118._Posterior_part_of_cenchri / membranous_and_inflected_and_devoid_of_hooks not_inflected_and_hooks_present_on_the_entire_surface, 167 339._Form_of_glandula_mucosa / more_or_less_round_and_oval_and_or_straight_and_or_slightly_curved with_a_bulbus_at_the_side U_shaped sigma_shaped with_three_blind_ends with_a_distal_appendage, 168 23._First_flagellomeres / not_broader_and_not_much_longer_than_the_length_of_any_of_the_following_flagellomeres distinctly_broader_and_much_longer_than_any_of_the_following_flagellomeres distinctly_enlargened_and_distal_flagellomeres_absent, 169 40._Anterolateral_corners_of_pronotum / not_conspicuous_and_anterior_surface_of_pronotum_without_any_differences_in_surface_sculpture with_prominent_projections_and_separating_smooth_anterior_surface_of_pronotum_from_distinctly_sculptured_dorsolateral_parts, 170 343._Form_of_abdominal_sternum_eight_in_males / not_constricted_and_almost_rectangular rectangular_and_but_in_some_cases_distal_margin_concave somewhat_constricted_medially_or_very_narrow_overall strongly_constricted_and_with_scleritous_bridge_in_the_middle separated_into_two_sclerites_connected_only_by_membrane, 171 294._Orientation_of_the_male_genitalia / not_inverted_at_rest_orthandrous rotated_180_degrees_around_the_longitudinal_axis_when_at_rest_strophandrous, 172 97._Configuration_of_mesopseudosternal_sulci / not_reaching_the_anterior_margins_of_the_mesepisterna_and_terminating_in_the_discrimen_of_the_mesothorax terminating_in_the_anterior_margins_of_the_mesepisterna, 173 198._Lamnia_of_1st_valvulae / not_reduced_in_size_relative_to_radices_and_comprising_the_larger_part_of_the_1st_valvulae reduced_in_length_and_height_distally_and_taper_to_narrow_points, 174 246._Vein_Sc / R_of_the_hindwing '/' present_and_at_a_distance_from_C present_and_running_along_the_anterior_wing_margin_and_adjacent_to_C_which_is_partly_dissolved present_and_the_basal_half_running_along_the_wing_margin_and_the_distal_half_at_some_distance_from_the_wing_margin, 175 32._Glossa / present_and_distinct reduced_or_absent, 176 278._Muscle_k / present_and_inserting_on_the_valvura_or_the_ergot_or_at_the_base_of_the_valviceps present_and_inserting_within_the_valviceps_in '-' between_medial_and_lateral_side_of_one_valviceps present_and_long_and_directed_distally_and_inserting_within_the_valviceps absent, 177 66._Prospinasternal_apodeme / present_and_short present_and_large_and_projecting_far_into_thorax rudimentary_or_absent_and_at_most_external_part_of_prospinasternurn_developed, 178 275._Muscle_h / present_and_straight_and_inserting_on_apex_gonostipitis present_and_fanned_out present_but_inserting_on_the_gonostipital_arm_instead_of_the_apex_gonostipitis, 179 230._Larval_abdominal_legs / present_and_well_developed present_as_bulges completely_absent, 180 54._Propleural_sulci_and_ridges / present_and_well_developed reduced_or_absent, 181 81._Mesolaterophragmal_lobes / present_and_well_developed reduced_or_absent, 182 314._Harpe / rather_straight_and_about_as_broad_as_the_distal_edge_of_the_gonostipes_and_dorsal_and_ventral_edge_more_or_less_parallel rather_triangular_and_with_a_dorsal_flap_which_extends_into_the_gap_created_by_the_constriction_between_the_parapenis_and_the_rest_of_the_gonostipes, 183 98._Anterior_mesofurcal_arms / short_and_inconspicuous elongate, 184 100._Lateral_mesofurcal_arms / short_and_not_extending_towards_the_mesopleura elongate_and_extending_towards_the_mesopleura, 185 84._Shape_of_anterior_apodemes / short_and_rounded elongate, 186 4._Position_of_labrum / situated_anteriorly_of_the_tips_of_the_mandibles_and_partly_covering_them situated_posteriorly_of_the_tips_of_the_mandibles_and_at_least_partly_covered_by_them, 187 102._Mesospina / small_and_does_not_project_posteriorly_between_the_anterior_metafurcal_arms elongate_and_projects_posteriorly_between_the_anterior_metafurcal_arms absent, 188 190._Female_abdominal_spiracles / spiracles_on_abdominal_segments_1to8_well_developed spiracles_on_abdominal_segments_2to7_reduced, 189 191._Male_abdominal_spiracles / spiracles_on_abdominal_segments_1to8_well_developed spiracles_on_abdominal_segments_2to7_reduced spiracles_on_abdominal_segments_2to8_reduced, 190 146._Metathoracic_paracoxal_sulci_and_ridges / straight_or_curving_slightly_anteriorly_and_if_curving_posteriorly_then_only_for_a_short_distance_at_the_extreme_lateral_ends curved_posteriorly strongly_curved_posteriorly_and_extending_subparallel_to_metathoracic_discrimen, 191 268._Form_of_muscle_a / straight_and_inserting_only_on_spiculum fan '-' shaped_and_inserting_on_speculum_and_to_the_left_and_right_of_the_spiculum_on_the_basal_margin_of_the_ninth_sternite, 192 41._Posterolateral_margins_of_pronotum / straight_and_not_covering_anterior_thoracic_spiracles with_concavities_for_the_accommodation_of_the_anterior_thoracic_spiracles straight_and_covering_anterior_thoracic_spiracles straight_and_anterior_thoracic_spiracles_visible_in_lateral_view_but_surrounded_by_pronotayprepectal_cuticle, 193 222._Cornea_of_larval_eyes / subdivided_and_one_lens_for_each_ommatidium only_one_common_lens_and_covering_all_ommatidia_and_for_each_larval_eye, 194 239._Maxillary_palp / thin_and_not_leg '-' like thick_and_leg '-' like_and_with_a_setae '-', 195 288._Parts_of_muscle_u / two_parts_which_start_next_to_each_other_basally_but_are_separate_distally two_completely_separate_parts one_part, 196 56._Medioventral_propleural_margins / widely_separate_and_not_inflected parallel_and_inflected_and_closely_abutting, 197 88._Pseudophragma_of_mesopostnotum / 2nd_phragma_without_any_anteriorly_projecting_lobes 2nd_phragma_with_a_pair_of_lobes_pseudophragma_projecting_anteriorly_of_the_connection_between_the_mesoscutellum_and_the_mesopostnotum, 198 233._Larval_suranal_hook / absent hook '-' like_cuticular_structure_present_on_suranal_lobe, 199 27._Sitophore / absent present_but_does_not_extend_distally_of_the_functional_mouth present_and_extends_distally_of_the_functional_mouth, 200 46._Cervical_apodemes / absent short_and_barely_delimited_from_cervical_prominences elongate_and_distinct_and_projecting_posteriorly_from_cervical_prominences, 201 36._Rod_like_sensilla / absent specialized_sensilla_on_distal_labial_palp_segment_present_and_not_situated_in_invagination present_and_situated_in_invagination, 202 10._Epipharyngeal_lobe / absent distinct_lobe_present, 203 232._Larval_supraspiracular_glands / absent glands_with_openings_situated_dorsally_of_abdominal_spiracles_present, 204 28._Hypopharyngeal_lobe / absent membranous_lobe_situated_distally_of_the_sitophore_present, 205 30._Hypopharyngeal_pectens / absent minute_microtrichian_hairs_present_on_hypopharyngeal_rods distinct_brushes_formed_by_large_and_closely_appressed_microtrichian_hairs, 206 308._Basal_inflection_of_gonostipes / absent on_gonostipital_arm on_gonostipital_arm_and_parapenis on_parapenis_only, 207 76._Paddle_shaped_setae_on_anterior_part_of_probasitarsi / absent one_row_of_paddleshaped_setae_present several_rows_of_paddleshaped_setae_present, 208 258._Patch_of_bristle_shaped_denticles_on_the_ventral_face_of_some_or_all_plantulae / absent present, 209 25._Multiporous_plate_sensilla / absent present, 210 55._Propleural_arms / absent present, 211 57._Cervical_grooves / absent present, 212 63._Posterodorsal_profurcal_apodemes / absent present, 213 82._Anterior_apodemes_of_mesolaterophragmal_lobes / absent present, 214 83._Excision_between_apodemes_and_lobes / absent present, 215 116._Cenchri / absent present, 216 203._Ctenidia_on_1st_valvulae / absent present, 217 234._Larval_suranal_process / absent present, 218 259._Secretion_pores_on_plantulae / absent present, 219 301._Distally_directed_branch_on_lateral_side_of_cupula / absent present, 220 306._Notch_in_the_median_margin_of_parapenis / absent present, 221 307._Medial_inflection_of_parapenis / absent present, 222 311._Spine_at_the_distal_end_of_the_gonoforceps / absent present, 223 315._Ridge_on_distal_edge_of_harpe / absent present, 224 318._Hairs_on_valviceps / absent present, 225 320._Pseudoceps_and_valvispina / absent present, 226 321._Row_of_teeth_on_valviceps / absent present, 227 330._Basivolsellar_apodeme / absent present, 228 331._Gonossicular_apodeme / absent present, 229 334._Thorns_on_basivolsella / absent present, 230 348._Strengthened_line_on_ninth_sternite_parallel_to_its_basal_margin / absent present, 231 270._Muscle_running_from_the_left_to_the_right_edge_of_the_ninth_sternite_across_the_external_genitalia / absent present, 232 199._Sawteeth_on_1st_valvulae / absent present_along_the_ventral_margins, 233 209._Styli / absent present_distally_on_the_3rd_valvulae, 234 212._Sawteeth_on_2nd_valvulae / absent present_on_the_dorsal_margins, 235 201._Serrulae / absent present_on_the_sawteeth_of_the_1st_valvulae, 236 322._Ergot_on_penisvalva / absent normal_and_short_tip scale '-' like very_long wing, 237 78._Parapsidal_signa_on_mesoscutum / absent weakly_developed conspicuous, 238 8._Lateral_epipharyngeal_brush / absent present_and_connected_to_one_of_the_tormae, 239 11._Labral_apodemes / absent distinct_apodemes_connected_to_the_tormae_present, 240 7._Median_epipharyngeal_brush / absent present_medially_on_epipharyngeal_wall, 241 122._Apodemes_for_the_lateral_metanoto_metapleural_muscles / absent_or_small large, 242 52._Dorsal_propleural_apodemes / absent_or_weakly_developed present_and_distinct_, 243 161._Cell_1M_of_forewing / angular_anteriorly round_anteriorly, 244 162._Shape_of_1st_abcissae_of_forewing_Rs / anterior_ends_more_proximal_than_posterior posterior_ends_more_proximal_than_anterior, 245 193._Cordate_apodemes / apodemes_situated_on_the_anterior_margin_of_T9_in_the_females_absent present_and_fused_at_most_for_a_short_distance_to_the_median_walls_of_T9 present_and_fused_to_the_median_walls_of_T9_for_a_distance_corresponding_to_at_least_half_the_length_of_the_anterior_flanges, 246 91._Occlusor_muscles_of_the_anterior_thoracic_spiracles / arise_from_the_prepecti arise_from_anterior_margins_of_the_mesopleura arise_from_posterior_parts_of_the_pronotum, 247 110._Mesofurco_mesocoxal_muscles / arise_posteriorly_of_the_paracoxal_ridges_from_the_base_of_the_mesofurca arise_anteriorly_of_the_paracoxal_ridges_from_the_discrimenal_lamella_of_the_mesothorax, 248 39._Posterodorsal_margin_of_pronotum / at_most_slightly_curved deeply_curved, 249 45._Position_of_cervical_prominences / at_the_anterodorsal_corners_of_the_propleura ventrally_of_the_anterodorsal_corners_of_the_propleura, 250 121._Lateralmetanoto_metapleural_muscles / attached_to_the_lateral_margins_of_the_metascutum at_least_part_of_the_muscles_attached_to_the_1st_axillaries, 251 302._Gonocondyle / basally_directed distally_directed_loop, 252 89._Lateral_attachment_points_of_the_mesopostnotum_with_the_mesepimera / broad_and_exposed invaginated reduced_and_mesopostnotum_internal, 253 312._Gonomacula / can_be_covered_with_a_lid cannot_be_hidden, 254 295._Left_and_right_parts_of_the_genitalia / cannot_be_folded_towards_each_other can_be_folded_towards_each_other_to_hide_the_volsellae, 255 341._Coiling_of_the_part_of_the_vas_deferens_that_forms_the_vesicula_seminalis / coiled zigzagging_but_not_coiled straight_at_least_basally, 256 94._Posterior_thoracic_spiracles / concavities_in_posterior_margins_of_mesepimera_opposite_spiracles_undeveloped_or_quite_shallow spiracles_accommodated_in_distinct_concavities_of_the_mesepimera spiracles_covered_laterally_by_mesepimera, 257 346._Distal_margin_of_ninth_sternite / convex_or_triangular drawn_out_into_a_long_and_pointed_tip more_or_less_straight_from_the_right_to_the_left_side_and_as_if_cut_off concave, 258 345._Basal_margin_of_ninth_sternite_seen_from_above / convex_or_triangular rather_straight_from_left_to_right_but_each_half_concave triangular_but_clearly_concave_left_and_right straight_from_left_to_right_as_in_Cephidae_and_Siricidae, 259 184._Jugal_lobes_of_hindwings / distinct_from_vannal_lobes not_distinct_from_vannal_lobes, 260 261._Structure_of_posterior_margin_of_second_abdominal_sternum / distinctly_overlapping_anterior_margin_of_third_abdominal_sternum weakly_or_not_overlapping_third_abdominal_sternum, 261 296._Position_of_foramen_genitale / dorsal basal ventral, 262 211._Ventral_wall_of_2nd_valvulae / evenly_thickened more_strongly_thickened_medially, 263 316._Median_face_of_harpe / extends_about_as_far_cranially_as_the_lateral_face extends_much_farther_cranially, 264 264._Ovipositor / external concealed, 265 111._Basal_rings_of_femora / femora_not_subdivided femora_subdivided_and_with_small_proximal_parts_and_the_basal_rings_and_set_off_from_the_rest, 266 22._Male_antennae / flagellomeres_without_lateral_projections flagellomeres_with_distinct_and_slender_lateral_projections_many_times_longer_than_the_base_of_the_flagellomeres_and_projections_not_flattened_and_appressed flagellomeres_with_flattened_and_appressed_lateral_projections, 267 33._Shape_of_glossa / flat longitudinally_folded_into_an_inverted_U '-' shape_and_or_hoodlike, 268 35._Number_of_labial_palp_segments / four three two one, 269 226._Larval_epicranial_sulcus / fully_developed reduced_or_absent, 270 147._Termination_points_of_metathoracic_paracoxal_sulci / in_the_posterior_margins_of_the_metepisterna in_or_close_to_the_metapleural_sulci_at_the_anterior_margins_of_the_metepisterna in_the_metepisterna_some_distance_from_both_the_anterior_and_posterior_margins, 271 37._Ventral_premental_adductors / inserting_separately_on_the_prementum inserting_through_a_common_tendon_on_the_posterior_end_of_the_prementum, 272 5._Shape_of_labrum / labrum_broader_or_just_as_broad_as_high labrum_several_times_higher_than_broad, 273 260._Structure_of_anterolateral_apophyses_of_abdominal_sternum_three / laterobasal_corners_of_sternum_slightly_or_not_at_all_modified_as_apophyses sternum_with_distinct_horn '-' like_or_finger '-' like_basal_extensions_directed_forward_or_upward, 274 338._Glandulae_mucosae / left_and_right more_or_less_dorsal_and_ventral_or_distal_and_basal_of_each_other, 275 6._Epipharyngeal_wall / membranous sclerotized_and_continuous_with_labrum_, 276 96._Mesopseudosternal_sulci / mesepisterna_subdivided_by_oblique_longitudinal_grooves_corresponding_to_internal_ridges at_most_shallow_grooves_without_corresponding_internal_ridges_present, 277 105._Mera_of_the_mesocoxae / mesocoxae_with_distinctly_demarcated_mera_dorsally mera_not_delimited_on_mesocoxae, 278 155._Metafurca / metafurcal_arms_arise_posteriorly_on_the_discrimenal_lamella_of_the_metathorax metafurcal_arms_arise_anteriorly, 279 309._Inclination_of_parapenis / more_or_less_parallel_to_the_median_axis_of_the_genitalia inclined, 280 350._Eighth_tergite / more_or_less_rectangular apically_extended_to_cover_anus apically_extended_to_cover_male_genitalia_with_apical_excision_with_apical_appendage, 281 305._Basal_margin_of_parapenis / more_or_less_straight_from_left_to_right drawn_out_basally, 282 237._Number_of_articles_in_female_antenna / more_than_15 13to15 12_or_less, 283 175._Number_of_distal_hamuli / more_than_four_per_wing four_or_less, 284 26._Mandibles / neither_highly_asymmetric_nor_extremely_long_and_curved highly_asymmetric_and_with_distinct_molar_process_on_the_right_mandible_corresponding_to_a_concavity_on_the_left elongate_and_curved_and_lying_in_distinct_clypeal_furrow_at_rest, 285 1._Ocellar_corona / no_conspicuous_structures_around_median_ocellus median_ocellus_surrounded_by_ring_of_cuticular_teeth, 286 257._Denticles_on_plantulae / no_denticles_on_plantulae denticles_present_somewhere_on_plantulae, 287 185._Constriction_between_1st_and_2nd_abdominal_segments / no_distinct_constriction constricted_dorsoventrally constricted_laterally_as_well_as_dorsoventrally, 288 256._Plantulae / no_plantulae integrated distal, 289 342._Relative_size_of_vesiculae_seminales / normal very_small_and_lumped_together_and_squeezed_between_the_glandulae_mucosae, 290 333._Extension_of_volsella / not_beyond_gonoforceps well_beyond_gonoforceps, 291 323._Valvura / not_broader_at_apex flattened_and_broadened_at_apex, 292 24._Apical_flagellomeres / not_conspicuously_modified clubshaped reduced_and_with_flattened_apex, 293 133._Metapleural_arms / not_covered_laterally_by_metepimera covered_laterally_by_posterodorsal_parts_of_metepimera, 294 327._Cranial_end_of_basivolsella / not_covering_gonostipes covering_gonostipital_arm, 295 253._Female_foretibia / not_divided subdivided, 296 252._Fore_basitarsus_of_the_female / not_drawn_out distally_drawn_out_into_a_kind_of_spur, 297 263._Tergite_nine_of_the_female / not_elongated distal_part_distinguished_from_basal_part_of_the_tergite_and_distal_part_elongated_into_a_tip_that_extends_beyond_the_cerci as_previous_state_but_the_tip_is_further_drawn_out_into_a_long_and_pointed_and_strongly_sclerotized_structure_the_cornus, 298 93._External_parts_of_mesothoracic_basalares / not_enlarged conspicuously_enlarged_and_extending_ventrally_behind_the_anterior_thoracic_spiracles, 299 3._Clypeus / not_inflected inflected, 300 238._Female_antennae / not_modified modified_into_hammers, 301 112._Insertion_point_of_mesoscutello_metanotal_muscles / not_on_any_conspicuous_structure on_the_anterior_margin_of_the_metanotum flanked_by_small_projections_from_the_anterior_margin_of_the_metanotum_and_projections_not_continuous_ventrally on_distinct_projections_from_the_anterior_margin_of_the_metanotum_and_forming_entire_ring_of_sclerotized_cuticle_around_the_insertion_point, 302 347._Spiculum / not_prominent drawn_out extremely_long_and_directed_cranially, 303 349._Strengthening_of_basal_margin_of_ninth_sternite / not_strengthened thickened with_inflection_ventrally with_protruding_inflection_ventrally, 304 106._Mesocoxae / not_subdivided subdivided_by_distinct_transverse_grooves, 305 194._Lines_of_fusion_of_cordate_apodemes / not_visible_externally visible_externally_as_distinct_furrows, 306 329._Position_of_volsella / on_medial_face_of_gonostipes_or_gonoforceps on_ventral_side_of_genitalia, 307 289._Distal_insertion_point_of_muscle_u / on_medial_face_of_harpe on_lateral_face_of_harpe, 308 243._Tip_and_appendix_of_radial_cell_of_forewing / on_the_wing_margin away_from_the_wing_margin, 309 231._Number_of_segments_in_larval_abdominal_legs / one two, 310 240._Number_of_maxillary_palp_segments / one_to_five six, 311 21._Corpotendon / point_of_origin_of_the_ventral_cibarial_and_pharyngeal_dilatators_on_tentorial_bridge_not_conspicuous distinct_apodeme_present_anteriorly_on_tentorial_bridge, 312 73._Tips_of_apical_protibial_spurs / pointed_and_sclerotized blunt_and_membranous, 313 303._Apices_gonostipitum_respectively_gonostipital_arms / pointing_medially pointing_cranially, 314 9a._Tormae / present absent, 315 15._Occipital_sulci_and_ridges / present absent, 316 125._3rd_phragma / present absent, 317 136._Anapleural_clefts_in_the_metathorax / present absent, 318 140._Articular_inflections_on_the_metepimera / present absent, 319 145._Metathoracic_paracoxal_notches / present absent, 320 172._Basal_hamuli / present absent, 321 173._Distal_hamuli / present absent, 322 176._Secondary_hamuli / present absent, 323 186._1st_abdominal_sternum / present absent, 324 221._Larval_eyes / present absent, 325 235._Larval_subanal_appendages / present absent, 326 274._Muscle_g / present absent, 327 313._Internal_membrane_between_the_harpe_and_the_gonostipes / present absent, 328 200._Distribution_of_sawteeth_along_1st_valvulae / present_along_at_least_half_of_the_length present_for_a_short_distance_distally_only, 329 266._Larval_abdominal_legs / present_on_abdominal_segments_1to10 present_on_2to8_and_10 present_on_2to7_and_10 present_on_2to6_and_10, 330 74._Probasitarsal_notches / probasitarsi_unmodified probasitarsi_with_distinct_notches_opposite_the_calcars, 331 75._Probasitarsal_combs / probasitarsi_without_any_specialized_setae_proximally with_few_and_well_separated_setae_in_a_row with_many_and_closely_set_setae_or_row_of_platelike_structures, 332 297._Apex_gonostipitis / proximal_of_parapenis distal_of_basal_margin_of_parapenis, 333 224._Number_of_larval_antennal_segments / seven six five four three two one, 334 340._Ductus_ejaculatorius / short extremely_long_so_that_the_glandulae_mucosae_are_far_away_from_the_genitalia, 335 223._Position_of_larval_eyes / situated_dorsally_of_the_bases_of_the_antennae situated_ventrally_of_the_bases_of_the_antennae, 336 107._Medial_articulations_of_mesocoxae / situated_proximally displaced_distally_along_the_medial_surfaces_of_the_mesocoxae, 337 324._Phallotrema / situated_ventrally situated_apically, 338 244._Structure_of_forewing_tip / smooth_or_coriaceous corrugated_and_wrinkled, 339 265._Labrum_of_the_larva / symmetrical asymmetrical, 340 227._Intrasegmental_annulation / thoracic_and_abdominal_segments_subdivided_by_superficial_transverse_grooves grooves_on_the_body_segments_absent, 341 229._Claws_on_larval_thoracic_legs / two_claws_present_on_each_leg one_claw_present claws_absent, 342 72._Anterior_apical_protibial_spurs / unmodified modified_into_curved_calcars_with_velum_for_use_in_antennae_cleaning absent, 343 236._Segmentation_of_subanal_appendages / unsegmented segmented, 344 77._Median_mesoscutal_sulcus / well_developed reduced_or_absent_and_internal_ridge_at_most_developed_on_1st_phragma, 345 126._Metalaterophragmal_lobes / well_developed weakly_developed_or_absent, 346 138._Metepimera / well_developed_throughout posterior_parts_of_metepimera_reduced, 347 202._Annuli_of_1st_valvulae / without_hairs with_narrow_fringes_of_hairs_on_their_margins, 348 213._Ventral_margins_of_2nd_valvulae / without_hairy_lobes with_hairy_lobes, 349 298._Ectophallic_membrane / without_pockets forms_two_flat_pockets_on_ventral_side_of_genitalia, 350 319._Tip_of_valviceps_or_penisvalva_viewed_from_medial / rounded_and_not_drawn_out drawn_out_into_a_pointed_thread '-' like_structure, 351 262._Anterior_margin_of_abdominal_sternites / straight_or_smoothly_curved_or_forming_a_slight_angle with_median_incision_or_angular_notch, 352 197._Banding_pattern / neither_1st_nor_2nd_valvulae_have_alternating_zones_of_differing_degrees_of_sclerotization 1st_or_2nd_valvulae_have_alternating_zones_of_differing_degree_of_sclerotization, 353 114._Lateral_metanotal_processes / blunt_or_inconspicuous slender_and_projecting_ventrally_or_medially, 354 29._Infrabuccal_pouch / small_or_with_inconspicuous_armature large_and_with_distinct_cuticular_teeth_developed_for_grinding_pollen, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Abia 0110011111110110?11110010101110001100001010110000100?10??000000001011111011100001011100001000001?01011100000?100001001111101111100101101101111100012211011??1010000001000{13}1?0001100100011000001110?00012001001?{01}?1011010000000000000000100120000110?12?10?00?00100000?00100001000001110002000102001101000010103001?0110100100?110110111010005000010010?01?11000100 Acantholyda 11101111111101111111110??11?1100??11000?11?1??????1?11???1?1?????????1111?11??11?011100001011000011?101110001111001011111111101110011111111110100010010000??1?00001000000000100?0?210011010001?0?0110?2210?00?1001111010?000?000000000?1110?000001001??01?0010?002001??0100000?00001110?0002000100?000??0010??1001?0?1010000000000100??0?0000010000021101000000000 Acordulecera 0110011110000010?111000?????11011011000001????????0?00????????????????110111???0??011000011?000??01??1?000?1?1000???11100???1??01011?1?1?0111??000111??00????????00?11000011?000?020100?00?000?1?0200??2?????10??10??01?0??000000000000????20?????0??2?10??0?0?10000??001000?1?000?11?0002100?02?00000000010??1??100?1?00?0???110110??1?2000600?10?0?0?0?0??000??? Aglaostigma 1110011111110?1011111101????110001110001010??????????????????????11?11110111???01?111000?100000??0?0111100?0?1001??101111???11?110111101111111?0001031?001??1??0?0??001001110000?000?1???0?00001?020??110000?200010???11?0000000000000010112000???0?12?10?0??01200000?001000010000011?100200010200?000000010?11?011011?00?0?11110010?1101000200?000011?0?0000001?0 Apoidea_A 0110011010000110?011000?????000000000000?10??????????00??????????00?11110110???1?1011101000?110??00??110?0?101??0?0?01110???0001?0111??0?0101??10?0?0??1?012??1?20??1?00000??00220?10?11112?0??2?0?1??220????30001101?0??00000?00100000????????????1???01?12?02?020020??1010???010?1110101000022000000000010?11??1?0?1?00?10???1001??0???121?1??100??1?111??000??? Arge 1110011111110010?1111100111111011011000001?1000001000100000100010111?011011110000011100001100001101001010000?11000101111110110111011110110111110001211100?00?110000?01320011000010000001002000?1102100120000000001011010001000100000000100120000010?12010000?0?103000000100001000100110002000002?01?1000001021100101010000000?110110111030006000100010?00010000110 Athalia 0110011111110110111111011??1000001100001010111000100?10??011011001011111011111101011100001000001101011110000?110101001111101111110111111111111100010211011??1010000?00000311000110000101?020000110?00012000002?0?1010?10000000000000000101100000000?10?10000?00201000?00100001000001110002000002000000000010011001?0110000000111001010101000200000?010?00000000110 Blasticotoma 11100111111101101111100??11?????????????0??1?????????11????1??????1?????????1??0001110001?000001101?10110001?1110011011111001111101?11?1?111111000101110?1000?0000010001000?000?0?200001000001?110?000121000001??10110100?00?00000??0??1010?0000011?1??10?00?0020?0?0000100002?0?001110002000?0200?000??00100??001?0?1?00000011100100??0?0001000?1001110000001?0?0 Calameuta 01100111111101101111110?0???000010000010?101?????????1?????1?????00?111000101?110011101000000011000?10111001111100101111101?11111101101011111110001?101000??0?0?00??1?000000010220?10?10010002?0?0?11?1220?0001001111000?00000?0000000010102000000001??01?01?02003000??01000?0?00010110000000112?00000000010000001?0?11002010111001??1?0?110?0?0?0???1?01000001010 Cephalcia 1110111111110111111111000111110011110001111110101111?11??11101100100011110111011?0111000010110000111101110001011001011111111101110011111111110100010010010011100001010000000100100210010010001?0101101221000011001111010000000000000000111030000010011?0000010000200101010000000000111000002000100000000001000100100?1010000000000100010?0000010000021101000000000 Cephus 0110011111110110111111000111000010000010?1011110100111111101001000011111001011110011101000000011000110111001111101101111101?11111101101011111110001?111000110000002?1?000000010220210?10010002?0?0?1101220?00010?1111000100000?00000000101020000000011001001?020030000?0100000000010110000000112?00000000010001001?0?11002010111001000?0?11030?0?00021001000001010 Chrysopidae 1??11?????11??11??????1??11????????????????1?1111?1??1011111?11???11????????1?11?001???????00010?01?0??1110?10?0?10?0????1111?1?11??111?01??0??0001???100?????100100???00????01?0?200?00002000?000?0000000000????001110?0???????????????0???000000000??010?0???0??1????000?112?0??0100?????00?00???00?000?00???0?????00??000001111001????00?4?10???002?000??????00 Cimbicinae 011001111111011011111001110?1100010100010101100??100?10??0000000????1111011110001011100001000001?01011100000?100001001111101111100111001101111100012211001??101000000130021?00011101000110000011102000120010010001011010000000000000000100020000110?1?010000?00100000?00100001?1?00111000200010200010100001020300100110100000?11011011101000500001?010?01?11000100 Cladius 1110011111110010?111100?????000001100001010??????????????????????10?11110111???0??111000010?000?10???11100?0?1100?1101111??111?110111101?0111??0001121?01????????0??00300111?002?200?1???0?00001?0?0??120????101011???1??000100000000001???10?????0????11?0??00100000?001000?1?000?11?0102000102001000000010?11??1?011?0000???11011??0?0200030??000?10?0?0??0101?? Corynis 0110011111110110?111100?1???1100111100010101?????1???10??1?0????????11110111??011011100001000001101?11100000111010100111100?111?10101101101?11100011111021??101?00?001000311000112001001?0200001?0200?11?01001?0?101??10?0000001000?000100120000010?10?10?00?00200000?001000?1?0000111020200010200010100001001300100?1000?001?110?1??1?0100050?001?0?0?11?10000100 Cynipoidea 0010010000000100?000000??10???????00???????11????00?01000??1????????????????0?010101100000?00100?00?0?10?00?110000000111000?00111001011010?11111010?1?112?11??1?203?1??000??000?2?210?11012110?1?0?11?1201?02?0?111011000?00?0?000??0??001??200000?122111?02???2??1011?0101111100??1110201100?20?00000??001001?101?0?0100?100?110111?????120???0?1?1?1?101?000?000 Decameria 0110011110010010?111010?????11011011000001?10????10?000??0???????00???110111???0?0011000011?0001?01??1?00000?1000??001100??11?11101101?1?0111??000111??00?????1??0??11000011?01000?01001?02?00?1?020??120????0000101??1??0?000000000000????20?????0????10????00?0300??0010?3?1?000?011000?000002101000??0010?11??101???000000?11011???1??000?0??10???0?000??000??? Diprion 01100111111100101111100?111?000000?00001010??????10??1?????1?????10?11110111??10001110000?000001101?111100???1110?1?01111?01111?1010?1?1?11?11?0001231100???1?11?0??0120031100010000010?0000?011?0?00?120????10001011011?00000000100000000?00???010?1??10?00?00101000?001100?1?000?11?1000000102001000000010001?01?011?100000011001??11?100040??00?010?0000110011? Dolerus 111000111111?110111111??111?11000110000101?11????10?01100011??????1??01101111??01011100011000001101?11010000?11010110111110111111011111111111110001030100???1010000?000002110001010001010000000110?100120000010001111010000000000000000101100000010?1?010?00?00200000?00100001000001110012000102001000000010012001?0110000000111001011?010002000001011?00010000110 Evanioidea 0010000000000100?010010????????0??00??0????1????????0??????????????????????????1?001100?000?0100000?1?1000010?11000001111?0?0001101?0?1010101111010?1?112???1?1?20??1?0000??000?0??10?01112110?2?0?11?1201?0??0?11111000??0010?000??00?101??2000000112?11?12?0??01101???1010?1?0??11110001100?20?0?000???010001100?0?1100?101?11011??0?1?120???100???1?101?000?00? Gilpinia 111001111111001011111001111100000000000101011000010001100011001000011111011111100011100001000001101011110000?11010100111110111111010111111111110001231100???1011000?012003110000000001010000000110?00012000001?001011011000010000100000000?30000010?10010000?00102000?00110001000001111000000102001000000010001001?01101000010110?10111?1000400000?010?00001100110 Hartigia 011001111111011011111100?11?000010000010?111??????0??1?11??1?????00?11110010??11001110100000001?000?11111001111111101111101?11111101101011111110001?101000110?00002?1?000000010221210?10010002?0?0?1101220?0001001111001100000?0000000010102000000001?001?01?020030000?0100000?00010110?00000012?00000000010000001?0?1?002010111001000?0?11030?0?00021001000001010 Hoplocampa 1110011111110110?111100?????000001110001010?????????0????????????11?01110111???0??111000010?000?10???11100?0?1100?1101111??111?11011?101?0111??0001131?011?????0?00?0000021??001?000?1???0?00001?020??12?????1?101????1??00000000000000????10?????0????10?0??00100002?00100001?000?11?1002000102001000000010?12??11011?0000?0?110010?01?2??0300?00001??0?0??010??? Ichneumonidae 01100?1100000100?011000?????110001000100?11?????????0000?????????00?11100111???1?0011000000?0100?00??010000100000?0?01111???0??1101101?010101??0011?1??120????1?20??1?00000??00201?10?11?02?00?1?0?1??120????00?11111?0???1000?00000000????22?????01???11?12?12230110??01010?1?00??1110000000?20?000000000?0?10??1?0?0?00?10??11011??0???120?0??00???1?101??000??? Lophyrotoma 1110001110000010?111010?1???00011011000001???????1?????????1?????00???1101110?000001100001100001?0?001?000?0?1000??001101?01101110111111?01011?000111?000?????1??0??11000011000002?0?0???02?00?2?020??110????1000101??10?0?0000000000001?0130???000?2??10????0010300??001200?1?000?01?000?000002100001??0010211?1101?1?000000?11011???10?000?0??11???0?0000000011? Macroxyela 111111111111111111111110011?11100111000011111????111?11??1?11????101111111111?100001100001000000001?11111000?111101111111111111111111111111111100010100000100000000?00010100001000110010000000?0101000111000011001101010000000010000000000?1010001001??00?0000000000100010?000000000110002010001000000000010001000000000000000000010000?000010000000110010?0000001 Mecoptera 1??11?????111?11???11?1??11?????????????0??1010?10011100111011111100??????????11?0010???1???0000?01??????10?1??00110011011111?1110??011101110??0001???101?????100200???0000??01???000?00?02000?000?0001000000????1100?0?0??????00???????????0000000????001?0???00?10???000?2020?000100?0?0?00?00???00?000?00???0????0000?000011111101????00?4?00???010?110????0?00 Megalodontes_sp._c 0110011111110110111111000??1110001110000111111100100?11??10100000101011000001011?0111000010110000011111010001110101011111101101110101111111110100012011020??010000??10400101100000210010010001?0?011012210?0012001111010000010000000000110040000000112?01000100002000?10120000?1000011010002000100100000001001200100?1100000001101100010?0000010010021100000000000 Megalodontes_sp._sk 01100111111101101111110?????1100011?000?1111?????????1???1???????00?0?100000???1?0?11000010?100?001??110?0?01110??1?11111??11?1??010?1?1??1?1??0001201?020????0??0??10?00101?00200?1001??1000??0?0?1??221????120?111??1??0001000000?000????4??????01???01?0?10?0020?0?101200?0?10??0110100020001?01000??0?10?12??1?0?110000???11011??????000????010??1??????000??? Megalyra 00?0000000000100?000000???????????00????????1?????0?1??00?????????????????????0101011?0000?0?000?00?1??0000?0?00000001111?0?000010100??0?0111111110?1?11??101?1?20??1??000??020?0??10?111?2110?3?0?11?1201?12?0?11101000??00?0?000??00?001??000000010?111?12????011001??1?11?11????1110201100?20?0?000???0100201?1?0?0?012?01?11001??????120???1?0???1?011?0?0?000 Metallus 0110011100010110?111100?????00000110000101???????????????????????11??0110111???0??111000010?000?10???10100?0?1100??101111??11??11011?111?1111??0001031?001???????0???0000111?001?200?1???0??0001?0?0??12??0??10?010???1???0000000000000????10?????0????11?0??02200000?001000?0?000?11?1302?00?02?01000000010?11??1?00100000?1111011??11?100020??000010?0?0??0?1??? Micropterigidae 1??11?????????00??????1????0????????????0??1?101?????????1?011??1100?????????0?0?0010???????0010001????1110??110?1?????0?1111?1?11??111?01??0??0001???000?????100200???00?0??00?1?000?01?0200??001?0001020?00?0??00???0?0???????????????????0101000????001?0??????1?????00021?00??0100???0?10?00???00?000?00???0????0000?001011111001????00?4?10????10?011????0?11 Monoctenus 1110011111110010?111100?111?00000110000101011????10??11??0?1????????1011011111?00011100001000001101?1111000??11010100111110111111010110111111110001231100???1?11000?010003110001000001010000000110?0001200?0?10001011011000000000100000000?20000010?10010?00?00100000?00110001?00001111000000102000000000010012001?01101000000110?10111?10004000000010?00001000110 Monophadnoides 1110011100010110?111110?????11000111000101???????????????????????11??1110111???0??111000110?000?10???10100?0?1100?1101111??11??11011?1?1?1111??0001031?001???????0???000021??000?100?1???0??0001?020??120?0??101010???1??00000000000000????10?????0????10?0??02100000?001000?1?000?11?1002000102?00000000210?11??11011?0000?1111011??11?100020??00?011?0?0??000??? Nematinus 1110011111010010?111100?????000001110001010??????????????????????11?11110111???0??111000010?000?10???11100?1?1101?1101111??11??110111111?1111??0001131?01??????0?0??0000011??001?000?1???0?00001?020??120????10?010???1???0000001000000????20?????0????10?0??00100000?00100001?000?11?1102000?02000000000010?12??11011?0000??1110010?01?200030??00001??0?0??010??? Nematus 11100?1111010010?111100?1???00100111000101011????10??11??0?1????????111101111?101011100011000001101?11110001?11010110111100110111011110111111110001131101???1011000?00000{12}11000100000101?0000001102000120000010101110?100000{01}00010000001001{12}0000000?10?10?00?00100000?001000?1?0000111140200010200100000001001200110110000000111001010102000300000?011?00010000110 Neurotoma 01101111111101111111110?????1100??11000011?1?????????11????1????????0111101???11?011100001011000011?1011?0011010001011111111101110111111111110100010010020??1?0000?01000000?100??0210010010001?0?011012210?0??1??11110100?00?00000??0001100?000001001??01?00100002000?10100000?0?001110000020?0100?00000001000100100?1000000000000100??0?000001001002110100000?100 Onycholyda 11101111111101111111110?????1100011100001111??????1?111??1????????0?01111111???1?0111000010?1000011?1011100010100?1011111??11?1110111111?1111010001001?020????0??0?010000000?00000?10010?10001?0?011??221????110?111??1??00000000000000110?30?????001??00?0?100002001?101000?0?000?1110000020001000000000010?01?0100?10000000000001??01??000?0??01???1?010??0001?? Orussus 0100010000010100?001000?0??1???000000000?1111010100110100001010000011111?11111010101100000000000?0011010100000011000011110011011101101100110111111101?1120100000203?1030000?000021211?11?10120?2?0?10?1201?0012?1100??100?0000?00000111001?32010000?20111002?1?2000000?11011?1110011110202101?00010000110111011110???01000100111001110???12060?00001?1?11100000000 Pamphilius 111011111111011111111100011?110001?1000011?11?1??11?11101111????????01111011??11?01110001?0110000111101110011110001011111111101110011111111110100010010000011100001010000000100?00210010010001?010110122{01}0000?10011110100000?00000??0001100?0000010011000?00100002001010100000?0?001110000020001?0?000000010001001?0?10000000000001000?0?0000?1001002110100000?000 Paraneoptera 00?1?00100???010??00?0??????????????????0??11?101111?11111?1??1?011???????????00?00?0??????10000??1????0111????001?00??0?111111?11??111101??000000????000?????0?00?0??000?0?1?1????01?00?0200??0???00?1000?00?0??000??00???????????????1?00?000101?00??0???0???0??1?????00?3???0?00110?????10?0????00????00????0??????00?0100111110????0?00??0?0?????0?01100???00? Paremphytus 11100111111101101111100?111?00000111000001111????10???1???11????????11100111??10?0111000?10?0001101??0110001?11100110??111001?111011111111111??0001?111001????0?00?100?2000?000200?00011000001?1?0200?12?0?0?0???1011010??010000000010010?010000011????10?00?002200?0?001000?2?0?001110002000?0??01?00??00100010?110??0000000111001??010?000?0?001???1?0000001?000 Perga 011001111000?010?11101?11??????1??1?000?01?10?1??10??00??0?1???1??1???11?11?11000011100001100001?01001?00000?10100101110?001101?1011000110??1110001?1?100????11?000?11?00?110?0000010001?02000?1?0?10?1200?001?0?1110?1000?0000000??0001001?0000000?12?10??0?0?1?300??001001?1?0??00110?02000002?0?100??01102?1001??0?0000000?1101101??0?000??0011?010?0001000?110 Phylacteophaga 0110011110000010?111010?1??000001010000?01?100100100?000000101000111??1?011100000011100001000001?01001?0000??10100?0111000011011101111001011111000121?100????11?00??11000010000000?00001?02000?1?0?10?1200?00?0001110?10?0?000000???000101120000000?12?100?0?0223300??001001?1?00001110012000002?00000??0010211001?0??0000000?11011???10?000?0?010???0?00000000110 Polyphaga_sp. ?1???????1????01???11???0110???????????????10111101110111111?1110???????????1111?0000???????0000001????01?1?1?100100011??1111?1?11??111101110??001????01??????0?0??0???0000???1?2??11?00?02?00?0?0?00?0??0?00????0?0??0????????00???????????000101????00?0???0?00?0?????00?1?2?0??1110???2?00?00???00?????10???0?????0?0?0100111110??????00????0?????0?000????0?01 Raphidioptera 1??11?????11?111?1?1?110?110????????????0??10?111????1???1?11???0111?????????111?0010???0??10010?01?0??1110?10?011000110?1111?1?11??0?1?10??0000001???101?????100100???00?0?001?0?200?00?02000?000??000000000????0010?000??????0???????01???000000000??000?0???0??1?????00?101?00?0100???0?00?00???00?????00???0?????0???010011111001????00?4?00???102?110????0000 Runaria 11100111111101101111100?1??1101011110000011110101101?110011100110011111101111010?0111000110?000110?01011000??1110?11011111001??11011111111111??0001?111?01???00??0?100020000000200?000???0000??1?0200?12?0?000100101??10?00100000000000100010000011??201000??00220001?001000??00?001110002000002000?00??0010001??110?10000000111001??0?0?000?0??01???1?000000100?0 Selandria 111001111111?110011111??????00000111000101011?????0?0100?0???????11?11110111???0?0111000010?0001101??1110001?1100??101111???1?111011110111111??0001131?001????1??0??00400111?00102000100?00?0001?020??1200?0?1010101??1??00000000000000????2000???0????11?0??00200000?001000?1000001110002000102000000??0010?12??110?1?00?011111001??11?100020??000?11?000??000??0 Sirex 01001111111101101111110?0??11100??1100?01110111?0001?11??101010000011?11101111110011101001?0001010011?11100?001110100111111?111111?1011011?111101010110000010000?02?00001?0?000021210010?11000?0?001101220000?10?100??101100000000??00?1010?000001012??1000110?01300100010011201??11110?00000001?0?00000201001?01101?00012001011011110?1?000???0?11121?10000000000 Stephanidae_A 0100000000000100?000010?????000010000000?01?????????1???????????????11110111??010001100100000000000?10100000011010000011100100101011011010111111110?0?112???1?00203?1?00010?000201210?11110120?2?0?10?1201?113???101100?0?0000?00010100101?2201000012??11?12?1?201110??01010?1?1?011110000001?20?01001?00110020100?0?11000101?110111?1?1?12061?000???1?011?0000000 Sterictiphora 0100000111110010?111100??11?11001011000001?10????100?10????1?????????1100111??000011100001100001101?01000000?110001?01111?0110111011110110111110001211100????11?000?01320011000000000001002000?110210?1200?00000?1011010001000100000000100100000010?1??10?00?00100001?00100001?00100110002000002100?1000001021200100010010000?110?10011020006000100010?00010000110 Strongylogaster 111001111111?110011111???11?000001110001010???????0?01???????????11?11110111???0??111000010?000?101??11100?1?1100?1?01111???11?11011?1?1111111?0001030?001?????0?0??0030011??001?000010?00?00001?0200?120?0??10101??101??0000000010000010??20?????0?1??10?00?00200000?00100001?000?11?1002000102000000000010?12?011011000?0?11110010?0101000200?000011?0?0??0001?? Syntexis 010000110011011011111100011111001110000?0?1110101011111??111011000011011?11?10110011100001?00011000110111000111110100111?11?111?11111010111?1110000111000?000100102?10?00?00000020210010010001?0?0?1001221?0002?111010101?00100000??0??101030000000020?00001?0?00?00100010111001??10110?00000?00?000000010100??011?0?1000201001100111111?0104??0001121?01100001000 Taxonus 11100111111101101111110?????1110011100010101?????10?011??????????11?11110111???0?0111000110?0001101??1110000?1101?1101111??11?11101111?111111??0001030?001????10?00??0000211?0010000010??00?0001?021??120?0??100011???1??00010000000000????20?????0????10?0??00200000?00100001?000?1110002000102000000000010?12??11011?00000?1110110?11?1000200?000011?000??000??? Tenthredo 011001111111?110111111???11?11000111000101010????10?011001?1?????11?111101111?101011100011000001101011110000?110101101111101111110111101111011100010301001??1010000?0020021100010000010100000001102100120000010101111010000000000000000101100000010?1?010?00?00200000?00100001000001110002000102000000000010012001101100000011110?1011?010002000000011?00000000110 Tremex 11001011111101101111010??01?110001110000111???????0?11?11????????????11?1011??110001101001?00010100?10111001001110100111011?1111111001101111111010101?000???0?00002?00?0110?000021210010?11000?0?0011012200001100110??1011000000001?0001010?000001012?010?0110?013000?00100212?1??11110101000001?0100000201000101111?00012001011011110?1?0005??0?11121?10000000000 Trigonalidae 0?1001100?000100?01?010??10?1100??00?0000001????????00000???????????1111?11?0?01?1011000010?0000?00??1100001000000000111100?0001101011?010101??1010?1?112001??1?20??1?00000??20000?10?01112?2??2?0?10?1201?11?0?0110100???00100000??000?????20000001??111?12?0?10110200?1001???1?0?1110000000?22?01000???0100211?1?0?11002101?11001??01??120???100???1?111??00??00 Urocerus 110011111111011011110100?01?11001111001011101??0000111111101????000?1101101111110011101001000010100?1011100?001110100111111?11111111011011111110101011000001010?00??0000110?000021?10011011000?0?0011?1220?0011001101010?100000000100001010100000101200100011000130010001001?2?1001111000000000100000000201001101101?00012001011011??0?1?000?0?0?1???1?10000000000 Vespidae 0110011010000110?011010?0100000000001000?11110000000000000000000000?10110111000100011001000?00?0000010100000001000000111000?000110101?1000001101010?0?1120??0111203?1?50010?000120210?11112000?2?0?110120101221?111110000?0011?00010000100002000000102111012?02001111???101001111011110101000?200000000000100211?0?0?1100?101?11011110?1?12160?0?001?1?11100000000 Xeris 11001111001101101111010?????110001110010111??????????????????????00?11111011???1??111010010?001??0???011?00?00111?1?01111???11?1110101?0?1111??010101??000?0?????0??00001101?000?1?1?0??????00?0?001??122????11?011???1???000000001?000????10?????01???10?0?1000130010001001?2??00?11?0100000?00000000??2010?11??101?0?01?0??0110?1??01??000?0???11??1?1?0??000??? Xiphydria 111001111111011011111101011100001011000011101010000111101101000000011111011111010111100001000010000110111000111110000111110111111011111011111111101011100010110020201000000?000000210011010001?1?001101221?0012001001110100010000000000101021000000020010012100120000000100110010011110000000002000000000010?0100100?0000010001001111011?00040?0010121?00100000000 Xyela 111111111111011111111110011111100111000011?11110111111111111111011011111111111100001100001000000001111111000?11110?011111111111111111111111111100010100000000000100000010210000000110010000000?0111000111000011011001010000000010000000010?10100000010000000000000001000100000000000110002010001000000000010001000000000000000000010100?00001010000011?01000000001 ; END; [ The following blocks are output data for analysis step 14055 ] BEGIN TREES; TITLE Hymenoptera_Morphological_62_taxa_1c.con; LINK TAXA = TaxaForAnalysisStep14055; TREE Fig._1c = [&R] (((Paraneoptera:0.1503251360284045,Polyphaga_sp.:0.09049889086642904):0.09493448407833482,((Chrysopidae:0.0313252431564337,Raphidioptera:0.08808364339316355):0.04434520989225255,(Micropterigidae:0.1670249118897951,Mecoptera:0.06616987459536583):0.08243505681430174):0.05469878197833511):0.2556329940875146,((Xyela:0.02785873482777035,Macroxyela:0.004164479624417794):0.05545739515339543,((((Runaria:0.01434722293271218,Paremphytus:0.02464420258894395):0.02249624460221433,Blasticotoma:0.006932063224623697):0.0490343616179143,(((Tenthredo:0.02287468141834048,Dolerus:0.0379127680071687):0.005511794586039453,Taxonus:0.01534674105965883):0.02796900472957997,Aglaostigma:0.05365685175527035,(Selandria:0.04160919718649558,Strongylogaster:0.02738079464056198,((Hoplocampa:0.02012703819355954,Cladius:0.07366999331734014):0.02195981990479412,(Nematinus:0.01414703684933671,Nematus:0.03212824650204205):0.02514288484299632):0.04326143291250173):0.05138406957157593,(Monophadnoides:0.02663719882434312,Metallus:0.08261967334498921):0.06841739043083328,(Athalia:0.06806068946062127,(Monoctenus:0.009509796412601523,(Gilpinia:0.02289682265065962,Diprion:0.02524587571632375):0.03072393966902143):0.08085945359905478,(((Cimbicinae:0.03112135205100852,Abia:0.02837103898982041):0.05787375139254736,Corynis:0.0631161392906493):0.05048385488182616,((Arge:0.02608787516370437,Sterictiphora:0.05660313417379448):0.03265763329027016,((Perga:0.03797796089770467,Phylacteophaga:0.07075852222815981):0.028701192014013,((Lophyrotoma:0.08164037033593241,Decameria:0.02715056614450995):0.0206934286211462,Acordulecera:0.0425271652672885):0.03713480806565439):0.06082104728862725):0.1179870230549199):0.06841410132902748):0.04003405901194101):0.1260274458423301):0.152695087752944,(((Neurotoma:0.01493344709280175,Onycholyda:0.004328413304073964,(Megalodontes_sp._c:0.0054187309615914,Megalodontes_sp._sk:0.01373851539457993):0.1478917283679591):0.03370702463054354,(Pamphilius:0.009857799443758086,(Cephalcia:0.01169464807293563,Acantholyda:0.01303125175266806):0.01620336514761123):0.01963522568620846):0.1039879153691479,((((Cephus:0.008827771648069471,Calameuta:0.01730711700846832):0.008576575528012387,Hartigia:0.02277087421402169):0.1447796230270977,Syntexis:0.1032857211318408):0.07749108625172862,((Sirex:0.01465686915629738,((Xeris:0.03054615591576326,Tremex:0.04577169571296902):0.00921485126783339,Urocerus:0.01454279438859729):0.006984142011795758):0.1489053004777152,(Xiphydria:0.02465759978621151,(Orussus:0.1588575970572678,(Stephanidae_A:0.1419853322304897,(((Megalyra:0.1263711708470729,Cynipoidea:0.105156189121115):0.07267418225807609,Evanioidea:0.04000417449676341):0.1004827925582878,Trigonalidae:0.1591402971299185,Ichneumonidae:0.1648035596623973,(Apoidea_A:0.150069431285863,Vespidae:0.1267201824265516):0.1476007062145401):0.1287422652990272):0.126212907468366):0.3693937887908082):0.1352591707451611):0.04233480928266231):0.1006975917133421):0.03130975978002692):0.06074484744028275):0.08568107699601413); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr89449] END;