#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on November 10, 2026; 8:41 GMT TreeBASE (cc) 1994-2008 Study reference: Donoghue M.J., Eriksson T., Reeves P., & Olmstead R. 2001. Phylogeny and Phylogenetic Taxonomy of Dipsacales, with special reference to Sinadoxa and Tetradoxa (Adoxaceae). Harvard Papers in Botany, 6(2): 459-479. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S981] [ The following blocks are input data for analysis step 1683 ] BEGIN TAXA; TITLE TaxaForAnalysisStep1683; DIMENSIONS NTAX=15; TAXLABELS Adoxa_moschatellina Diervilla_sessilifolia Sambucus_australasica Sambucus_australis Sambucus_caerulea Sambucus_ebulus Sambucus_racemosa Sinadoxa_corydalifolia Tetradoxa_omeiensis Viburnum_acerifolium Viburnum_dentatum Viburnum_lentago Viburnum_plicatum Viburnum_sieboldii Weigela_hortensis ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M524] TITLE ITS; LINK TAXA = TaxaForAnalysisStep1683; DIMENSIONS NCHAR=679; FORMAT DATATYPE=DNA SYMBOLS= "0 1 G T" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 370 380 390 400 410 420 430 440 450 460 470 480 490 500 510 520 530 540 550 560 570 580 590 600 610 620 630 640 650 660 670 680 690 700] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .] Adoxa_moschatellina T1G000-11TG10T0G10G00TG0111GTG001TTGTTTTTT?T1T1GTGG1T1GT1GG01T0GG10TG100GTG-11T1G0T1G0100G1-TT1GGT10GGG1G110T1--GGTGT10TG0110001001G001111GG{1G}01GGT1TGTG1100GG000TTT0T-0T000G0G1GTGT1T0TTGTTG1111GTT1G1GGTG-G101G1---T0GG10TG-1G11TTTG000101T0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT{0T}G1G000TG1G0T0TTTGGTGTG00TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G110TT0GG1100GGG101GT1TG11TGGG1GT101010TTG11GT1G111110TT1100-1T111TTTG{1T}TTGGG-0--GT-TTGTTGT---------------------------0G--GGG1GG000TTGG11T111GTG1T1T1TGGTG1GGTTGG111000TG0G0GT0111G01001GT01GT10100100GTGGTGGTTG000-0G11GT0T-0T10TGTTGTG10110GT11GTTGT10-1GGG101TG01TTG0111T00-0G1GT1GT1TTTG0TGT1G1T11G01T000110111010000011011 Diervilla_sessilifolia T1G000-11TG1010G10G001G0111G1G00101GTT1GT01011--GGG0-0G11GGT11GGG1G1GT10G--10111GG11GG1G1T1110TGGT1GGGG0G11110--GG1T1111G0T1G000-11G001111GG1G1G0T1TG1G1100GG00TT0100010000GGG11TG11T111GTTG1111GT11G1GGTG1G1G1GGGTGT0G-1T1G11T1TTT1G000101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G1101T0GG11G0GGG101GT1TG11TGGG1GT101G10T1G1-GT1G1111----1100---111------------------1G-1GT111-010GGGT1G1G1G1GG1GGGGGG0G-----1GG0G00TGG11T111GTG111T1GGG1G1GG1TGG11100001TG0GT1111GG1GG1GG01GT101G0100GTGGTGGT1G000G0G111T1TT0T101GT1GTG1GGTT1111GT1G1--T1GGG1GG1100GTG0111TG01G1GT1GT1GTT1G01GG1G1T11G011111011000100100100100 Sambucus_australasica T1G000-11TG1010G10G00TG0111G1G001T1GTTTT10T0TT-GGGG1T1GT1GG11T0GG1G1G100G1G--1T1GGT1GG1GG01-TT1GGT10GGG1G111T1--GGTGTG1TG0110001001G001111GG1G1G0T11G1G1100GG00TTTTT0-11G00G0G1GTGT1TT11GTTG{1GT}111GTT1G1GG1GTG101GG---T0GG111G-1G11TTTG000101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTT{GT}G001G100GTTG1G111G00G110TT0GG11G0GGG101GT1TG11TGGG1GT101010TTG11GT1G11111-TT1100TTT1110TT-1TTGGTG0-1GT--TG--GT---------------------------GGT1GGG1GG0T0TTGG11T111GTG1T1T1G0G1G1GGTTGG1110000G1G0GT1111G01001GG01GT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG101100TT10TTGT11-1GGG10T1G0GTTG0111TG0-1G1GT1GT1T11G01GT1G1T11G0T1100110101010100010011 Sambucus_australis T1G000-11TG1010G10G00TG0111G1G001T1GTTTT10T0TT-GGGG1T1GT1GG11T0{0G}GTG1G100G1G--1T1G0T1G01GT01-1T1GGT10GGG1G111T1--GGTGTG1TG0110001001G001111GG1G1G0T1TG1G1100GG00TTTTT0-1TG00G0G1GTGT1T0T1GTTG1111GTT1G1GGTGTG101GG---T0GG11T0-1G11TTTG000101-0001G01T1T1GG1001GG0T0T1T1GG{1G}T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTT{GT}G0?1G100GTTG1G111G00G110TT0GG11G0GGG{01}01GT1TG11TGGG1GT101010TTG11GT1G11111?TT1100TTT1110TT-1TTGGGG0-1GT--TG--GT---------------------------0GT1GGG1GG0T0TTGGT1T111GTG1T1T1G0G1G1GGTTGG1110000G1G0GT1111G01001GG0TGT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG101100TTTGTTGTT0-11GG10T1G0GTTG0111TG0-1G1GT1GT1T1TG01GT1G1T11G0T1100110101010100010011 Sambucus_caerulea T1G000-11TG1010G10G00TG0111G1G001T1GTTTT10T0TT-GGGG1T1GT1GG11T0GGTG1G100GTG--1T1GGT1GG1GG01-TT1GGT10GGG10111T1--GGTGTG1TG0110001001G001111GG1G1G0T1TG1G1100GG00TTTTT0-1TG00G0G1GTGT1T0T1GTTG1111GTT1G1GGTGTG101GG---T0GG1{01}TG-1G11TTTG000101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G110TT0GG11G0GGG{01}01GT1TG11TGGG1GT101010TTG11GTTG11111-TT1100{0T}TT1110TT-1TTGGGG0-10T--TG--GT---------------------------0GT1GGG1G00T0TTGGT1T111GTG1T1T1G0G1G1GGTTGG1110000G1G0GT1111G01001GG01GT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG101T00TT1GTTGT10-1GGG10T1G0GTTG0111TG0-1G1GT1GT1T?1G01GT1G1T11G0T1100110101010100010011 Sambucus_ebulus T1G000-11TG1010G10G00TG0111G1G001T1GTTTT10T0T1-GGGG1T1GT1GG11T{0G}GGTG1G100G1G--1T1GG11GG1GG01-TT1GGT10GGG1G111T1--GG1GTG1TG0110001001G001111GG1G1G0T1TG1G1100GG00TTTTT0-1T000G0G1GTGT1T0T1GT1G1111GTT1G1GGTGTG101GG---T0GG11TG-1G11TTTG000101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTT{GT}G001G100GTTG1G111G00G110TT0GG11G0GGG101GT1TG11TGGG1GT101010TTG11GT1G11111-TT1100TT11110TT-1T1TGGG001G1--TG--GT---------------------------0GT1GGG1GG0T01TGG11T111GTG1T111G0G1G1GGTTGG1110000G1G0GT11-1G01001GG01GT101G0100GTGGTGGTTG000-0G11TT1T00T11TGT1GTG101100TT1GT1GT1G-1GGG10T1G0GTTG0111TG0-1G1GT1GT1T11G01GT1G1T11G0T1100110101010100010011 Sambucus_racemosa T1G000-11TG1010G10G00TG0111G1G001T1GTTT010T0TT-GGGG1T1GT1GG11T0GGTG1G100GTG--1TTGG11GG1G001-TT1GGT10GGG10111T1--GGTGTG1TG011000100TG001111GG1G1G001TG1G1100GG00TTTTT0-1TG00G0G1GTGT1T0T1GTTG1111GTT1G1GGTGTG101GG---T0GG11TG-1G11TTTG000101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTT{GT}G001G100GTTG1G111G00G110TT0GG{1T}TG0GGG001GT1TG11TGGG1GT101010TTG11GT1G11111-TT1100TTT1110TT-1TTGGGG0-{1T}GT--TG--GT---------------------------0G{1T}1GG{1G}1GG0T0TTGGT1T111GTG1T1T1G0G1G1GG{0T}TGG1110000G1G0GT1111G0100{1G}GG01GT10TG0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG101100TT1GTTGT10-1GG?10T1G0GTTG0111TG0-1G1GT1GT1T11G01GT1G1T11G0T1100110101010100010011 Sinadoxa_corydalifolia T1G000-11TG10T0G10G00TG0111GTG001TTGTTTTTT{GT}01T1GTGG1T1GTTGG01T0GG10TG100GTG-11T1GGT1G0100G1-TT1GGT10GGG1G110T1--GGTGT10TG011000100TG001T11GG101GGT1TGTG1100GG000TTT0T-0T000G0G1GTGT1T0TTGTTG1111GTT1G1GGTG-G101G1---T0GG10TG-1G11TTTG000101T0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T0TTTGGTGTG00TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G110TT0GG1100GGG101GT1TG11TGGG1GT101010TTG11GT1G111110TT1100-1T111TTTT1TTGG0-0--GT-TT0TTGT---------------------------TG--GGG1GG000TTGG11T111GTG1T1T1TGG1G1GGTTGG1110000G0{0G}0GT01T1G01001GT01GT101G0100GTGGTGGTTG000-0G11GT0TT0T10TTT1GTG1011GGT11GTTGT10-TGGG101T0010TG0111T00-TG1GT1GT1TTTG0TGT1G1T11G01T000110111010000010011 Tetradoxa_omeiensis T1G000-11TG10T0G10G00TG0111GTG001TTGTTTTTTTT1T1GTGG1T1GT1GG01T0GG10TG100GTG-11T1G0T1G0100G1-TT1GGT10GGGTG110T1--GGTGT10TG0110001001G001111GG101GGT1TGTG1100GG000TTT0T-0T000G0G1GTGT1T0TTGTTG1111GTT1G1GGTG-G101G1---T0GG10TG-1G11TTTG000101T0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T0TTTGGTGTG00TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G110TT0GG1100GGG101GT1TG11TGGG1GT101010TTG11GT1G111110TT1100-1T111TTTG1TTGGG-0--GT-TTGTTGT---------------------------0G--GGG1GG000TTGG11T111GTG1T1T1TGGTG1GGTTGG111000TG0G0GT0111G01001GT01GT10100100GTGGTGGTTG000-0G11GT0T-0T10TG1TGTG1T110GT11GTTGT10-1GGG101TG01TTG0111T00-0G1GT1{GT}T1TTTG0TGT1G1T11G01T000110111010000011011 Viburnum_acerifolium T1G000-11TG11T0G10G001G0111G1G00101GTTT00TT01T-0GGGTG1011GGT1GGGG1G1GT10{1G}---1111GG11GGT{1G}1111TT1GGT1GGG01G1T1T11GGG1GG111GG111001001G001111GG1G1G0T1TG1G1100GG000TTT00-1TG00G0G101G111111GTTG1111GTT1G1-G1GTG1G1GG---GGTG1TTG-1{0G}1TTT1G00T101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001G---------------------------TG10G00T111GTG00110T1G--TTTTTG001G100GTTG1G111000G110TT0GG11G0GGG101GT1TG11TGGG1GT101G10TTG1-GT1G1111----1010---111------------------TG-TGT1111G000GG1G0GG10--11GGG1G-0GG-GGG1GG0T0TTGG11T111GTG1TT11GGGTG1GGTTGG1110000G1G0GT1111GG10G1GG01GT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG1GGT11T11GTTG110T1GGG101T111TTG0111TG0-1G1G11GTT11TG01GG1G-TT1G011100000101010100110001 Viburnum_dentatum T1G000-11TG1T10G10G001G0111G1G00101GTTT001T01T-0GGGTG10T1GGT1GGGGTG1GT10G--1111TGG11GGTG111-1T1GGT1GGGG1G1T1TT1G0G1G0111GG111001001G001111GG1G1G0T1TG1G1100GG000TTT00-1TG00G0G10TG11111TG1TG1111GTT1G1GGTG1G1G1GG---TTGG1TTG-1G1TTT1G00T101-?????????????????????????????????????????????????????????????????????????????????????11GTG00110T1G0GTTTTTG001G100GTTG1G111000G110TT0G?1-G0-GG?01GT1TG11TGGG1GT101G10TT---GT1G1111----1010---111------------------TG-TGT11110000G0GG---10T0TTGGG1G-0GG-GGG1GG0T0TTGG11T111GTG1TTT1GGGTG1GGTTGG111000001G0GT1111GG1001G-01GT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG1GGT11T11GTTG110T1GGG101T1T1TTG0111TG0-TG1G11GTT11TG01GG1G1TT1G0--100100101010111110001 Viburnum_lentago T1G000-11TG1110G10G001G0111G1G00101GTT1001001T-GGGGTG1?11GGT1GGGGTG1GT10G--111T1GGT1GGTG1111111GGT1GGG01G1T?TT1G0G1G?1?TGG11-001001G001111GG1G1G0T1TG1G1100GG000TTT00-1TG00G0G10TG111111GT1G1111GTT1G1GGTGTG1G1GG---TTGG11TG--G1TTT1G00T101-???????????????????????????????????????????????????????????????????????????TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G110TT0GG11G0GGG101GT1TG11TGGG1GT101G10TTG1-GT1G1111----1010---111------------------TG-TGT11110000GGG----10--11GGG1G-1G--GGG1GG0T0TTGG11T111GTG1T1T0GGG1G1GGTTGG1110000G1G0GT1111GG1001GG01GT101G01G0G1GGTGGTT?000-0G11TT1TT0T11TGT1GTG1GGT11T11G1TG11011GGG101T111TTG0111TG0-TG1GT1GTT11TG01GG1G1TT1G0--100000101011100110001 Viburnum_plicatum T1G000-11TG11T0G10G001G0111GTG0010TGTT100TT01T-0GGGTG10TTGGT10GGGT{0G}1GT10{1G}---111TGGT1GGTG111-1T1GGT1GGG01GTT1TT1G0G1GG111GG111001001G001111GG1G1G0T11G1G1100GG0{0T}0TTT00-1TG00G0G10TG1111T1GTTG1111GTT1G1GGTGTG101GG---TTGG11TG-1G1TTT-G00T101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001G--------------------------TTG10G0{0T}T{1T}11GTG00110T1G0GTTTTTG001G100GTTG1G111000G110TT0GG11G0GGG{1T}01GT1TG11TGGG{1T}GT101G10TT01-GT1G1111----1010---111------------------TG-TGT1111000-GGGG---10--TTTGG10-0G--GGG1GG0T0TTGG11T111GTG1TTT1TGGTG1GGTTGG1TT0000G1G0GT1111GG1001GG01GT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGTTGTG1GGT11T11GTTG110T1GGG101T111TTG0111TG00TG1G11GTT11TG01GG101TT1G011100100101010100110000 Viburnum_sieboldii T1G000T11----T0G10G001G0111G1G0010T?TT100TT01T-0GGGTG10TTGGT1GGGGTG1GT10G--1111TGG11G0TG111-1T1GGT1GGG01G1T1TT1G0G1GG101GG111001001G001111GG1G1G0T1TG1G1100GG000TTT00-1TG00G0G10TG1111110TTG1111GTT1G1GGTGTG101GG---TTGG1TTG--G1TTTTG00T101-???????????????????????????????????????????????????????????????????????????????0G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111000G110TT0GG1-G0-GG-01GT1TG11TGGG1GT101G10TTG1-GT1G1111----1010---11T------------------TG-TGT111T000-GGGG---10--TTGGG1G-0G--GGG1GG0T0TTGG11T111GTG1TTTT1GG1G1GGTTGG1TT0000G1G0GT11T1GG100TGG01GT101G0100GTGGTGGTTG000-0G11TT1TT0T11TGT1GTG1GGT11T11GTTGT10T1GGG101T111TTG0111TG0-TG1G11GTT11TG01GG101TT1G0--100100101011111110001 Weigela_hortensis T1G000-11TG1010G10G001G0111G1G00101GTT1GT01011--GGG0-1GT1GGT11GGG1G1GT10G--11110GG11GGTG1T1110TGG11GGGG0G1111T--GG1T11T1G0T1G000-11G001111GG1G1G0T11G1G1100GG00TT0100010G00GGG1TTG11T111GTTG1111GT11G1GGTG1G1G1GGG0GT0G-1T1G11T1TTT1G000101-0001G01T1T1GG1001GG0T0T1T1GG1T1T1G10T1G0TG00G001GT0G1G000TG1G0T01TTGGTGTG00TTG10G00T111GTG00110T1G0GTTTTTG001G100GTTG1G111G00G1101T0GG11G0GGG101GT1TG11TGGG1GT101G10T1G1-GT1G1111----1100---111------------------1G-1GT111-000GGGT1G1G1G1GG1GGGGGG0G-----1GG0G00TGG11T111GTG1T111GGG1G1GG1TGG1110000T1G0GT1111GG1G01GG01GT101G0100GTGGTGGTTG000G0G111T1T10T000GT1GTG1GGTT1111GT1G1--T1GGG1GG1100GTG0111TG01G1GT1GT1GT11G01GG1G1T11G011111011000100100100100 ; END; [ The following blocks are output data for analysis step 1683 ] BEGIN TREES; TITLE Tb6459; LINK TAXA = TaxaForAnalysisStep1683; TREE Fig._5 = [&R] ((Weigela_hortensis,Diervilla_sessilifolia),(((Viburnum_dentatum,((Viburnum_sieboldii,Viburnum_plicatum),Viburnum_acerifolium)),Viburnum_lentago)Viburnum,(((Adoxa_moschatellina,Tetradoxa_omeiensis),Sinadoxa_corydalifolia)Adoxina,((((Sambucus_caerulea,Sambucus_racemosa),Sambucus_australis),Sambucus_ebulus),Sambucus_australasica)Sambucus)Adoxoideae)Adoxaceae); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5791] END;