#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on November 10, 2026; 8:40 GMT TreeBASE (cc) 1994-2008 Study reference: Donoghue M.J., Eriksson T., Reeves P., & Olmstead R. 2001. Phylogeny and Phylogenetic Taxonomy of Dipsacales, with special reference to Sinadoxa and Tetradoxa (Adoxaceae). Harvard Papers in Botany, 6(2): 459-479. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S981] [ The following blocks are input data for analysis step 1685 ] BEGIN TAXA; TITLE TaxaForAnalysisStep1685; DIMENSIONS NTAX=16; TAXLABELS Abelia Adoxa Diervilla Dipelta Dipsacaceae Kolkwitzia Leycesteria Linnaea Lonicera Pittosporum Sambucus Symphoricarpos Triosteum Valerianaceae Viburnum Weigela ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M521] TITLE rbcL_plus_Judd_et_al.; LINK TAXA = TaxaForAnalysisStep1685; DIMENSIONS NCHAR=1463; FORMAT DATATYPE=DNA SYMBOLS= "0 1 2 3 4 5" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 370 380 390 400 410 420 430 440 450 460 470 480 490 500 510 520 530 540 550 560 570 580 590 600 610 620 630 640 650 660 670 680 690 700 710 720 730 740 750 760 770 780 790 800 810 820 830 840 850 860 870 880 890 900 910 920 930 940 950 960 970 980 990 1000 1010 1020 1030 1040 1050 1060 1070 1080 1090 1100 1110 1120 1130 1140 1150 1160 1170 1180 1190 1200 1210 1220 1230 1240 1250 1260 1270 1280 1290 1300 1310 1320 1330 1340 1350 1360 1370 1380 1390 1400 1410 1420 1430 1440 1450 1460 1470 1480 1490 1500 1510 1520 1530 1540 1550 1560 1570 1580 1590 1600 1610 1620 1630 1640 1650 1660 1670 1680 1690 1700 1710 1720 1730 1740 1750 1760 1770 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Abelia ??????????????????????????????????????????????????????????????????0133033030131132013032000110002030132030313322102103311202300131131001132202331101132002002102222112122302132112003133100132230103220100132323220112032201330110211332031233010002221203213011010312021112332132202002000031003030?32133032302133011103302011333332002002233132330130010323330133130332322230032303332223310002111321212131301231322002031321200311132312133030130000133311002211121131032210311002332020202030003320010023032231211111323322203230130330001130003322223303132130000013032230202122333032003231301232232201332033330110002032032020012320013111001103330321233220202031233313303333232112002111333030002101002132000102232000310002221033013320032130132102230103232002000320310000222132303332110202003322202331130312300321012013013300102222203310132100030130213322131003033211202030032211301331331010311011212100321032102330332030201020020031032230301013331232301302130002123301230323132232202031030331013112230112302302230001332002222000202020310133322213332332033301321212032033330332000002031200231212230333033310131002033222313131301122232331321112322133122222230331012333221030321112131320112020313332222032033132301301023312212202200133302201011133222200032121112232112302130031202312131302002103232301002131230032022201232031332131232022230032000330311232222133210003?????????????????????????????????????????????????????????????????????????????010101110021311100112101211010-0011 Adoxa ??????????????????????????0023213220331000213223233000203301000332013303303013113201303200011000203013203031332210210331120230013113100113220233110113200200210222211212230213211200313313013223010322010013232322013203220133011001133201123301000222120321301101031202111233213220200200013100333033213303230213301110330201133333200200223313233013001032333013311033232223003230333222331000211132121213130123132200203132121033113233213303233000013331100221112113103223031100233202020203000332001002303223123111132332220323013033000113000332220330313213000001301223020212233303200323131121223220133203333011000201203202001233001313100110333032123322020203123331330333323211200211133303000210100213200010221200031000222103301332003213013212223010321200200032032000020213030333211020200332220233113031230032101201301130011223220331013210003013011332213103303321120203003221130133133101031101121210032103210233033203020102002003103223030101333123230130213000212330123032313223220203103033101311223011230230223000133200222200020202031013332223333233203330130121203203333033200000203120023121223033303331013100203322231313130110223233132112230213312222223033101233322103032113213132011202031333222203203311230130102331223220220013130220101113322220003210112223211230213003120230213130200210323230100213123003202220123?03?3321312320222300320003303312320221331100032202311320013021321321332320223032200220003100033320033320021003220301333230011111?00?11020101000102{12}0??110-0?00 Diervilla ??????????????????????????002321322033100021222123300020330100033201330330301311320130320001100020301320303033221021033112023001311310011322023311011320020021022221121223021321120031331001322301032201001323232201120322013301102113320312330100022212032130110103120211123321322020020000310030303321330323021330111033020113333320020022331323301300103233301331303323222300323033322233100021113212121313012313220020313212003111321121330321300001333110022111211310322103110023320202020300033200100230322312111113233222032301303300011300033222233031321300000130122302021223330320032313012322322013320333301100020320320200123200131110011033303212332202020312333133033332321120021113330300021010021320001022320003100022210330133200321301321022301032320020003203100002221323033321102020033222023311303123003210120130133001022222033101321000301302133221330330332112020300322113013313310103110112321003210321023303320302010200200310322303010133312?23013021300021233012303231322322020310303310131122301123023022300013320022220002020203101333222133323320333013212120320333303320000020312002312122303330333101310020332223131313011222323313211123221331222222303310123332210303211121313201120203133322220320331123113010233122122022001333022010111332222000321211122321123021300312023121313020021032323010021312300320222012320313321312320222300320003303112322221132100032202311320013021321321332320223032200220203100033320033320021003220301333200301010110002131100?12210001001010010 Dipelta ??????????????????????????0023213220331000212221233000203301000332013303303013113201303200011000203013203031332210210331120230013113100113220233110113200200210222211212230213211200313310013223010322010013232322011203220133011021133203123301000222120321301101031202111233213220200200003100303033213303230213301110330201133333200200223313233013001032333013313033232223003230333222331000211132121213130123132200203132120031113011213303233000013331100221112113103221031100233202020203000332001002303223121111132332220323013033000113000332222330313213000001303223020212233303200323130123223220133203333011000203203202001232001311100110333032123322020203123331330333323211200211133303000210100213200010223200031000222103301332003213013210223010323200200032031000022213230333211020200332220233113031230032101201301330010222220331013210003013021332213103303321120203003221130133133101031101121210032103210233033203020102002003103223030101333123230130213000212330123032313223220203103033101311223011230230223000133200222200020202031013332221333233203330132121203203333033200000203120023121223033303331013100203322231313130112223233132111232213312222223033101233322103032111213132011202031333222203203313231130102331221220220013330220101113322???0032121112232112302130031202312131302002103232301002131230032022201232031332131232022230032000330311232222133210003?????????????????????????????????????????????????????????????????????????????01010111002131110?112102211010-0?11 Dipsacaceae 03231011010001020201300021002321322033100021222123300020330100033201330330301311320130320001100020301320303{12}33221021033112023001311310011{13}22023311011320020021022{02}21121023021321120031331001322301032201001323232201120322213301102313320312330{13}00022{23}12032130110103120211123321{13}2202002000031003{03}303321330323021330{13}11033020113333320020022331323301300103233301331303323222300323033322233100021113212121313012313220020313212003{13}11323121330323300001333110022111211310{13}22103110023320202020300033000100230322312{01}11113{02}33222032301303300011{03}00033222233231321300000130322302021{01}23330320032313012322322013320333301100020320320200123200{13}31{01}10011033303212{13}32202020312333133033332321120021113330{13}000210100{02}13200010223200031000220103301332003213013{02}1{02}2230103232002000320310000222132{13}{13}33321302020033222{01}{02}3311303123003210320{13}30133001022222033101321000301302{13}13221310330332{13}120203003221130133133101031101121210032103210233033203020102002003103223030101333123230130213000212330123032313223220203103033101311223011230230223000133200222200020{12}{03}203{01}01333222133323320333013212120320333303320{01}00020312002312122303330333101310020332223{13}31313011{02}22323313211123221331222222303310123332210303211{13}213132011202031333222203203313231130102331221220220013330220101113{23}{23}222000321211122321{13}2302130031202312131302002103232301002131230032022201232031332131232022230032000330311232122133{12}100032202311320013021321321332320223032200220203100033320{02}33320021003220301333230011{01}11?1?2?2131110011210??1?011-1??? Kolkwitzia ??????????????????????????002321322033100021222123300020230100033201330330301311320130320001100020301320303133221021033112023001311310011322023311011320020221022221121223021321120031331001322301032201001323232201120322013301102113320312330100022212032130110103120211123321322020020000310033303321330323021330111033020113333320020022331323301300103233301331303323222300323033322233100021113212121313012313220020313212003111321321330323300001333110022111211310322103110023320202020300033200100230322312111113233???032301303300011300033222233031321300000130322302021123330320032313012322322013320333301100020320320200123200131110011033303212332202020312333133033332301120021113330300021010021320001022?????????022210330133200321301321022301032320020003203100002221323033321102020033222023311303123003210120130133001022222033101321000301302133221310130332112020300322113013313310103110112121003210321023303320302010200200310322303210333312323013021300021233012303231322322020310303310131122301123023022300013320022220002020203101333222133323320333013212120320333303320000020312002312122303330333101310020332223131313011222323313211123221331222222303310123332210303211?2131320112020313332222032033132311301023312212202200133302201011133222200031121112232112302130031202312131302002103232301002131230032022201232031332131232022230032000330321????????????????????????????????????????????????????????????????????????????????????????????010101110021311100112101211010-0011 Leycesteria ??????????????????????????002321322033100021222123300020330100033201330330301311320130320001100020301320303133221021033112023001311310011322023311011320020021022221121223021321320031331001322301032201001323232201120322013301102113320312330100022212032130110103120211123321322020120000310030303321330323021330111330020113333320020022331323301300103233301331303323222300323033322233100021113212121113012313220020313212003111323121330323300001333110022111211310322303110023320202020300033200100230122312111113233222032301303300011300033222233231321300000130122302221223330320032313012322322013320333301100020320320200123200131110011033303212332202020312333133033332321120021113330300021010021320001122320003100022210330133200321301321022301032320020003203100002221323033321102020033222023311303123003210120130133001022222033101321000301102133221310130332112020300322113013313310103110112121003210321023303320302010200200310322303010133312323013021300021233012303231322322020310303310131122301123023022300013320022220002020003101333222133323320333013212120320333303320000020312002312122303330333101310020332223131313011222323313211123???331222222303310123332210303211?21313201120203133322020320331123133010233122122022001333022010111332222000321011122????230213003120231213130200210323230100213123003202222123203133213123202223003200033031121??22133210003?????????????????????????????????????????????????????????????????????????????01010120002{01}301000112{01}020100102011? Linnaea ??????????????????????????002321322033100021222?2330002033010003320133033030131132013032000110002030132030313322102103331202300131131001132202331101132002002102222112122302132112003133100132230103220100132323220112032201330110211332031233010002221203213011010312021112332132202002000131003330332133032302133011103302011333332002002233132330130010323330133130332322230032303332223310002111321212131301231322002031321200311132312133032330000133311002211101131032210311002332020202030003320010023032231211111323322203230130330001130003322223303132130000013032230202122333032003231301232232201332033330110002032032020012320013111001103330321233220202031233313303333232112002111333030002101002132000102232000310002221033013320032130132102230103232002000320320000222132303332110202003322202331130312300321012013013300102222203310132100030130113322131003033211202030032211301331331010311011212100321032102330332030201020020031032230301013331232301302130002123301230323132232212031030331013112230112302302230001332002222000202020310133322213332332033301321212032033330332000002031200231212230333033310131002033222313131301122232331321112322133122222230331012333221030321112131320112020313332222032033132311301023312212202200133302201011133222200032121112232112302130031202312131302002103232301002131230032022201232031332131232022230032000330311232022133210003?????????????????????????????????????????????????????????????????????????????01010111002{01}31110?112?0{12}211010-0?11 Lonicera 03231011010001020201300021002323322033100021{23}22323300020{23}30100033201330330301311320{02}30{13}{12}0001100{02}20301320303{01}33221021033112023001311310011122023311211320020021222221{13}212230213213200313310011223010322010013232322011203220133011021133203123301000222120321301{12}{02}10312021112332132202002000{01}31003{03}30332133032302133011103302011333332002002233132330130010323330133130332322230032303332223310002111321212131301231322002031321200311132{13}{13}21330323300001333110022111211310322{23}03{13}10023320202020300033200100230122{13}12{13}111132332220323013033000113000332222330313213000001301223022212233303200323130123223220133203333011000203203202001232001311100110333032123322020203123331330333323211200211{01}333{03}300021010001320001022320003100022210330133200321301321022301032320020{01}03203{12}000022213230333211020200332220233113031230032101201301330010222220331013210003011011332213101303321120203003221130133133{13}0103110112121003210321023303320302010200200310322303{12}10{13}33{13}12{13}23013021{03}00021233012303231322322020310303310121122301123023022300013320022220002020003101333222133323320333013212{13}20320333303320000{02}203120023121223033303331013100203322231313130112223233132111232213312222223033101233322103032111213332011202031333220203203311231130102331221220220013330220101113322220003210113223211230213003120231213130200210323230{01}0021312300320{02}22{02}1232031332131{02}{23}20{01}22300320003303{13}12{13}2{01}221332100032202{13}11{23}20013021321321332320{02}23032200{02}202031{02}{03}0333{02}0{03}3332{01}021{03}{02}3220{13}{02}{13}3332{23}0{03}0101{01}120102{01}3{01}100011200{012}01001020110 Pittosporum ????10110100010200013000210023233220331000213223233000203301000332013303303013113201303100011000203013203031332210210331120230013113100113220233112113200200210222211212230213211200313313013223010322010013232322011201220133011021133203123301000222120321301200031202111233213220200200003100333033213303230213301110330201133333200200223313233013001032333013311033230223003230333222331000211132121213130123132200203132120031113233213303233000013331100221112113103221031100233202020203000332001002301221121111132332220323013033000113000332222330313213000001301221020212233303200323131123223220133203333011000201203202001232001311100110333032123322020203123331332333323211200210133303000210102233200010223200031000222103301332003213013212223010321200201032032000022211230333211020200332220233113031230032101201301330010222220331013210003013011332213103303321120203003221130133133101031101121210032101210233033203020102002003103223032101333123230130213000212330123032313221220203103033101211223011230230223000133200222200020201031013332221333233203330332123203203333233200000203120023121223033303331011100201322231313130110223233132111232213312222223033101233322103032113213132011202031333222203203311230130102331223220220013330220101113322220003210111223211230213003120230213130200210323230100213123003202220123203133213023200223003200033033123202213213000322021111200130213213213323202030322002202031000333200333?????????????????????00001??00?000?00??0?001000?00000??? Sambucus ?????????????????????????????32132203310002132232330002033010003320133033030131132013032000110002030132030313322102103311202300131131001132202331101132002002102222112122302132112003133130132230103220100132323220132032201330110211332031233010002221203213011010312021112332132202002000231003330332133032302133011103302011333332002002233132330130010323330133110332322230032303332223310002111321212131301231322002031321200331132{23}3213303233000013331100221112113{13}0322303110023320202020300033200100230322312311113233222032301303300011300033222233031321300000130122302021223330320032313112122322013320333301100020120320200123200131310011033303212332202020312333133033332321120021{02}{01}3330300021010021320001022{13}20003100022210330133200321301321222301032120020{01}0320320000202130303332110202003322202331130312300321012013013300102222203310132100030130113322131033033211202030032211301331331010311011212100321032102330332030201020020031032230321013331232301302130002123301230323132232202031030331012112230112302302230001332002222000202030310133322233332332033301321212032033330332000002031200231212230333033310131002033222313131301102232331321122322133122222230331012333221030321132131320112020313332222032033112301301023312232202200133302201011133222200032101112232112302130031202302131302002103232301002131230032022201232031332131232022230032000330311232022133210003?????????????????????????????????????????????????????????????????????????????{01}111100011100?101100102{12}00111030000 Symphoricarpos ??????????????????????????????????????????????????????????????????????????????????????????????????????2030313322102103311202300131131001132202331121132002002102222112122302132132003133100132230103220100132323220112012201330110211332031233010002221203213011010312021112331132202002000131003330332133032302133011103302011333332002002233132330130010323330133130332322230032303332223310002111321212131301231322002031321200311132312133032330000113311002211121131032230311002332020202030003320010023012221211111323322203230130330001130003322223303132130000?1301223022212233303200323130123223220133203333011000203203202001232001311100110333032123322020203123331330333323211200211033303000210102213200010223200031000222103301332003213013210223010323200201032032000022213230333211020200332220233113031230032101201301330010222220331013210003011011332213101303321120203003221130133133101031101121210032103210233033203020102002003103223032101333123230130213000212330123032313223220203103033101311223?112302302230001332002222000202000310133322213332332033301321???032033330332000002331200231212230333?333101310020332223131313011222323313211123221331222222303310323332210303211121313201120203133322020320331123113010233122022022001333022010111332222000321011122321?230123003120231213130200210323230100213123003202222123203133213123202223003200033031123?022133210003220231132001302132132133232022303220022020310003332003332002100322030133323000101{01}1{12}0102{01}301000?120{01}2211010-0110 Triosteum ??????????????????????????002323322033100021222?233000203301000332013303303013113201303200011000203013203031332210210331120230013113100113220233110113200200210222211212230213213200313310013223010322010013232322011203220133011021133203123301000222120321301101031202111233213220200200023100333033213303230213301110330201133333200200223313233013001032333013313033232223003232333222331000211132121213130123132200203132120031113231313303233000013331100221112113103223031100233202020203000332001002301223121113132332220323013033000113000332222330313213000001301223022212233303200323130123223220133203333011000203203202001232001311100110333032123322020201123331330333323211200211133303000210100213200010223200031000222103301332003213013210223010323200203032032000022213230333211020200332220233113031230032101203301330010222220331013210003011011132213101303321120203003221130133133101031101121210032103210233033203020102002003103223032101333123230130213000212330123032313223220203103033101211223011230230223000133200222200020200031013332221333233203330132121203203333033200000203120023121223033303331013100203322231313130112223233132111232213312222223033101233322103032111213132011202031333220203203311231130102331221220220013330220101113322220003210111223???2302130031202312131302002103232301002131230032022221232031332131232022230032000330311232122133210003?????????????????????????????????????????????????????????????????????????????11010?2000213010001120021?1010-0110 Valerianaceae 03231011010001020{02}013000210023213220331000212221233000203301000332013303303013113201303{12}0001100020301320303133221021033112023001311310011{23}22023311{03}{12}1320020021022221121{02}2302132112003133100132230103220100132323220112032201330{12}{12}02113320312330100022212032130110103120211{13}233213220200200{02}{12}3{01}0033303321330323021330{13}11033020{13}1333332002002233132330{13}{13}00{13}0323330133{13}{13}033232223003230333222331000211{13}321{02}{123}213130123{13}3220020313212003111323{13}21330323300001333110022111211310{13}223031100233202020203000{13}3200100230122312{13}11113{02}33222032301303300011{13}00033222233{02}31321300000130322302{02}21223330320032313012{13}22322013320333301100020320320200123200131110011033303212332202020{13}12333133033332321120021{01}{013}33303000{23}10100{023}1320001022320003100022210330133200321{23}01321{02}22301032320020003203100002221323033321102{02}20033222023311{23}03123003210{13}201301{13}3001{02}22222033101{13}21000{13}013{01}{12}1{13}3{23}21310330332112020300322{13}13013313310103110112121003210321023303320302010200200310322303010133312323013021{13}00021233012303{23}31322322020310303{03}1{03}131122301123023{01}2{12}300013320022{02}20002020203{01}{01}133322213332332033301321212032033{03}3033200000203120{02}2312122{13}03330333101{13}100203322231313{01}3{02}11222323313{02}11{13}23{02}21331{12}22222303310123332210303211{13}21313201120203133322220320331323113010233122122022001333022{02}{12}01113322220{02}{02}3{02}1211{03}2232112302130031202{23}{01}21313020021{03}{03}2323010{02}{12}{13}{23}12300320222012320313321312{13}202223003200033031123212213{23}{12}100032202311320013021321321332320223032200{02}202031000333200333{23}{01}{02}{01}1023020301{123}{23}{23}{23}{03}{03}{013}11{01}11?10{02}?21311100112101211011-1??? Viburnum 0323101101000102020130002100232132203310002132232330002033010003320133033030131132003032000110002030132030313322102103311202300131131001132202331101132002002102222112122302132112003133130132230103220100132323220132032201330110211332031233010002221203213011010312021{12}{13}23321322{01}200200{02}{012}310033303321330323021330111033020113333320020022331323301300103233301331103323222300323033322233100021113212121313012313220020313212003{13}11323321330323300001333110022{13}11211310322103110023320202020300033200100230322312{03}111132332220321013033000113000332222330313213000001301223020212233303200323131121223220133203333011000201203202001232001311100110333032123322020203123331330333323211200210{01}33303000210100{02}1320001022320003100022210330133200321301321222301032120020{03}03203{12}00002221323033321102020033222{01}23311303123003210320130133001022222033101{13}2100030130{12}133221310330332112020300{13}22113013313310103110112{13}210032103210233033203020102002003103223032101333123230130213000212330123032313223220203103033101{23}1122301123023022300013320022220002020{123}03{01}013332221333233203330132121203203333{02}332000002031200231212230333033310131002033222313131301102232331321112{13}2{12}{12}331222222303310123332210303{12}111213132011202031333222203203311230130102331223220220013330220101113322220003{12}1{02}11{01}22321123021300312023021313020021032323010021312300320222012320313321312320222300320023303112320?213021000322023113200130213213213323202230322002200031000333200333200210032203013332300010{01}00001110{01}?10010010213010103000{01} Weigela ??????????????????????????0023213220331000212221233000203301000332013303303013113?013032000110002030132030313322102103331202300131131001132202331101132002002102222112122302132112003133100132230103220100132323220112032201330110211332031233010002221203213011010312021112332132202002000031003030332133032302133011103302011333332002002233132330130010323330133130332322230032303332223310002111321212131301231322002031321200311132112133032130000133311002211121131032210311002332020202030003320010023032231211111323322203230130330001130003322223303132130000013012230202122333032003231301232232201332033330110002032032020012320013111001103330321233220202031233313303333232112002111333030002101002132000102232000310002221033013320032130132102230103232002000320310000222132303332110202003322202331130312300321012013013300102222203310132100030130213322133333033211202030032211301331331010311011212100321032102330332030201020020031032230301013331232301302130002123301230323132232202031030331013112230112302302230001332002222000202020310133322213332332033301321212032033330332000002031200231212230333033310131002033222?1313130112223233132111232213312222223033101233322103032111213132011202031333222203203311231130102331221220220013330220101113322220003212111223211230213003120231213130200210323230100213123003202220123203133213123202223003200033031123?022133210003?????????????????????????????????????????????????????????????????????????????01010110002131100?12210001001010010 ; END; [ The following blocks are output data for analysis step 1685 ] BEGIN TREES; TITLE Tb6461; LINK TAXA = TaxaForAnalysisStep1685; TREE Fig._3 = [&R] (Pittosporum,(((Adoxa,Sambucus),Viburnum)Adoxaceae,((((Lonicera,Symphoricarpos),Triosteum),Leycesteria)Caprifolieae,((Diervilla,Weigela)Diervilleae,((Abelia,Dipelta,Kolkwitzia,Linnaea)Linnaeeae,(Dipsacaceae,Valerianaceae))Linnina))Caprifoliaceae)); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5794] END;