#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 05, 2025; 18:15 GMT TreeBASE (cc) 1994-2008 Study reference: Boisserie J., & Bibi F. 2021. "Hippopotamidae from the late Miocene Baynunah Formation, Abu Dhabi Emirate, U.A.E." In: Bibi F., Kraatz B., Beech M., & Hill A., eds. Sands of Time: Fossils from the Late Miocene Baynunah Formation, U.A.E. pp. . Cham, Switzerland, Springer. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S21683] [ The following blocks are input data for analysis step 21825 ] BEGIN TAXA; TITLE TaxaForAnalysisStep21825; DIMENSIONS NTAX=58; TAXLABELS Aepinacodon Afromeryx Amphimeryx Amphiraghatherium Anthracokeryx_tenuis Anthracokeryx_thailandicus Anthracotherium_chaimanei Anthracotherium_magnum Archaeomeryx Archaeopotamus_harvardi Archaeopotamus_qeshta Bothriodon Bothriogenys_andrewsi Bothriogenys_fraasi Bothriogenys_gorringei Bothriogenys_orientalis Brachyodus_aequatorialis Brachyodus_depereti Brachyodus_onoideus Bunophorus Cebochoerus Choeropotamus Chorora_hippopotamine Chororatherium Dacrytherium Dichobune Diplobune Elomeryx_borbonicus Elomeryx_crispus Entelodon Epirigenys Gobiohyus Gujaratia Hallebune Heptacodon Hexaprotodon Homacodon Indohyus Kenyapotamus_coryndonae Kenyapotamus_ternani Kenyasus Khirtaria Libycosaurus_anisae Libycosaurus_bahri Lophiomeryx Merycoidodon Merycopotamus_dissimilis Merycopotamus_nanus Microbunodon Mixtotherium Morotochoerus Myaingtherium Palaeochoerus Paroxacron Perchoerus Siamotherium Sivameryx Xiphodon ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M57689] TITLE Hippopotamidae_late_Miocene_Baynunah_Formation; LINK TAXA = TaxaForAnalysisStep21825; DIMENSIONS NCHAR=164; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4" MISSING=? GAP= -; CHARSTATELABELS 1 number_of_lower_incisors / three two one, 2 lower_incisor_morphology / not_caniniform at_least_one_caniniform_lower_incisor, 3 relative_dimensions_of_lower_incisors / all_of_equal_size one_or_two_more_developped, 4 most_developped_incisor / 'I/2 ' 'I/3 ' 'I/1', 5 transverse_section_of_lower_incisors_crowns / strongly_irregular about_rounded, 6 lower_incisor_cervix_morphology_ / no_deep_indentation 'indented cervix, indentation as long as the root diameter on the cervix' deep_identation_longer_than_the_root_diameter_at_cervix, 7 crown_of_lower_I1 / straight__ 'spatulate (with convex mesial and distal border) ', 8 'presence of a median lingual pillar (lingual rib) on lower i1' / yes no, 9 Lower_canine_cross_setcion_at_cervix / subcircular elliptical_, 10 cristids_on_lower_canine_enamel_caps / none one_distal 'two, one mesial and one distal' one_mesial, 11 wear_on_lower_canine / distal_wear_facet_contact_with_canine_ mesial_wear_facet_contact_with_I3_, 12 groove_on_labial_side_of_lower_canine_ / no yes, 13 groove_on_lingual_side_of_lower_canine / no yes, 14 lower_canine_in_male / 'fang-like' premolariform incisiform, 15 crown_of_lower_canine_in_male / small_near_premolar_size at_least_twice_the_premolar_size 'prolonged growth to ever-growing ', 16 'p/1 roots' / one two, 17 paraconid_on_lower_premolars / no yes, 18 'elongated p/3 ' / 'no (shorter or equal than M/1 lenght)' 'yes (longer than M/1 lenght)', 19 'three lobed P/3' / no yes, 20 accessory_cusp_on_the_preprotocristid_of_lower_premolars / none one_or_more, 21 'lingual contour at cervix of P/4 in occlusal view' / convex_to_straight concave, 22 'labial wall on P/3 or P/4' / convex concave, 23 'Orientation of postprotocristid on P/3' / distal distolingual distolabial, 24 'high cingulid on labial face of P/3' / no yes, 25 'preprotocristid mesiolingualy curved on P/3' / no yes, 26 'mesial accessory cusp on preprotocristid on P/3' / no Shoulder_like_structure_on_lateral_view_of_preprotocristid_of_lower_premolars complete_accessory_cusp, 27 Change_in_the_orientation_of_the_preprotocristid_mesialy_to_the_junction_of_accessory_mesiolingual_crest_on_lower_premolars / no yes, 28 'Orientation of the endoprotocristid on P/4' / absent separated_from_postprotocristid_at_the_protoconid_apex_and_then_strait_and_distolingual fused_with_postprotocristid_in_part_and_then_curved_mesiolingually, 29 'entoconid on P/3' / never at_least_on_some_specimen, 30 'P/3 hypoconid' / no yes, 31 'distolingual cingulid on P/4 in lingual view' / forming_a_continuous_wall_lingually_until_the_distostylid_ reaching_the_level_of_the_distal_basin_and_keeping_be_shallow_until_the_distostylid 'reaching the level of distal basin and then being high when joigning the distostylid (distolingual notch of cingulid)', 32 'presence of a preentocristid on P/3 and/or on P/4' / no yes, 33 'mesiolingual secondary cristid on P/4 (cristid connecting lingual margin and preprotocristid)' / no yes, 34 'labial cingulid form a V (indentated) on P/4 before to reach the distal cingulid' / no_ yes_, 35 'marked postprotofossid on P/4' / absent present, 36 'postectoprotocristid on P/4' / no_ yes_, 37 'hypoconid on P/4' / no_ 'yes (even incipient) ', 38 'ectoprotofossid on p/4' / absent frequent, 39 premetacristid_on_lower_molars / strong reduced_or_missing_, 40 'paraconid on lower molars, almost on unworn specimens' / yes no, 41 lower_molar_trigonid_ / equal_in_height_with_talonid higher_than_talonid, 42 connection_between_premetacristid_and_preprotocristid_on_lower_molars / yes_ no_, 43 postectoprotocristid__on_lower_molars / absent 'reduced in the valley to fully developped at least on M/1', 44 'postprotofossid on lower molars at least on m/3' / no_ yes_, 45 'postmetacristid on M1-2' / curving_toward_postprotocristid_forming_a_transverse_bridge_with_it orientated_toward_the_centre_of_the_tooth forming_a_rounded_postmetaconulid_not_preferentiallyorientated joins_prehypocristid, 46 ectoprotofossid_on_lower_molars / absent present, 47 Ectometafossid__on_lower_molars / yes_ no_, 48 endometacristid__on_lower_molars / no_or_slightly_expressed_much_more_like_an_enamel_fold present, 49 postectometacristid_on_lower_molars / lightly_marked_to_absent always_present_and_well_marked, 50 premetafossid_on_lower_molars / present absent, 51 preentocristid_ / absent present, 52 preentocristid_connects / endohypocristid prehypocristid__toward_the_cuspid_apex prehypocristid_toward_its_mesial_extremity, 53 postectoentocristid__on_lower_molars / absent_ present_but_more_like_a_keel_on_cusp present_and_well_individualized_from_the_cusp, 54 ectoentocristid / present absent, 55 postentocristid__on_lower_molars / absent present_, 56 prehypocristid_dividing_in_two_mesial__arms_on_lower_molars / yes_ no_, 57 'prehypocristid inflated (not sallient when unworn) ) in transverse valley of lower molars' / no_ 'yes (even to form a conulid) ', 58 prehypocristid_reaches / 'median part of transverse valley (eventually labial)' lingual_part_of_transverse_valley labial_part_of_the_transverse_valley, 59 main_arm_of_prehypocristid_connects / 'trigonid distal walls (junction between cristids from metaconid and protoconid)' postmetafossid lingual_margin_of_transverse_valley postmetacristid, 60 posthypocristid_joins / nothing_or_distostyle postentocristid postectoentocristid, 61 endohypocristid_on_lower_molars / absent present, 62 posthypofossid_on_lower_molars / absent present, 63 entostylid__on_lower_molars_that_could_sometimes_be_linked_to_an_entocristylid / never frequently_present, 64 ectostylid__on_lower_molars / no_cingulid_ a_shallow_and_constant_cingulid_in_front_of_the_transverse_valley 'frequently developped cingulid in a /some stylid at least on m1', 65 ectocrystilid__on_lower_molars_ / no_ yes_even_if_variable, 66 'cingulid surrounding m/3 hypoconulid' / no_specimen_exhibiting_such_extension occasionally_bordering_the_labial_wall_, 67 'ectohypocristulid on M/3' / absent not_complete present_joigning_the_summit_of_hypoconulid, 68 'distostylid on M/1-M/2' / median lingual none, 69 'mesial part of Loop-like hypoconulid' / open_ pinched_, 70 entoconulid / no yes, 71 Number_of_upper_incisors / 3_ 2, 72 central_upper_incisor / 'morphologically similar to I2/I3 ' 'peg-like, morphologically different from others ' caniniform, 73 'I3/ reduced in size compared to I1/' / no_ yes_, 74 upper_canine_morphology / strong_with_circular_or_elliptic_cross_section_ 'strong and laterally compressed (blade-like)' premolariform, 75 canine_size_root / equivalent_to_slightly_longer_than_the_crown_ at_least_twice_the_size_of_the_crown_ prolonged_to_continous_growth_of_root_ prolonged_and_continuous_growth_of_crown, 76 dimorphic_upper_canine / no_ yes_, 77 'diastem C-P1/' / yes no, 78 'diastem P1/-P2/' / no yes, 79 number_of_upper_premolar / 4_ 5_ 3, 80 distolabial_crests_of_upper_premolars / simple with_a_maximum_of_two_accessory_cusps with_more_than_two_accessory_cusps_at_least_on_one_premolar, 81 'number of mesial crests on P1-3' / one two, 82 'disto-lingual basin in P2' / yes no, 83 'accessory cusp on disto-lingual cingulum of P3/' / none one_cingular_style 'protocone (surrounded by a cingulum)', 84 'Metacone on P3/' / no yes, 85 P3_root_pattern / 'one mesial root, two distal root not fused' one_mesial_root_and_fused_distal_ones, 86 P4_paracone / simple_with_crest complex_with_fossa very_complex_with_more_fossae, 87 'orientation of preparacrista on P4/' / mesial labial, 88 'postprotocrista on P4/' / absent present, 89 'Postprotocrista on P4/ joins' / base_of_paracone distostyle metastyle, 90 'preprotocrista on P4/ joins�' / mesiostyle base_of_the_paracone_then_mesiostyle parastyle, 91 'postectoprotocrista on P4/' / absent present, 92 P4_protocone_ / rounded crescentic, 93 'in lingual view protocone of P4/ is' / displaced_mesially median_, 94 'P4/ mesial margin' / concave convexe, 95 'strong development of distostyle on P4/' / no yes, 96 'mesial accessory cusp on P4/ that can be linked to mesiostyle' / no yes, 97 P4_metacone / absence presence, 98 P4_paracone_higher_than_the_protocone / slightly_higher_than_protocone much_higher_than_protocone, 99 P4_endoparacrista / absence_ presence, 100 'distal accessory cusp on postprotocrista of P4/ that can be linked to distostyle' / no yes, 101 height_of_lingual_cingulum_compared_to_unworn_protocone_height__on_upper_molars / one_third_ half_ no_cingulum, 102 'mesio-distal ribs development of labial cusps of upper molars' / almost_half_the_molar_lenght_ 'pinched (inferior to one third of molar lenght)' 'enlarged (superior to half the molar lenght) ', 103 postectoprotocrista / absent pr�sent, 104 postprotocrista / present absent, 105 Protocone_and_metaconule_junction_on_upper_molars / none_ 'premetaconule-postectoprotocrista ' 'premetaconule-postprotocrista ' postprotocrista_and_lingual_part_of_metaconule, 106 Premetacristule_divided_in_two_mesial_arms / no_ yes_, 107 ectometacristule_on_upper_molars / absent present_at_least_on_M1 not_frequent_and_only_on_M2_or_M3_, 108 postmetafossule / absent present, 109 secondary_cristule_labial_to_metaconule_eventually_an_endometacristule_or_enamel_knob / no_ yes_, 110 Distostyle_on_upper_molar / yes no, 111 distostyle_position_on_upper_molars_levels / metaconule_ metacone_, 112 secondary_ectometafossule_lingual_to_ectometacristule / absent_or_very_light present_mesially_at_least_on_M1_linked_to_ectometacristule, 113 paraconule_on_upper_molars / present_ absent, 114 'M2/ paraconule when present' / similar_in_size_with_protocone smaller_than_protocone, 115 postparacristule_extend_to_connect_ / none base_of_the_paracone transverse_valley, 116 preparacrista_connects_the_parastyle_ / 'no, separated by a groove ' yes_lingually yes_labially, 117 endoparacrista_on_upper_molars / absence presence, 118 ectoparafossa_on_upper_molars / no yes, 119 Ectocristyle / frequently_present absent, 120 premetacrista_and_postparacrista_connect / no_connection 'direct connection, forming a centrocrista' 'connection to mesostyle (direct, or indirect through ectocristyle)', 121 endometacrista_and_endometacristule_forming_a_transverse_crest_ / absence presence, 122 parastyle_development / enamel_knob smaller_or_equal_than_mesostyle larger_than_mesostyle, 123 premetacristule_invade_labial_part_of_the_transverse_valley / no yes, 124 position_of_metaconule_on_upper_molar / labial_side_of_the_protocone distal_side_of_the_protocone, 125 'M2/ metaconule' / similar_in_size_with_protocone smaller_than_protocone, 126 mesostyle_on_upper_molars / no yes, 127 mesostyle / enamel_knob half_to_the_size_of_labial_cusp larger_than_labial_cusp, 128 metastyle / reduced_to_enamel_knob_or_absent fully_developped_, 129 root_fusion_on_upper_molars / four_roots_with_occasional_fusion_close_to_cervix_the_apices_always_remaining_free fully_fused_lingual_roots three_roots, 130 lingual_cingulum_on_upper_molars / no yes developped_in_entostyle, 131 'hypocone on upper molars (at least M2/' / yes no, 132 'shape of M1/' / triangular quadrate, 133 'shape of M3/' / triangular quadrate, 134 'M3/ size' / 'Larger than M2/' 'equal in size with M2/' 'reduced (less than 60%)', 135 Mesiolingual_style_on_upper_molar_mesial_cingulum / no_ yes_, 136 symphysis_morphology_in_sagittal_section / elliptic dorsally_concave ventrally_concave, 137 'diastem C-P/1' / absent pr�sent, 138 bone_fusion_at_symphysis_in_adult_specimens / no_ yes_, 139 maximal_thickness_of_the_symphysis_in_sagittal_section / in_the_middle_part_ in_the_rostral_part_ in_the_nucal_part, 140 symphysis_extension / 'extends nuchally between C and P/1' 'extend nuchally between P/1 and P/3' 'extends nuchally to P/3', 141 number_and_position_of_main_external_foramen / numerous_ only_one_below_the_anterior_part_of_of_the_premolar_row_ 'two, one below the anterior part and the other below the posterior ', 142 'transverse constriction of mandible at C-p/1 diastema' / no_ yes_, 143 'P/1-P/2 diastema' / absent_ present_, 144 'P/2-P/3 diastema' / yes no, 145 Opening_of_internal_choanes / 'at M3/ ' 'nucal to M3/ ', 146 enamel_ornementation / no yes, 147 Schmelzmuster_composed_of_ / two_layers_ three_layers_ one_layer, 148 inner_radial_enamel_ / absent_ present_, 149 HSB_percent_of_Schmelzmuster / absent_ less_than_75%_ more_than_76%, 150 Outer_radial_enamel_ / less_or_equal_to_20%_ more_than_20%_, 151 Hsb_zone / thin_with_bands_always_less_than_100�m_ 'large (equal or more than 100�m) ', 152 'regular aspect (constant width)' / yes_ no_, 153 'HSB varaible (SD>20)' / no_ yes_, 154 HSB_angle_with_EDJ_ / '>70� ' '<70 ��', 155 orientation_of_HSB / straight_ bent_, 156 'HSB definition (decussation angle and size of transition zone)' / clear_ blurry_, 157 division_of_HSB / anastomosis_ bifurcation no_division, 158 HSB_configuration / curved_ transverse_, 159 synchronous_prism_undulation_on_horizontal_section / no_ yes_but_few__ yes_but_more_or_equal_to_4, 160 IPM_in_inner_portion / closed_sheath_ Inter_row_sheets_, 161 IPM_in_middle_portion / closed_sheath_ Inter_row_sheets_ no_IPM, 162 IPM_in_outer_portion / _closed_sheath no_IPM_, 163 prism_angle_with_EDJ_ / equal_or_more_than_60�_ less_than_60_�_ tend_to_diminish_in_the_inner_part, 164 prism_diameter / mean_between_3_and_3.9_�m small_diameter_mean_below_3_�m_ large_diameter_mean_above_or_equal_to_4, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 ] [ . . . . . . . . . . . . . . . . . . . ] Aepinacodon 0?10?0??????00100000100011010020011001110100001110122001011200011??1??00020001000100?111000111000000011000000000002201120101011{01}?11110011121101001?????????????????? Afromeryx {02}???????1????010000110101201001?011000110100000110122001000000011?01100?0?1100011000?1111001000000000110000000101--1011202010111021110021021210111001100011011101111 Amphimeryx 0011000110?0020011101020000001-0000010110100301001120111013100000001100??2???1000?2100012201110000002100200001-00022011201110110201111020001211000210---------000002 Amphiraghatherium ????????1????200111000000000112101101101000000101110000110001012002000???????0010121010110010010110012102011000001110{01}1{12}0001010021111000?021201{01}?1?????????????????? Anthracokeryx_tenuis 000-00001?1000000100000000010010000000010100000110101001000010000020000001{23}?01000100?10????10???00000010111000100111010202010110??11100?1021{12}11001?????????????????? Anthracokeryx_thailandicus ???????????????0000000000001001000100001000000011012100100000001002000????????????00?101000100000000?00021{01}000100121010201010111011110011021211101?????????????????? Anthracotherium_chaimanei ?0100101????????0?0000000001000010101001001100011010101010011110001001???01101000110?1111011000?000000101110101101210102010101110211101????{01}???101?????????????????? Anthracotherium_magnum 00100100000000100000000000010000101010010011000110102010100{01}110000200100001?110001101111111?000000000010111010{01}10121010202010111021110100101201101002000000001200000 Archaeomeryx 000-?0??1??002001110102000010100000010010000301001120101013000000001?000010?0120012{01}?0110201000000000101200001-01--1010202111111?111100?1??1?011?0?????????????????? Archaeopotamus_harvardi 001212011?0010200000112110-111210110111101113111001201{01}110300112112000000031010{12}002012011001101100110200200100{01}0{01}--0{01}10{012}00010100021111021102001{01}11?????????????????? Archaeopotamus_qeshta 001012011?0010200?00112110-11121011011110111311?001201111030?11211?00000103???0?0?201??110011?1?0??10200200100{01}011-0011000010{01}00?21?1?0211{01}200?0?1?????????????????? Bothriodon 0010000012000100000010001201000000100011000000011012200100120001?0011000020001000100?1111001010000000110011000000022011201010120011110011121111001?????????????????? Bothriogenys_andrewsi ??????????????????????????????????????01010000100011210100120001110?00??????????????????????????????0?????00???00??1?102?2???111?1????0??????????1?????????????????? Bothriogenys_fraasi ??????????0????0000010201112001?10100011000000011011101100011001110100???20?00001000?11101010000000?0010111000100121010202010111?11110011020111011?????????????????? Bothriogenys_gorringei {01}00?0011120001000000110012120010101000110000001{01}10122001000{12}0111110100000????????????1110101000000001110{12}1100010011101020201011101111001102121101111100001102??01?10 Bothriogenys_orientalis ???????????????0000011201211001010100001000001011011101100010102101100??????0000{01}000?11100011010000000101110?00001110102010101100111100010211?1111?????????????????? Brachyodus_aequatorialis 11110?1?1?0??1000000111011020011{01}0100011010000101111200000{01}200011111001212000000{01}000?1110101000000010111001000100121010202010111011111011120110111?????????????????? Brachyodus_depereti 21110?--?????100000000001102001100100011010000101112200101120000110100???????00????0?1111001000000000100000{01}00100121010202010111?11?1?0111?0?101?1?????????????????? Brachyodus_onoideus 111100111200010000000100110200101010001101000010111121010112000011010012120000001000?111010100000001011100000010012101020201011101111{01}011120110111202000011001201010 Bunophorus ????????????????1000000000000?200000001011000010110-001100000001000000???????????????000--001000000002003000001001000011000010-02100020??????????0?????????????????? Cebochoerus 000-000?10?0020100000000000001000000100100010100000-100100000102002000???20001000020010110000100000002002001000001010001020100-0001111020121?01100002000000000000001 Choeropotamus 000-???????0020001000000010100000010000100001000000-001000000012000000???0????000100?0011001010010000100201110000110010201010111?111110010010011?1001111100001000002 Chorora_hippopotamine ?0??1???1200102?0??0??????-1???10?10111101013111001{12}0111103001121??0?0???03????20?20120110011111?0110200200100{01}0???0010{02}00010100021???0??????????1?????????????????? Chororatherium ????????120??02?0??0??????-???????????110101?111001201?110??01???1{02}?00?????????20?101??110011?1?0011?200200100{01}00110110{02}00010100021???0??????????1?????????????????? Dacrytherium 0???00??1210020001000000110201210010100111000000010-1101012000100001000??20?100000201110-201011000010100000001-001020112021101112011100000?11001?0?????????????????? Dichobune 000-00??12?0010010000010020000000000100010000000010-000100001002000000???????1000?1?0000-0001100000000010000000001010011020010-02001020000?120100001200110010??0??02 Diplobune 001000011210010001100000110211010000101001000010110-1101112200000001000212001000012011112201010000002100000001-001020012011101112011100100012001001111000{01}0001000012 Elomeryx_borbonicus 00100010120001000001101012010011101000110000001{01}111220010112000211011000011100000000?111{01}0010000000001{01}0100100100122011201010111021110021021111111112000011001111110 Elomeryx_crispus 0010001012000100000{01}101010010011101000010000000110102001011210011101{01}{01}00011100000000?111100100000000011011110010012201120201011101111002101?111111?????????????????? Entelodon 001{01}0001000000110100000000000000000010101100101101100011000010000000-00000??1100012010010-000100000012010-00000001001010000010-0110102020021100001111110000021?00002 Epirigenys ????0111????????0?00010111-1002{01}1111100100{01}{01}00011012100010000112?1?100???????????????1111001101100011010{12}11010010120010201010110?111100??????????1002000011021101110 Gobiohyus ????????????????100000001001002001001000100000001110000100001001001000????????????2?01011001010000001001000001-001010111020100-02111110??????????0?????????????????? Gujaratia 000-000?1210010010000000000000000000001011000010010-00110000000100100000011?11000120?00112000100010000003000001001101011100010-0201001000011??0100210---------000001 Hallebune ???????????????011101020000011?10?001010010000101110000100001001002000?????????00?21?101100100001?0012103000000001110111000{01}10-0?10{01}010???????11?0?????????????????? Heptacodon 000-????110000100000002000010000001010110000000110101111000110000000000?001110000110?1111011000000000010101000110111010202010121?211101001211001?1?????????????????? Hexaprotodon 001212011?0010200000112110-111210110111101113111001{12}01{01}11030{01}1121120000000310{01}0{12}002012011001101100110200200100{01}0{01}--0{01}10{012}00010100021111021102001{01}11111111111121212111 Homacodon ???????????????????0??????????????????10010000100110000100001002000000?????????00?10000110001100000000103000000000010001020000-02101020??????????0?????????????????? Indohyus ????????????????0??000????01??0000001011100000101110000102001000001200?????????00?001000--00000001100200201001-001101011100110-0211?100????????????????????????????? Kenyapotamus_coryndonae ?0??12011?00102?000?112110-11121011{01}110100{01}1{13}101{01}012101110000112112000?????????10?10110110011011001112{01}020{01}100{01}10110110{02}000101000211110??1?2?0?{01}?120201011100121101{02} Kenyapotamus_ternani 00101???1??010200?00?12?10-??0????????0100{01}1{13}?01{01}01210111000011?1?2000?????????{01}0?10?2011001101?00111210201100{01}10110110{02}000101000211110{01}11020010?1?????????????????? Kenyasus 001{02}0100130100210000000000-000-00000101100012001000-11110010100100000001102?01000?00?10110010100100022010011000101001010100100-01011100201?1?00100112000000001001000 Khirtaria ????????????????0000000000010000001010111001-0111110001102001000001{02}0{01}??????????????????????????????02000010100001101011100110-02110021??????????1?????????????????? Libycosaurus_anisae {01}012100{01}11010021000110101201001000101011010000101112100101200002110110???020?0121000?1111001110000000100002000101--20112010101111?1110021020210111111111111121112110 Libycosaurus_bahri 201210011101002100011010120100100010101101000010111210010120000211011000102?0012100011111001010000000100002000101--201120101011111111002112021{01}111210---------112120 Lophiomeryx 001200??1????20011100020000101-000001011010030100112010101300002000110---2?00120011000112201110000000100200001-01--2011201110111?111100010?0201110?????????????????? Merycoidodon ????????010000100000002011011100101010010100-00001122101013200120001??00001?0000100000112201000000002000200001-01--2001202110111?21110010??1200110?????????????????? Merycopotamus_dissimilis 000-0???11010021000110101201001000101011010000101112100101200002110110???0110001100011111001010000000100002000101--2011201010111121110021101200111111111111121112112 Merycopotamus_nanus 000-0???110100110001101012010010001010110100001011121{01}010120000211011000001?0001100011111001010000000100000000101--20112010101111211100210011101110011111111211111?2 Microbunodon 000-00001110000001000010000100100000000100000001101211010000000100200000013110000100?1110001010000000100200000100121010201010111011110011121211101002101000000000000 Mixtotherium ????????1200001011100020000111-?0000100010000110110-101100010102000100???01?10000?2101111001000010000100200001-001020002020111112211110?1?????11?011100000000??00002 Morotochoerus ????????????????0???01????-1??2?0{01}11?10101111101101{01}100110000112102000??????????????????????????????{01}200201100000110010{02}00010100?111100??????????1?????????????????? Myaingtherium 000-000?0?????1100000000000100?000101001001100001110100100001000002001???????????????101001100000000?2101110000001100111000101001211101010?2?101?1?????????????????? Palaeochoerus 001{02}0100130110200000000000-000-00000101100110000110-1001101000010000000?1021{01}00000101000-0001101100022010011100001000010100100-01011110211?1?001?00020000000{01}0000000 Paroxacron 000-000?1????2?111101020000201000000100110000010010-0101012000000001-1???????000012101111201010001002101000001-00002001201101111201112020011100000210---------000002 Perchoerus 001{02}0?0013011020000000?0??-0?0?00000101100110000100-?1?1001010010000000??02?{01}1000?00?000-000010010002201001000?001000011000100-0?011110?11??2?11?0?????????????????? Siamotherium 0???00??0000?00000000000000100000010010100000000001010010000110000200000001?0?0001{01}01100-000010000001000210000100111010100010100011110000021200101002000000001000002 Sivameryx 000-????110?00110001101012010010001000010000010110122001012{02}0101110110??????00?11000?1110001000000000110000000100112011201010111?211100210202101?1210---------112110 Xiphodon ????????????????11101020000211010000101101000010110-200101100001000110???????001010111111201010000002100000001-000220012011101102211111???????01?01111000100{01}1100012 ; END; [ The following blocks are output data for analysis step 21825 ] BEGIN TREES; TITLE Hippopotamidae_late_Miocene_Baynunah_Formation; LINK TAXA = TaxaForAnalysisStep21825; TREE Strict = [&R] (Homacodon,((Gujaratia,Bunophorus),(Entelodon,(Khirtaria,Indohyus))),(Dichobune,(Gobiohyus,(((Amphiraghatherium,Hallebune),(Mixtotherium,((Diplobune,(Dacrytherium,Merycoidodon)),(Xiphodon,(Paroxacron,(Amphimeryx,(Lophiomeryx,Archaeomeryx))))))),((Cebochoerus,(Kenyasus,(Palaeochoerus,Perchoerus))),(Choeropotamus,(Siamotherium,((Anthracokeryx_tenuis,(Myaingtherium,(Heptacodon,(Anthracotherium_chaimanei,Anthracotherium_magnum)))),(Anthracokeryx_thailandicus,(Microbunodon,((Afromeryx,(Sivameryx,(Merycopotamus_nanus,(Merycopotamus_dissimilis,(Libycosaurus_anisae,Libycosaurus_bahri))))),((Bothriodon,Aepinacodon),(Elomeryx_crispus,Elomeryx_borbonicus),((Bothriogenys_andrewsi,Brachyodus_aequatorialis,Brachyodus_onoideus,Brachyodus_depereti),(Bothriogenys_fraasi,Bothriogenys_gorringei,(Bothriogenys_orientalis,(Epirigenys,(Morotochoerus,Kenyapotamus_coryndonae,Kenyapotamus_ternani,(Chororatherium,(Chorora_hippopotamine,(Archaeopotamus_qeshta,(Archaeopotamus_harvardi,Hexaprotodon))))))))))))))))))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr128345] END;