#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on November 09, 2026; 2:50 GMT TreeBASE (cc) 1994-2008 Study reference: Kjer K., Blahnik R., & Holzenthal R. 2001. Phylogeny of Trichoptera (Caddisflies): Characterization of Signal and Noise Within Multiple Datasets. Systematic Biology, 50(6): 781-816. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S852] [ The following blocks are input data for analysis step 1406 ] BEGIN TAXA; TITLE TaxaForAnalysisStep1406; DIMENSIONS NTAX=89; TAXLABELS Agapetus_pinatus Agarodes_distinctus Agathiphaga_queenslandensis Alloecella_grisea Anagapetus_debilis Anisocentropus_bicoloratus Apatania_zonella Apsilochorema_gisbum Archaeopsylla_erinacei Arctopsyche_grandis Atopsyche_sp Atriplectides_dubius Austrheithrus_ronewa Austrotinodes_sp Banksiola_crotchi Beraea_pullata Brachycentrus_americanus Caloca_saneva Catocala_Lymantria Ceraclea_resurgens Cheumatopsyche_oxa Chimarra_feria Chimarra_rossi Chimarrhodella_ulmeri Cochliopsyche_vazquezae Contulma_talamanca Culoptila_n_sp Cyrnellus_fraternus Cyrnus_trimaculatus Dicosmoecus_atripes Diplectrona_modesta Dipseudopsis_varians Dolophilodes_distinctus Ecnomus_tenellus Ernodes_articularis Glossosoma_nigrior Goera_stylata Gumaga_okinawaensis Helicopsyche_borealis Heterobathmia_pseuderiocrania Homoplectra_doringa Hydropsyche_occidentalis Lepidostoma_togatum Leptocerus_americanus Leptonema_salvini Limnephilus_externus Macrostemum_zebratum Merope_tuber Micropterix_calthella Molanna_uniophila Neophylax_concinnus Nyctiophylax_affinis Oeconesus_maori Olinga_feredayi Oxyethira_rossi Paduniella_mahanawana Palaeagapetus_neacticus Pangulia_faziana Panorpa_latipennis Parapsyche_elsis Philanisus_plebeius Philopotamus_montanus Phryganea_cinerea Phryganopsyche_latipennis Phylloicus_aeneus Phylocentropus_placidus Plectrotarsus_tasmanicus Polycentropus_interruptus Protoptila_tenebrosa Pseudoneureclipsis_thuparama Psilotreta_frontalis Psychomyia_pusilla Ptilocolepus_granulatus Pycnocentrodes Rhyacophila_brunnea Rhyacophila_fuscula Smicridea_talamanca Smicridea_turrialbana Stenopsyche_marmorata Stenopsychodes_mjoebergi Sthenopis_purpurascens Taschorema_evansi Tasmanthrus_angustipennis Tinodes_waeneri Trichovespula_macrocera Triplectides_flintorum Wormaldia_gabriella Wormaldia_triangulifera Xiphocentron_n_sp ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M305] TITLE nuclear_RNA; LINK TAXA = TaxaForAnalysisStep1406; DIMENSIONS NCHAR=1078; FORMAT DATATYPE=DNA SYMBOLS= "0 1 2 3 4 5 6 7 8 9 e f i j l o p q t x z ! @ # : % ^ * & ¥" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 370 380 390 400 410 420 430 440 450 460 470 480 490 500 510 520 530 540 550 560 570 580 590 600 610 620 630 640 650 660 670 680 690 700 710 720 730 740 750 760 770 780 790 800 810 820 830 840 850 860 870 880 890 900 910 920 930 940 950 960 970 980 990 1000 1010 1020 1030 1040 1050 1060 1070 1080 1090 1100 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Agapetus_pinatus 22033111120230212212021200012220010211102121320031132122111122--12022112112220003230212331222022-020-01310010303010--01012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3111-03111002023122233213320202-321021113002322232230001311031300221300030130110-120201120302120010100220231302103232121202310-332220????????302011102022121003200023200-0????????????????????????????-22200203212220312?-12311---121121033111202212333120200-0-21-10-1121-21203303122201210132020212301012132220111200020322-32001303211322?013000233233212233000002131230233200033232313321223231223-13011201132122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300233332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211201233-31000-3330011122322210213-11122200011-000213331 Agarodes_distinctus 22033111120230212212021200012220020211102121320031132112111222--12022122112220003230212331203322-020-23300033--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112102022232110221112322320122302112-211212012231-3110-03031102023122233213320202-321021113002022{02}32230001311031300221300030130110-122201120302120010100220231302103232121202310-332220-131-20-300011102022121003200023200-02212202210131210021110120110-22200203212322212?-12131---121121033111202212333120200-0-23-10-1123-21203303120201210132020212301012132220111200020322-32001303211322???3000233233212233000002131230233200033232111321223231223-13011201332122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101312113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31000-3330011122322210213-11122200011-000213331 Agathiphaga_queenslandensis 2203331131023021221202120010220002021110212112003111212233????????-20312112220203232232330222022-331????????????????????313123123-----------?121212-31312311002331231332001222211--213330112302022232130231112302120102212112-212212020220-0111-03113102023122233213320202-321021113002122232230001311031300221300030330112-120201120302120010??????????0210-232321202310-332020????????20001320002-123001200023200--?????????1131??????????1?13?2?220?2032-2122212?-121?1---2131210131?1302212313123312------10-0311-2313--03210?21?12131030210300032132220111200030322-32001303?113223013000033233212233000002131230233210333232121121213231223-{02}{12}-3121?????-12{03}0-0132210{02}23332122221030312312232021?013031203222-----1120100-333013332001000330202321310002122213??????32133200303?13-232103220030030200303203131223313033332--3322333312-2001-1111202030032033003022201001322222103312303321201233020223200033133220312312100201220103102120002103332110002232333310310031002001200023302022331-200221203302030112111302331300110300032103211021102120311211201233-0033--2032213122123310213-3112????????????????? Alloecella_grisea 22033111120230212212021200012220020211102121320031132112111222--12022122112220003230212331223022-020-23000033--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112102022232110221112322320122302112-211212012231-3111-03011102023122233213320202-321021113002122232230001311031300221300030130110-122201120302120010100220231302103212121202310-3322201010-01-200011102022121003200023200-02210202210131210021110220110-22200203212122212--12111---121121033111202212333120200-0-21-10-1123-21203303120201210132020212121012132220111200020322-32001303211322????000233233212233000002131230233200033232311321223231223-13011201332122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Anagapetus_debilis 22033111120230212212021200012220010211102121320031132122111122--12022132112220003230212331222022-020-23300010303010--?1012-121223--103302310012-213-21132311002331031332001222211--213301112302022232110221112322320122302112-211212012231-3131-03111302023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322200201-00-302011102022121003200023200-0212???????1312???????????21?-222002032121222122-12111---121121033111202212333120200-0-21-10-1121-21203303120201210132020212301012132220111200020322-32001303211322???????????????????????????????????????????3-1321223231223-13011201132122301201022101231321222213313121122321231311211221222112332122132-33301333200100033020232131000212221333231332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-?00221203302030112132302331300110300032103211021102120311211201233-31030-3330011122322210213-11122200011-00??13331 Anisocentropus_bicoloratus 22033111120230212212021200012220020211102121320031132112311212--12022122112220003230212331223022-020-23300033--3030--?0012-121223--103302310012-213-21102311002331031332001222211--213301112302022232110221112322320122322112-211212012231-3111-03011102023122233213320202-321021113002322232230001311031300221300030130110-120201120302120010100220231302103232121202310-332220???????-300011102022121003200023200-02212200210131220121310120110222200203212122012?-12311---123121033111202212331120200-0-21-13-1123-21203303122201210132020212301012132220111200020322-32001303211322???????????????????????????????????????323231-321223231223-1301120133212230120122210123132122221331312112232123--11211221222112312122132-33301333200100033020232131000212221300101312113200303323-012103220030030200302201131223313233332--3322133313-2030-111020223003203300302220100132222210331230332120123302022320003313322031231210020122013000212000210333211000223233331033003100200120002330202?331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330013122322210213-11122200011-000213331 Apatania_zonella 22033111120230212212021200012220020211102121320031132122111022--12022112112220003230212331222022-020-233030-0333010--?3012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111302023122233213320202-321021113002322232230001311031300221300030130110-120201120302120010??????????????????121202310-3322201013-20-300011102022120003200023200-02211-222101312000211-1220310-222002032121222122-12111---121121033111202212333120200-0-21-11-1123-21203303120201210132020212301012132220111200020322-32001303211322?????????????????????????????????????033232311321223231223-13011201032122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-11102022300320330030222012013222221033123033202012310202232000331332203123121002012201300021200021033321100022323333103300310020012000233020?2331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Apsilochorema_gisbum 22033111120230212212021200012220120211102121320031132122111122--12022112112220003230212331222022-020-23303000--3030--?0012-121223--103302310012-213-21102311002331031332001222211--213301112302022232110221112322320122302112-211212012231-3131-03111102023122233213320202-321001113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322200031-20-300011102022121003200023200-0????????????????????????????-222002032121222322-12111---121121033131202212333120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-32001303211322?????????????????33000002131230233200033232311321223231223-13011201{01}32122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200203-31030-3330311122322210213-11122200011-000213331 Archaeopsylla_erinacei ??????1131023021221202?20010222002021110210130003111212211????????022011232220003230232330222022-031????????????????????313122321--?-------??12--12-21202331002311031332003222211--03330111030202223211022111230212011233-2121230312020220-3131-03113102023120233213320202-32102131300232223223000131103130022130003032011???????????????????????????????????????????????????????????????00011300022121003200023000--?????????????????????????????222032032-2011210?-32123---211121003111222212313101313------10-33---2120--20020221210111020212301012132220111200020322-320013032113223033000233233212233000002131230233200313232311101013231223-3101121?????3233100132210323132322201231130112232221?013011223222-----1122301233301333200100033020232133000210221310--3312113200303323-232103220030032200302201131223313033332--3322333312-2003-3111202230032033003022201001322222103312303321201233020223200033133220312312100201220102002120002103332110003232333310310031002001200023302023331-200221203102030112111302331300110300012103211021302120311211200233-1131-02032013122122210213-31122200011-000213333 Arctopsyche_grandis 22033111120230212212021200012220020211102121320031132112111122--12022122112220003230212331222022-020333300-33--??301300-12-121223--103302310012-212-2110231100231103133200222221111213303110302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200231-00-3020111020221210032000232001021231022101312300231200102102222002032-2122112--12311---2131-1033111202212333120200-0321-13-1123321203303120201210132020212121012132220111200020322-32001303211322????????????????????????2131230233200033232311321221231223-1301120133212230120102212123102122221331312112232123--11211221222111312122132-33301333200100033020232131000212221330103332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333002332033122322210213-11122200011-000213331 Atopsyche_sp 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Chimarra_rossi 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Chimarrhodella_ulmeri 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Cochliopsyche_vazquezae 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Contulma_talamanca 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Culoptila_n_sp 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Cyrnellus_fraternus 22033111120230213212021200012220220211102121320031132122111222--12022112112220003230212331222022-020333300-33--30103300012-121223--103302310012-212-2110231100231103133200222221111213303112302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002022232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200212-00-30201110202212000320002320010212????????????????????1?21?2222022032-2121212?-12121---213121013131202212333120200-0321-10-1123121203303120201210132020212121012132220111200020322-32001303211322????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????322133313-2010-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31333-2332033122322210213-13122200011-000213331 Cyrnus_trimaculatus 22033111120230212212021200012220010211102121320031132122111222--12022112112220003230212331222022-020323300-33--30103300012-121223--103302310012-212-2110231100231103133200222221111213303112302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-0322200231-21-20201110112212000320002320010212????????????????????1?21?2222022032-2121212--12121---211121013111202212333120200-0321-10-1123321203303120201210132020212121012132220111200020322-32001303211322???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????3313233332--3322133313-2010-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31330-2332033122322210213-11122200011-000213331 Dicosmoecus_atripes 22033111120230212212021200012220020211102121320031132122111122--12022112112220003230212331222022-020-23303010--3010--?0012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111302023122233213320202-321021113002322232230001311031300221300030130110-120201120302120010?????0231302103232121202310-3322201013-20-300011102022120003200023200-02211-222101312000211-1220310-222002032121222122-12111---121121033111202212333120200-0-21-11-1123-21203303120201210132020212301012132220111200020322-32001303211322?????????????????????????????????3200033232311321223231223-1301120103212230120102210123132122221331312112232123131121122122-112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201201322222103312303320201233020223200033133220312312100?????0130002120002103332110002232333310330031002001200023302022331-200221223302030112132302331300110300032?03211021102120311211200233-31030-333001112?32221021??????????????????????? Diplectrona_modesta 22033111120230212212021200012220010211102121320031110112311202--12020122312220003230212331222022-020333300-33--????3300-12-121223--103?02310-12-212-2110231100233103133200022221111213303112302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-332220???????-300011100022121003200023200102123101110-312100222200112102222002032-2122112?-12311---213121033111202212333120200-0321-13-1123321203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321221231223-1301120103212230120102212123132122221331312112232123-211211221222311312122132-33301333200100033020232131000212221300303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-33330-2332033122322210213-11122200011-000213331 Dipseudopsis_varians 2203311112023021221202120001222001021110312112003111212231131?--12022112112220003232212331222222-020333000-33--??301300-12-121223--?????????312-212-2110231100231301133200122221111213303112102022232110221112302120122312112-211212012231-3111313111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3122200213-12-30201112202212100120002320010212???????13121????????1?21?2222002032-2111212--12122---231121013111202212333120200-0321-13-1021321203303120201210132020212101012132220111200020322-320013032111223013000233233212233000002131230233200013232311121221231223-1101123113212232120122212123102122221331312112232123--11211221222111312122132-33301333200100033020232131000212221333103312113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Dolophilodes_distinctus 22033111120230212212021?00012220020211102121320031132122111022--12022112112220003230212331222022-020323300-33--30300300012-121223--103302310012-212-2110231100231103133200222221111213303110302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200013-12-3000111020221210032000232001021231023303330300001202112102222002032-2122312?21231133-211121003111202212333120200-0321-10-1123321203303120201210132020212101012132220111200020322-320013032113223013000233233212233000002131230233200033232311321221231223-13011201332122301201022121231321222213313121122321231-11211221222111312122132-33301333200100033020232131000212221303303332113200303323-21210322003003020030220113?223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333322332033122322210213-11122200011-000213331 Ecnomus_tenellus 22033111120230212212021200012220100211102121320031132122311132--12022112112220003230212330222022-020333320-33--3?300300112-121223--1?301233?212-212-0312233100231103133200122221111213303112302022232110221112302-00122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200013-20-30001110202212100320002320010212???????3312?????????1?21?2222022032-2121212?-12121---211121013111202212333120200-0321-10-1123321203303120001210132020212121012132220111200020322-3200130321132230130002{03}3233212233000002131230233200033232331321221231223-13011201332122301201022121231321220213313121122321230-31211221222111312122132-33301333200100033020232131000212221303103332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-33330-2332033122322210213-11122200{02}11-000213331 Ernodes_articularis 220331110202302122120212000122200202111021213200311321121112202112-22122112220003230212331221022-020-23003033--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112102022232110221112322120122302112-211212012231-3111-03211102023122233213320202-321021113002022232230001311031300221300030110110-122201120302120???100220231302103232121202310-332220???????0300011102022121003200023200-02211202010131220023110220311022200203212122212?-12111---121121033111202212333120200-0-21-10-1123-21202303120201210132020212101012132220111200020322-32001303211322??03000233233212233000002131230233200033232311321223231223-13011201332122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221303101312113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Glossosoma_nigrior 22033111120230212212021200012220000211102121320031132122111122--12022112112220003230212331222022-020-23300010303010--?1012-121223--103302310012-213-21132311002331231332001222211--213301112302022232110221112322320122302112-211212012231-3131-03111002023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322200201-00-301011102022121003200023200-0??????????1312??????????????-222002032121222121-12111---121121033111202212333120200-0-21-10-1121-21203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232313321223231223-1301120113212230120122210123132122221331312112032123-301211221230212330122132-33301333200100033020232131000212221332231332113200303323-012103220030030200302201131223313233332--3322133033-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211201233-31030-3330011122322210213-11122200011-000213331 Goera_stylata 22033111120230212212021200012220020211102121320031132122111122--12022112112220003230212331222022-020-2330--10333010--?-012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322201213-20-300011102022120003200023200-02231-222101312000211-1200310-22200203212122012?-12311---121121033111202212333120200-0-21-11-1123-21201303120201210132020212301012132220111200020322-32001303211322??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Gumaga_okinawaensis 22033111120230212212021200012220020211102121320031132112111222--12022122112220003230212331223322-020-23300030--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112102022232110221112322320122302112-211212012231-3110-03011102023122233213320202-321021113002122232230001311031300221300030130110-122201120302120010100220231302103232121202310-332220???????-300011102022121003200023200-02212202210131210023110120110-22200203212122212?-12111---121121033111202212333120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-320013032113223013000233233212230000002131230203200033232111321223231223-1301120133212230120102210{13}23132322221331312110232{13}03-311211231220-02312120102-33302333200100033020232131000212211300101312133200303323-032103220030030210302001131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Helicopsyche_borealis 22033111120230212212021200012220120211102121320031132112111322--12022122112220003230212331223022-020-23300000103000--?1012-121223--101302310012-213-21122311002331031332001222211--213301112302022232110221112322120122302112-211212012231-3111-03011102023122233213320202-321021113002022232230001311031300221300030120110-122201120302120010????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????30?023323?2122330000?2131230233200013232311321221231223-1321120133212210120102212120132122221331312112232123-311211221222012312122112-33301333200100033020232131000212221300101012113200303333-012103220030032200302201131223313233332--3322133313-233001112202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-20022120330203011213230233130011030003210321102110212031121120023333131023330011122322210213-11122200011-000213331 Heterobathmia_pseuderiocrania ??????????????????????????????????021110210232003111212233????????-20312112220203232232331222023-331????????????????????313123123-----------?121212-31132311002331231332001222211--21333111230202223211022111230232012202?112-212212020220-0331-03113102023122233213320202-3210211130023222322300013110313002213??????????????????????????????100220231302103232321202310-332020????????200013200022123001200023200--22122021201312100121101201301222022032-2022312--12211---313121013111202212313123332------10-0313-2310--03210221212131020210300032132220111200020322-32001303211322301300003323321223300000213120323321033323212112121323122332101121?????1232030132010123332122021030312312232021?013031223222-----1120100-333?133320010003302023213300-2122213??????32133230323130-23210322003003020-303203131223313033332--3322333312-2001-3111202230032033003022203001322222103312303321201233020223200033133220312312100201220103102120002103332110002232333310310031002001200023302022331-20022120330203011211130233130011030003210321102110212031?211203233-0130--2032213122122210{12}{12}3-3112????????????????? Homoplectra_doringa 22033111?20230212212021200012220010211102121320031132312111222--12022122112220003230212331222022-020333300-33--30?00300312-121223--103302310012-212-2110231100231103133200222221111213303112302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322000013-00-300031102022121003200023200102123102???1312?????12?21?21?2222002032-01212120-12121---233121033111202212333120200-0321-13-1023321203303120201210132020212101012132220111200020322-32001303211322??????????????????????????????????????????????321221231223-1301120113212230120102212123132122221331312112232123--11211221222111312122132-33301333200100033020232131000212221300303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-333-0-2332033122322210213-11122200011-000213331 Hydropsyche_occidentalis 22033111120230212212021200012220020211102121320031130312131302--12022122110220003230212331222022-020?03300-03--???30320-12-121221--?????2310-12-212-2110231100233103133200022221113213303112302022232110221112302120122102112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200033-30-30001110202212100330002320010212310-010130--2023-220112102222002032-21221120-12311---211121033111202212333120200-0321-11-1123-21203303120201210132020212121012132220111200020322-32001303211322??????????????????????002131230233200013232311321221231223-112113-033212212120102012123132122221331312112232323-213213021220311312122132-33301333200100033020232131000212221320333312113200303323-212103220030030200302201131223310033332--3322133313-2010-11102022300320330030222010013222221033123033212012330202232000331332203123121002012201300021200021033321100022323333103300310020012000233020223311200221203302030112132302331300110300032103211021102120311211202223-03300-2332003122322210213-11122200011-0002????? Lepidostoma_togatum 22033111120230212212021200012220020211102121320031132122111122--12022112112220003230212331222022-020-23303010--3030--?0012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111302023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322203033-20-300011102022120003200023200-02212-222101312000211-1120310-222002032121222122-12111---121121033111202212333120200-0-21-10-1123-23203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-13011201032122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Leptocerus_americanus ???????????????????????????12220020211102121320031112112211212--12021122112220003230212330223022-020-23300010--1000--?0012-121223--103302310012-011-21102311002331031332001222211--213301112102022232110221112322320122322312-211212032231-3111-03011202023122233213320202-321021113002022232230001311031300221300030120110-120201120302120-10100220231302103232121202310-332220???????-302011102022121001200023200-02212212310131210020110120110-22202203212321212?-12123---121121013111202212333120200-0-23-10-1123-21203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-1321120123202230120102210123132322201331312112232123-311231221220012332122132-33301333200100033020232131000212221300101012113200303323-012103220030032200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201200130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31010{23}3330011122322210213-11122200011-000213331 Leptonema_salvini 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Limnephilus_externus 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Macrostemum_zebratum 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Merope_tuber 220331113302302122120212001022200002111021?132003111212031????????020012132220003230232333022002-031????????????????????311320033-----------?303230-03002333002311011332003222211--0-1321112302022232110221112302-00132330132-031332220220-3131-03113332023122233213320202-321021313002322232230001311031300221300030300110-120201120302120010100220231302103232121002310-3322200303-01-000011300022120003200023000--?????????????????????????????222032032-2233-32?-30300---111121003111222012313103313------10-33---2120--23020221230113020212301012132220111200020322-320013032113223033000233233212233000002131230233200313232311101013231223-0101121?????2213300132210323132322201231130112232223?013031203222-----1122301233301333200100033020232133000210221300033332113200303313-232103220030032200302201131233313033332--3322333312-2002-1311202230032033003022201001322222103312303321201233020223200033133220112312100201230102002120002103332110003232333310310031002001200023302022331-200221203102030112111302331300110300012103211021302120313212101233-1131-02032013122302210213-31122200011-000213333 Micropterix_calthella 2203331133023021221202120012220002021110212112003111212213????????-22112112220203232212331222022-231????????????????????313333121-----------?123312-11312311002331231332001222211--233301112102022232110221112302120102212212-21233202022031311-03111102023122233213320202-321021110002122232230001311031300221300030320112-120201120302120010100220231302103232121202110-332220????????200011101022121003200023200--2211-221211312-11121102201303222012032-2320212?-12331---311121013111222212313323112------10-0313-2312--20132221210112030212300032132230111200020322-320013032113223313020033233212233000002131230233210333232321121013211223-1103121?????2232120132210221332122221030312112232021?013232203222-----11221003333013332001200330202321310002122213??????20300022321032-002103220030032200303203103223333033332--33223333-0-20-2-0111002030032033003220021001322222103312303321201233020223200033133220312312100201220110132120002103332110002232333310310031002001200023302022331-200221203302030112111302331300110300032133213231300120311211201233-0013--3032213122122210213-301?????????????????? Molanna_uniophila ????????????????????????????????020211102121320031112112111212--12022122112220003230212331223022-020-23300033--3000--?0012-121223--103302310012-211-21102311002331031332001222211--213301112102022232110221112322320122322112-211212012231-3111-03011102023122233213320202-321021113002022232230001311031300221300030120110-120201120302120010100220231302103232121202310-332220???????-302011102022121003200023200-02222202210130210021310110110-22200203212122212--12111---121121033111202212331120200-0-21-10-1123-21201303122001210132020212301012132220111200020322-32001303211322????????????????????????????????????????????1?321?23231223-13011201332122301201222101231321222213313121122321231311211221220112312122132-33301333200100033020232131000212221300101312113200303323-012103220030030200302201131223313233{03}32--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Neophylax_concinnus 22033111120230212212021200012220020211102121320031132122111122--12022112112220003230212331222022-020-23303030--3010--?0012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111302023122233213320202-321021113002322232230001311031300221300030130110-1202011203{02}2120010??0220231302103232121202310-3322201013-20-300011102022120003200023200-02211-222101312000211-1220310-222002032121222122-12111---121121033111202212333120200-0-21-11-1123-21203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-13011201032122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-11102022300320330030222010013222221033123033212012330202232000331{13}32203123121002012201300021{12}0002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330013122322210213-11122???????????????? Nyctiophylax_affinis 22033111120230212212021200012220020211102121320031132122111222--12022112112220003230212331222022-020333300-33--3?303300{13}12-121223--103302310012-212-2110231100231103133200222221111213303112102022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-132220???????-30101110122212100320002320030212???????1312?????????1?21?2222022032-2121312?-12121---211121013111202212333120200-0321-10-3123121203303120201210132020212121012132220111200020322-320013032113223013000233233212233000002131230233200033232311321221231223-13011221332122301201022121231321222213313121122321231-11211221222111312122132-33301333200100033020232131000212221300303312113200303323-212103220030030200302201131223313233332--??22133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31330-2332033122322210213-11122200011-000213331 Oeconesus_maori 22033111120230212212021200012220020211102121320031132122111022--12022112112220003230212331222022-020-23303000--3030--?0012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010????????1302103232121202310-3322201203-13-300011102022120003200023200-02211-222101312100211-1220310-222002032121222122-12111---121121033111202212333120200-0-21-10-1123-23203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-?3011201032122301201222101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-33300111222?2210213-11??????????????????? Olinga_feredayi ??????????????212212021200012220020211102121320031132112111222--12022122112220003230212331223022-020-23000033--3000--?0012-121223--103302310012-213-21102311002331331332001222211--213321112102022232110221112322320122302112-211212012231-3111-03011102023122233213320202-321021113002122232230001311031300221300030130110-12220??20302120010100220231302103232121202310-3322201212-20-300011102022121003200023200-02231202210131210021110220110-22200203212122312--12011---121121033111202212333120200-0-21-11-1121-21201303120201210132020212101012132220111200020322-3200?303211322????????????????????????????????33200033232311321223231223-1301120133212230120102210123132122221331312112232123--11211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Oxyethira_rossi 22033111120230212212021200012220020211103121320031132120312322--12020312112220003230212313222022-020-23300033--3000--?0012-121223--103302310012-233-2110231100233103133200122221111210301110102022232110221112322-20122302112-211212012231-3131-03111202023122233213320202-321021113002122232230001311031300221300030110110-120201120302120010100220231302103232121202310-332220????????300011102022120003200023200-0231210-2-01332-101--20120210-22200203212321201?012101----23121033111202212333120200-0-23-10-1123-21203303120001210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311121221231123-1201120133212230120322212123132122201331312112232123-311211121220032312122133-33301333200100033020232131000212221323333312113200303323-012103220030030200302201131223313233332--3322133313-2010-1112202??0032033003022201001322222103312303321201233020223200033133220312312100201200130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200230-313331-033012122322210213-1112????????????????? Paduniella_mahanawana 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Palaeagapetus_neacticus 22033111030230212212021200012220020211103121320031132122111122--12022112112220003230212330222022-020-23303033303010--01012-121223--103302310012-213-2113231100233103133200122221311213321112102022232110221112322320122302112-211212012231-3131-03111002023122233213320202-321021113002322232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322200031-01-300011102022121003200023200-0?????????????????????????210-222002032123232122-12301---121121033111202212333120200-0-21-11-1123-23303303120201210132020212301012132220111200020322-32001303211322??13000233233212233000002131230233200033232311321223231223-1301120113212230120102210123132122221331312112232123-311211221222112312122132-33301333200100033020232131000212221300233312113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201002322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200033-01010-3330331122322210213-11122200011-000213331 Pangulia_faziana 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Panorpa_latipennis ??????????????????????????????????021110210132003111130211????????022030232220003230232303022022-033????????????????????311320203-----------?103230-21212333002311031332003222211--003301110302022232110221112300-00132330132-031332220220-3131-03113332023122233213320202-32102131300232223223000131103130022130003030011????????????????????101220231302103232121002310-3322200133-00-300011300022123003200023000--?????????????????????????????222002032-3030031?-20332---301121033111022012313103033------10-33---2120--03020221230110020212301012132220111200020322-320013032113223033000233233212233000002131203233200333232311103013231223-3101121?????323330013221032-130322201231130312232223?031031223222-----1120301233301333200100033020232133000210221300-31312113230303323-232103220030030200302201131233313033332--3322333312-2003-3111202230032033003022201001322222103312303321201233020223200033133220112312100201220102002120002103332110003232333310310031002001200023302022331-200221203102030112111302331300110300012103211021302120311211000233-1131-02032013322122210213-31122200011-000213333 Parapsyche_elsis 22033111120230212212021200012220120211102121320031132112111122--12022122112220003230212331222022-020333300-33--??301300-12-121223--103302310012-212-2110231100231103133200222221111213303112302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200231-00-3020111020221210032000232001021231022101312300211200102102222002032-2122112--12311---213121033111202212333120200-0321-13-1123321203303120201210132020212121012132220111200020322-32001303211322???????233233212233000002131230233200033232311321221231223-1301120133212230120102212123102122221331312112232123--11211221222111312122132-33301333200100033020232131000212221330103332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331--00221203302030112132302331300110300032103211021102120311211200233-3333002332033122322210213-11122200011-000213331 Philanisus_plebeius 22033111120230212212021200012220020211102121320031132112111222--12022122112220003230212331223022-020-20300003--3000--?0012-121223--103302310012-213-2112231100233103?332001222211--213321112102022232110221112322320122302112-211212012231-3111-03011102023122233213320202-321021113002122232230001311031300221300030130110-12220112030212001010022023{01}302103232121202310-332220???????1300011102022121003200023200-02212102211131210021120211110-22200203212122212?-12111---121121033111202212333120200-0-21-10-1121-21203303120201210132020212101012132220111200020322-32001303211322????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1??????????????????????????-??????????????????????????????????????????????? Philopotamus_montanus 22033111120230212212021200012220020211102121320031132122111022--12022112112220003230212331222022-020323300-33--30300300012-121223--103302310012-212-2110231100231103133200222221111213303110302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200013-00-30001110202212100320002320010212????????????????????1?21?2222002032-2122012?21231131-211121003111202212333120200-0321-10-1123321203303120201210132020212101012132220111200020322-32001303211322?????????????????????????????????3200??3232311321221231223-13011201332122301201022121231321222213313121122321231-11211221222111312122132-33301333200100033020232131000212221303303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333322332033122322210213-11122200011-000213331 Phryganea_cinerea ???????????????????????????12220020211102121320031132122111122--12022112112220003230212331222022-020-23303010--3010--?0012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111302023122233213320202-3210211130023222322300013110313002213??????????????????????????????100220231302103232121202310-3322201233-20-300011102022120003200023200-02211-222101312000211-1220310-222002032121220122-12311---121121033111202212333120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-1301120103212230120102210123132122221331312112232123131121122122-1123?21?2132-3330133320010003302023213100021222130010133{02}113200303323-01{02}103220030030200302201131223313233332--3322133313-2030-11102022300320330030222010013222221033123033212012330202232000331332203123121002012?0130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-111?????????????????? Phryganopsyche_latipennis 22033111120230212212021200012220020211102121320031132122131022--12002112112220003230212331222022-020-23003030--3010--?0012-121223--103302310012-213-211323110023310313320012222111-203301112102022232110221112322320122302112-211212012231-3131-03111302023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322201013-20-300011102022120003200023200-02231-220103312000331-1220110-222002032121220122-12111---121121033111202212333120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-32001303211322?????????????????????????????????3200033232311321223231223-13011201332122301201022101231321222213313121122321231311213021222112312122132-33301333200100033020232131000212221300301332113200303323-012103220030030200302201131223313233332--3322133313-2030-111020223003203300302220100132222210331230332120123302022320003313322031{12}312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-13122200011-????????? Phylloicus_aeneus 22033111120230212212021200012220020211102121320031112112111212--12022122112220003230212331223022-020-23300033--3000--?0012-121223--103302310012-211-21102311002331031332001222211--213301112302022232110221112322320122322112-211212012231-3111-03011102023122233213320202-321021113002322232230001311031300221300030120110-1?0201120302120010100220231302103232121202310-332220???????-302011102022121003200023200-02012200210131220121010120310222200203212122012--12311---121121033111202212331120200-0-23-10-1123-21203303122201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-13011201332122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221303101312113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-21030-3130013122322210213-11122200011-000213331 Phylocentropus_placidus 22033111120230212212021200012220010211102123120031132122111122--12022112112220003232012331222022-020303300-33--??303300-12-121223--103302310012-212-2100231100231103133200222221111213303112302022232110221112302220122312112-211212012231-3131-03111102023122233213320202-321021113002322232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200033-30-30001110202212000320002320010212???????13121????????1?21?2222002032-2120312?-12321---211121033111202212333120200-0321-10-1123321203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321221231223-1301123133212230120102212123132122221331312112232123--11211021220111312122132-33301333200100033020232131000212221330303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-33330-2332033122322210213-11122200011-000213331 Plectrotarsus_tasmanicus ??????????????????????????????????0211102121320031132122111122--12022112112220003230212331222022-020-23303010--3010--?0012-121223--103302310012-213-211323110023310313320012222111-213301112102022232110221112322320122302112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-332220-312-10-300211102022123003200023200-02211-222101312000211-1220110-222002032121222122-12111---121121033111202212333120200-0-21-13-1123-21203303120201210132020212301012132220111200020322-32001303211322????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330013122322210213-11122200011-000213331 Polycentropus_interruptus 22033111120230212212021200012220012211102121320031132122111222--12022112112220003230212331222022-020323300-33--30303300012-121223--103302310012-212-2110231100231103133200222221111213303112102022232110221112322320122312112-211212012231-3111-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-312220???????-30201112002212100320002320010212????????????????????1?21?2222002032-2022212?012311---311121033111202212333120200-0321-10-3121121203303120201210132020212101012132220111200020322-320013032111223013000233233212233000002131230233200033232311321221231223-13011221332122301201022121231321222213313121122321231-11211221222111312122132-33301333200100033020232131000212221300303332113200303323-212103220030030200302201131223313233332--3322131313-2030-11122022300320330030222010013222221033123033212012330202232000331332203123121002012201300021200021033321100022323333103300310020{02}1200023302022331-200221203302030112132302331300110300032103211021102120311211200233-1133-??33{02}033122322210213-?1122200011-000213331 Protoptila_tenebrosa 22033111330230212212021200012220130211103123120031132122111122--12022112112220003232012310222022-020-233000033-3000--02012-121323--103302310012-311-21112311002331031332001222211--213321112102022232110221112302120122322012-210212012231-3131-03111202023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-132220????????302011101222121003200023200-1????????????????????????????-22202203212132212?-12112---121121033111202212333120200-0-21210-1121?21201303122201210132120212301012132220111200020322-32001303211322??????0233233212233000002131230233200033232113321221231223-13011201132122301201022121231021222213313121122321233311211221222112312122132-33301333200100033020232131000212221310331312113200303323-012103220030032200302201131223313233332--3322133313-2030-1112202230032033003022201201322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211231233331031-3030011122322210213-11122200011-000213331 Pseudoneureclipsis_thuparama 22033111120230?12212021200012220010211102121320031132122113100--12022112112220003230212330222022-02033330?-??--?0103300212-121223--?03302310112-212-2100231100231103133200222221111213303112302022232110221112302-20122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120??2120010100220231302103232121202310-3322200033-02-300011102022121003200023??????????????????????????????????222002032-0312012?313312---33112103311120221233312000010121-31-0121-23303303120001210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311121221231223-13011201312122301201022121231321222213313121122321231-31211221200131312122133-03301333200100033020232131000212221300303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-33330-2332033122322210213-11122200011-000213331 Psilotreta_frontalis 22033111120230212212021200012220020211102121320031132112111222--12022122112220003230212331223022-020-23300010--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112102022232110221112322320122302112-211212012231-3111-03011102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-032220???????-300011102222121003200023200-02231201210130210021210230110-22200203212122012?-12311---121121033111202212331120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-13011201032122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201201322222103312303320201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322-102{13}3?????????????????????? Psychomyia_pusilla 22033111120230212212021200012220020211102121320031132122111122--12022112112220003230212330222022-020?23333-03--?0303320{13}12-121223--103302310012-212-2110231100231103133200222221111213303110302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200033-10-3000111020221210032000232001023211023303312-10201202112102222002032-2120012?-12131---231121033111202212333120200-0321-10-3123321203303120201210132020212101012132220111200020322-32001303211322???????????????????????????????????????????311?21221231223-13011201332122301201022121231321222213313121122321231-11211221222111312122132-33301333200100033020232131000212221301303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312302100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333322332033122322210213-11122200011-000213331 Ptilocolepus_granulatus ?203-111-2023021221202120001222002021110?121320031132122311322--12022112112220003230112331222022-020-2330-033303010--01012-121223--10330?310012-21?-211{13}2?110023310313320{02}?222213112133{02}11121020222321102211123223201223?2112-213212012231-3131-0311130202312223321332{01}002-3210011130023222322{13}00013110{23}13002213000301?0110-120{23}01120300120010100220231302103232121202310-3320200011-20-300013102022121003200023200-0212???????1312???????????210-22200203212120212?032131---121121033111202212333120200-0-21-13-1123-21203303120001210132020212301012132220111200020322-32001303211322???????????????????33{03}002131230233200033232311321223231223-13011201132122301201022101231021222213313121122321231331211221223112312122132-33301333200100033020232131000212221300330332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3030011122322210213-11122200011-000213331 Pycnocentrodes 22033111120230212212021200012220020211102121320031132112111222--12022122112220003230212331223022-020-23000033--3000--?0012-121223--103302310012-213-21102311002331331332001222211--213321112102022232110221112322320122302112-211212012231-3111-03011102023122233213320202-321021113002022232230001311031300221300030130110-122201120302120010100220231302103232121202310-3322201010-20-300011102022121003200023200-02231202210131210021110230110-22200203212122312--12011---121121033111202212333120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-32001303211322???????????????????????????????203200033232311321223231223-13011201332122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--332213331332030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Rhyacophila_brunnea 22033111120230212212021200012220020211102121320031132122111122--12022112112220003230212331222022-020-03303010--3010--?0012-121223--103302310012-213-21102311002331031332001222211--233301112302022232110221112322320122302112-211212012231-3131-03111102023122233213320202-321021113002122232230201311031300221300030130110-120201120302120010100220231302103232121202310-332220???????0300011102022121003200023200-022121022101312-1023120120110-22200203212122011?-22311---121121033111202212333120000-0-21-10-1123323303303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200033232311321223231223-13011201132122301201022101231321222213313121122321233331211221222112312122132-33301333200100033020232131000212221300233332113200303323-012103220030030200302201131223313233332--33221333133203031110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31033-3330011122322210213-11122200011-000213331 Rhyacophila_fuscula ?????????????????????????????????20211102121320031132122111122--12022112112220003230212331222022-020-01303010--3010--00012-121223--103302310012-213-21102311002331031332001222211--233301112302022232110221112322320122302112-211212012231-3131-03111102023122233213320202-321021113002122232230201311031300221-?????????????????????????????????220231302103232121202310-332220???????0300011102022121003200023200-022121022101312-1023120120110-222002032121220122-12111---121121033111202212333120200-0-21-10-112132330330312020121013202021230101213222011120002032--3200130321132?????000233233212233000002131230233200033232311321223231223-13011201132122301201022101?313212222133131211223212????????2212221123121????2-33301333200100033020232131000212221300233332113200303-23-010103220030030200302201131223313233332--3322133303-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200203-31033-3330311122322210213-1112????????????????? Smicridea_talamanca 22033111120230212212021200012220020211102121320031112112311202--02022122112220003230212331222022-020323300-33--????3300-12-121223--?????????-12-212-2110231100231103133200022221111213303112302022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103212121202310-122220???????02000111020221210012000232001021231020101312100231202112102222002012-2122312?-12211---211121033111222212333120200-0321-13-1123-21203303120201210112020212121012132220111200020322-320013032113223013000233233212233000002131230233200223232311321221231223-1321120103212232120122212123102122201331312112232323-221221211210311312122132-33301333200100033020232131000212221300101312113200303323-212103220030032200302201131223310033332--3322133313-2010-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-333-0-2332003122322210213-11122200011-000213331 Smicridea_turrialbana 22033111120230212212021200012220020211102121320031112112311202--12020122112220003230212331222022-020323300-33--??303300-12-101223--100010?10-12-212-2110231100233103133200020221111213303112302022232110221112302120122312112-211232012231-3131-03111102023122233213320202-321021113002322232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200033-30-3000111020221210032000232001021231020101312100231202112102222002032-01223121-12311---233121033111222212333120200-0321-13-1123321203303120201210112020212121012132220111200020322-32001303211322???3000233233212233000002131230233200033232311321221231223-1321120103212230120122212123102122221331312112232121-213311222220311312122132-33301333200100033020232131000212221300101332113200303323-212103220030030200302201131223313233332--3322133313-2010-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-33330-2332033122322210213-11122200011-000213331 Stenopsyche_marmorata 22033111120230212212021200012220100211102121320031132122111022--12022112112220003230212330222022-020323300-33--30?03300312-121223--103302310012-212-2110231100231103133200222221111213303112302022232110221112322320122312112-211212032231-3131-03111102023122233213320202-321021113002322232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200013-00-3000111020221210032000232001021211020101302100231202112102222002032-21221120-12311---211121033111202212333120200-0321-10-3123321203303120201210132020212101012132220111200020322-320013032113223013000233233212233000002131230233200033232311321221231223-1301120133212230120102212123132122221331312112232123--11211221222111312122132-33301333200100033020232131000212221303303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333322332033122322210213-11122200011-000213331 Stenopsychodes_mjoebergi 22033111120230212212021200012220020211102121320031132122111122--12?--112112220003230212331222022-020323300-33--30133300012-121223--103302310012-212-2110231100231103133200222221111213303112102022232110221112322320122312112-211212012231-3131-03111102023122233213320202-321021113002322232230001311031300221300030120110-120201120302120010100220231302103212121202310-3322200213-02-3020111020221210032000232001021211020101102100231202112202222002032-2122112?-12311---211121033111202212333120200-0321-10-3123321201303120201210132020212121012132220111200020322-3200130321132230130002{13}3233212233000002131230233200033232311321221231223-1301120133212230120102212123132122221331312112232123--11211221222111312122132-33301333200100033020232131000212221300303332113200303323-212103220030030200302201131223313233332--3322??????-????-???0202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333322332033122322210213-111222000111000213331 Sthenopis_purpurascens 2203331133023021221202120010220002021110210132003111312231????????-22312012220203232232331222022-331????????????????????313321223-----------?121210-31132311002331231332001222211--103331112302022232110221112302120122012112-211212020320-0331-03113102023122233213320202-321021113002322232230001311031300221300030320112-120211120302120010100220{12}333021032323212023100332020????????302013300022123{03}01200023200--22121021201312100121201201301222002032-2022112--12211---313121013111202212313123332------10-0311-2310--032302{02}121{02}131020210102032132220111200020322-32001303211322?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1021321303????????????????????????????????????????????????????????????????????????????201131223313233332--3322333310-2001-3111202230032033003022203001322222103312303321201233020223200033133220312312100201200103102120022103332110002232333310310031002001200023102022331-20022120330203011211130233132011030003-003231203302120311211203233-0130--2032213122122212213-31122200011-000033333 Taschorema_evansi 22033111120230212212021200012220120211102121320031132122111122--12022112112220003230212330222022-020-23303000--3030--00012-121223--103302310012-013-21102311002331031332001222211--213301112102022232110221112322320122302312-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322200231-01-302011102022121003200023200-0????????????????????????????-22200203212122212?-12111---121121033131202212333120200-0-21-10-1123-21203303120201210132020212101012132220111200020322-32001303211322????300233233212233000002131230233200033232311321223231223-13011201132122301201022101231321222213313121122321231311211221222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Tasmanthrus_angustipennis 22033111120230212212021200012220020211102121320031132112111202--12022122112220003230212331223022-020-23300003--3000--?0012-121223--103302310012-211-211023110023310313320012222111-013301112102022232110221112322320122322112-211212012231-3111-03011102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-332220???????0300011102022121003200023200-02221201210131210021210210110-222002032121222120-12111---121121033111202212333120200-0-21-10-1121-21201303120201210132020212301012132220111200020322-32001303211322?????????????????????????????????3200033232313321223231223-13011231002122301201022101231321222213313121122321233213211221222112312122132-33301333200100033020232131000212221300101312113200303323-012103220030030200302201131223310033332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-2330013122322210213-11122200011-000213331 Tinodes_waeneri ???33111120230212212021200012220020211102121320031132122111122--12022112112220003230212330222022-020?23300-03--?3303320{13}12-121223--103302310012-212-2110231100231103133200222221111213303112302022232110221112322320122312112-211212012-31-3131-03111-------22233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200013-00-3000111020221210032000232001021211020303312-10231202112102222002032-2111012?-12122---211121033111202212333120200-0321-10-3123321203303120201210132020212101012132220111200020322-320013032113223013000233233212233000002131230233200033232311121221231223-13011201332122301201022121231021222213313121122321230-11211221222111312122132-33301333200100033020232131000212221300303332113200303323-212103220030030200302201131223310033332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312302100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211220033-0333322332033122322210213-11122200011-000213331 Trichovespula_macrocera ?2033111120230212212021200012220020211102121320031132112111322--12022122112220003230212331223022-020-23301033--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112102022232110221112322320122302112-211212012231-3111-03011102023122233213320202-321021113002122232230001311031300221300030130110-120201120302120010100220231302103232121202310-3322201010-01-300011102022121003200023200-02231201210130210021210230110-22200203212122012?-12311---121121033111202212333120200-0-21-10-1123-21203303120201210132020212301012132220111200020322-32001303211322???????????????????????????????????????????????????????????????????????????????????????????????????312112232123131121122-222112312122132-33301333200100033020232131000212221300101332113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030-3330011122322210213-11122200011-000213331 Triplectides_flintorum 22033111120230212212021200012220020211102121320031112112311212--12020122112220003230212330223022-020-23300033--3000--?0012-121223--103302310012-213-21102311002331031332001222211--213301112202022232110221112322320122322112-211212012231-3111-03011202023122233213320202-3210211130021222322300013110313002213{02}0030130110-120201120302120010100220231302103232121202310-332220???????-300011102022120003200023200-02212203121131120121010120110-22200203012121212--12121---123121033111202212313120200-0-23-10-1123-21203303120201210132020212301012132220111200020322-32001303211322??????????????????????????????????????????????????23231123-1?{02}11201332122301201022101231?2122221331312112232123-311211221220012332122132-33301333200100033020232131000212221300101312113200303323-012103220030030200302201131223313233332--3322133313-2030-1110202230032033003022201021322222103312303321201233020223200033133220312312100201220130002120002103332110002-32333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-31030{23}3330011122322210213-11122-00011-0002133-1 Wormaldia_gabriella 22033111120230212212021200013220010211102121320031132120311002--12022312112220003230212331222022-020333300-33--30300300012-121223--?????????-12-212-2110231100233103133200022221111213303112302022232110221112302320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200013-12-3000111020221210032000232001021212002103312300210102112102222002032-2120012?212311-3-211121013111202212333120200-0321-11-1123-21203303120201210132020212101012132220111200020322-320013032113223013000233233212233000002131230233200033232331121221231323-132113-0332122301201222121231321222033313121122321230-11211021222111312122132-33301333200100033020232131000212221330303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1112202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032133211021102120311211230233-31330-2332013122322210213-11122200011-000213331 Wormaldia_triangulifera 22033111120230212212021200012220010211102121320031132120311002--12022312112220003230212331222022-020333300-33--??300300-12-121223--?????????-12-212-2110231100233103133200022221111213303112302022-32110221112302320122312112-211212012231-3131-03111102023122233213320202-321021113002122232230001311031300221300030120110-120201120302120010100220231302103232121202310-332220???????-30001110202212100320002320010212???????1312?????????1?21?2222002032-2122012?212311-1-211121013111202212333120200-0321-11-1123-21203303120201210132020212101012132220111200020322-32001303211322??????????????????????00213123023320{01}033232331121221231323-132113-0332122301201222121231321222033313121122321230-11211221222111312122132-33301333200100033020232131000212221330303332113200303323-212103220030030200302201131223313233332--3322133313-2030-1112202230032033003022201001322222103312303321201233020223-00033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032133211021102120311211230233-31310-3332013122322210213-11122200?11-000213??? Xiphocentron_n_sp 22033111120230212212021200012220020211102121320031130122311302--12022112312220003230212331222022-020?23300-03--?3303320{13}12-121223--?01102310112-212-2110231100231103133200222221111213303112302022232110221112302120122312112-211212012231-3131-03111102023122233213320202-321021113002322232230001311031300221300030120110-120201120302120010100220231302103232121202310-3322200233-10-3000111000221210032000232001021211022301312100211202112122222022032-21120121-12312---213121013131202212333120200-0321-11-3123321203303120201210132020212301012132220111200020322-320013032113223013000233233212233000002131230233200023232311321221231223-1301120133212230320102212123132122221331312112232123--31213221222111312122132-33301333200100033020232131000212221300303332113200303323-212103220030030200302201131223313233332--3332133313-2030-1112202230032033003022201001322222103312303321201233020223200033133220312312100201220130002120002103332110002232333310330031002001200023302022331-200221203302030112132302331300110300032103211021102120311211200233-3333322332033122322210213-11122200011-000213331 ; END; BEGIN CODONS; CODONPOSSET * CodonPositions (CHARACTERS = nuclear_RNA) = N: 1-1078; CODONPOSSET CodonPositions (CHARACTERS = nuclear_RNA) = N: 1-1078; END; [ The following blocks are output data for analysis step 1406 ] BEGIN TREES; TITLE Tb6182; LINK TAXA = TaxaForAnalysisStep1406; TREE Fig._5 = [&R] (Archaeopsylla_erinacei,(Panorpa_latipennis,Merope_tuber)Mecoptera,(((((Catocala_Lymantria,Sthenopis_purpurascens),Heterobathmia_pseuderiocrania),Agathiphaga_queenslandensis),Micropterix_calthella)Lepidoptera,(((((((Ptilocolepus_granulatus,Oxyethira_rossi),Palaeagapetus_neacticus),(Glossosoma_nigrior,Anagapetus_debilis),Agapetus_pinatus),(Rhyacophila_fuscula,Rhyacophila_brunnea)),((((Protoptila_tenebrosa,Culoptila_n_sp),Atopsyche_sp),Taschorema_evansi),Apsilochorema_gisbum)),(((Brachycentrus_americanus,(((Apatania_zonella,Limnephilus_externus,Dicosmoecus_atripes),((Banksiola_crotchi,Phryganea_cinerea),(Lepidostoma_togatum,Oeconesus_maori))),Neophylax_concinnus)),((Goera_stylata,Pangulia_faziana),Plectrotarsus_tasmanicus)),Phryganopsyche_latipennis),(Alloecella_grisea,(Agarodes_distinctus,Gumaga_okinawaensis),(Contulma_talamanca,Philanisus_plebeius),(Beraea_pullata,Ernodes_articularis),(Cochliopsyche_vazquezae,Helicopsyche_borealis),(Pycnocentrodes,Olinga_feredayi),Caloca_saneva,((Trichovespula_macrocera,Psilotreta_frontalis),(Atriplectides_dubius,(Tasmanthrus_angustipennis,Austrheithrus_ronewa))),((Leptocerus_americanus,Triplectides_flintorum,Ceraclea_resurgens),Molanna_uniophila,(Anisocentropus_bicoloratus,Phylloicus_aeneus)))),(((((((((Diplectrona_modesta,(Smicridea_talamanca,Smicridea_turrialbana)),(Hydropsyche_occidentalis,Cheumatopsyche_oxa)),(Parapsyche_elsis,Arctopsyche_grandis)),(Homoplectra_doringa,(Leptonema_salvini,Macrostemum_zebratum))),((Dolophilodes_distinctus,Philopotamus_montanus),(Chimarrhodella_ulmeri,(Chimarra_feria,Chimarra_rossi)))),(Stenopsychodes_mjoebergi,Stenopsyche_marmorata)),(((Tinodes_waeneri,Psychomyia_pusilla),Paduniella_mahanawana),(Pseudoneureclipsis_thuparama,Xiphocentron_n_sp))),(((Wormaldia_gabriella,Wormaldia_triangulifera),Dipseudopsis_varians),Phylocentropus_placidus,(Austrotinodes_sp,Ecnomus_tenellus))),(Polycentropus_interruptus,((Cyrnus_trimaculatus,Cyrnellus_fraternus),Nyctiophylax_affinis))))Trichoptera)); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5683] END;