#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on March 30, 2026; 13:36 GMT TreeBASE (cc) 1994-2008 Study reference: Barden P., Perrichot V., & Wang B. 2020. Specialized predation drives aberrant morphological integration and diversity in the earliest ants. Current Biology, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S26540] [ The following blocks are input data for analysis step 26811 ] BEGIN TAXA; TITLE TaxaForAnalysisStep26811; DIMENSIONS NTAX=46; TAXLABELS Acanthoponera_minor Amblyopone_pallipes Aneuretus Anomalomyrma_sp Apomyrma_stygia Aquilomyrmex Brownimecia_clavata Camelomecia_janovitzi Cerapachys_sexspinus Ceratomyrmex Chalybion_californicum Chonidris_insolita Dhagnathos Formica_fusca Gerontoformica_gracilis Gerontoformica_magnus Gerontoformica_pilosus Gerontoformica_spiralis Haidomyrmex_scimitarus Haidomyrmodes_mammuthus Haidoterminus_cippus Heterogyna Hypoponera_opacior Kyromyrma_neffi Lasius_californicus Leptanilla_swani Leptanilloides_nomada Leptogenys_diminuta Linguamyrmex_vladi Martialis_heureka Metapolybia_cingulata Myanmyrma_gracilis Myrmecia_nigriceps Myrmica_americana Nothomyrmecia_macrops Opamyrma_hungvuong Paraponera_clavata Platythyrea_punctata Pogonomyrmex_californicus Proceratium_stictum Protoceratomyrmex Scolia_verticalis Sphecomyrma_freyi Tatuidris_tatusia Tetraponera_punctulata Zigrasimecia_tonsora ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M55024] TITLE Formicidae; LINK TAXA = TaxaForAnalysisStep26811; DIMENSIONS NCHAR=65; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7" MISSING=? GAP= -; CHARSTATELABELS 1 Occipital_foramen, 2 'Mandible Orientation*', 3 'Clyeal Pegs*', 4 'Labral Pegs*', 5 Posterior_Clypeal_Sulcus, 6 'Clypeus Development*', 7 Subgenal_sulcus, 8 'Post Antennal Marg*', 9 'Torulus*', 10 'Frontal Lobe*', 11 'Frontal Lobe Notched*', 12 'Scape Length*', 13 'Antennal Segments*', 14 'Ocelli*', 15 'Compound Eye*', 16 Bidentate_Mandibles, 17 Basal_flange_present, 18 Inner_margin_toothed, 19 Lat_eye_position, 20 Eye_abutting_vertex, 21 Head_depth, 22 Clypeus_concave, 23 Clypeal_ridges_with_teeth, 24 'Clypeal Peg Structure*', 25 'Frontal Carina*', 26 Frontal_Triangle, 27 Frontal_triangle_fused, 28 Trigger_Setae, 29 Eye_shape, 30 Clypeal_Process, 31 Clypeal_brush, 32 Clypeal_Stalk, 33 Clypeal_Lobe, 34 Anterior_margin_of_clypeus, 35 Antennal_socket_vertex, 36 Pronotal_collar_lip, 37 Propodeal_spiracle_shape, 38 'Metapleural Gland*', 39 'Metapleural gland opening*', 40 Propodeal_Shelf, 41 Metatibial_setae, 42 'Post Pronotal Development*', 43 'Pronotal Lobe*', 44 'Propleuron*', 45 'Promesonotal Suture*', 46 'Metanotum*', 47 'Propodeal Suture*', 48 'Trochantellus*', 49 'Pretarsal Claw*', 50 'Pygidial Teeth*', 51 Subpetiole_spicule, 52 Subpetiole_lamella, 53 'Petiole*', 54 'Petiole Peduncle*', 55 'Petiole Expansion*', 56 'Post Petiole Face*', 57 'Petiole Fusion*', 58 'Helcium*', 59 'Third Abd Segment*', 60 'Sternal Abd Projection*', 61 Keel_flat_face, 62 'Constriction bt abd 3 and 4*', 63 'Abd seg fusion*', 64 'Sting*', 65 'Acidopore*', ; MATRIX [ 10 20 30 40 50 60 ] [ . . . . . . ] Acanthoponera_minor 100001010202{12}020--0000--10-000---010010001002100101011201100?1010 Amblyopone_pallipes 101000011212{12}020--0000-000-000---010010001001110011111110100?1010 Aneuretus 1000010100-2{12}020--0000--10-010---100010001012110000011101100?0010 Anomalomyrma_sp 1001000100-2{12}000----00--00-0-0---010011001011100001010210011?0210 Apomyrma_stygia 1001000100-2{12}001----00--00-0-0---010010001011200000011200000?0310 Aquilomyrmex 0100111010-2{12}?200011010-00-00100-11111001101??11100011101100-1110 Brownimecia_clavata 10??010??0-2{12}020--0000--00-000---?1001000101211010??1110?100?1110 Camelomecia_janovitzi 100001001301{12}120--0000--00-010---110010001011?11101011101101??110 Cerapachys_sexspinus 100000010102{12}000----00--10-0-0---010011001?00200011010200100?0010 Ceratomyrmex 0100110010-1{12}120100110--01110101111011000101101110101110110100110 Chalybion_californicum 0000010000-0{12}120--0100--00-010---010-0-0011120111000100-0000?0110 Chonidris_insolita 0100010000-???201100011-00-01100-101110111011?1110111110110111110 Dhagnathos 0100111100-1{12}?201100011-00-01100-101110111011?1110111110110111110 Formica_fusca 1000010000-2{12}020--0000--10-010---010110001012110000010300000?0001 Gerontoformica_gracilis 1010010000-1{12}121--0000-120-000---01001000101101110011110110100110 Gerontoformica_magnus 1010010000-1{12}121--0100-120-000---01001000101101110??1110110100110 Gerontoformica_pilosus 1010010000-1{12}121--0000-120-000---01001000101101110011110110101110 Gerontoformica_spiralis 1010010000-1{12}121--0000-120-000---01001000101101110011110110100110 Haidomyrmex_scimitarus 0100110000-1{12}020100110--01010110-110010001011011100011101100-0110 Haidomyrmodes_mammuthus 0100?1?000-1{12}120100110--01010110-110??000101101?10001110?100-0110 Haidoterminus_cippus 0100110000-2{12}020100110--01010110-110010001011011100011101100-0110 Heterogyna 0000010000-0{13}120--0100--00-010---01010-00111101110000-----00?0110 Hypoponera_opacior 100001002112{12}020--0000--10-000---01001?001011110200001201000?1110 Kyromyrma_neffi 1000010000-2{12}020--0000--10-000---01001000101211000??10?00000?0001 Lasius_californicus 1000010000-2{12}020--0000--10-000---010010001012110000010300000?0001 Leptanilla_swani 1000000100-2{12}000----00--00-0-0---010011001011200000010200011?0010 Leptanilloides_nomada 100000001002{12}000----00--10-0-0---010011001000200001110200010?0010 Leptogenys_diminuta 100001002112{12}020--0000--10-000---01011?001011110201011201001?1110 Linguamyrmex_vladi 0100110000-1{12}120100110--01110101111011000101101110001110110101110 Martialis_heureka 1000101110-2{12}000----00--00-0-0---010010001001111000011100100?1010 Metapolybia_cingulata 0000010000-1{12}120--0100--00-010---01000-001002010000011100100?0110 Myanmyrma_gracilis 1011?1???0-1{12}??1--0?00-1?0-0?0---1100100010?101110??1110110??1110 Myrmecia_nigriceps 100001011102{12}120--1000--10-000---010111001012110101011101101?1110 Myrmica_americana 100001010202{12}020--0000--10-000---010010001012110001011100110?0010 Nothomyrmecia_macrops 100001011102{12}020--0000--10-000---010111001012110101011101101?0110 Opamyrma_hungvuong 1001000100-2{12}001----00--00-0-0---010010001011210000011200100?0010 Paraponera_clavata 100001001102{12}020--0000--10-000---010110001002110111111200100?1010 Platythyrea_punctata 100001002112{12}020--0000--10-000---01011?001001210201010211000?1110 Pogonomyrmex_californicus 100001010202{12}020--0000--10-000---010110001012210000011100110?0010 Proceratium_stictum 100000010102{12}010--0000--00-0-0---01001?001002210001011210000?1010 Protoceratomyrmex 0100010010-1{12}020100110--01010101011001000101101110001110110101110 Scolia_verticalis 0000010000-0{13}120--0100--00-010---010?0-000002011000011110100?0110 Sphecomyrma_freyi 1000010000-1{12}121--0000--00-000---010010001011011100?1110?100?0110 Tatuidris_tatusia 1000101000-27020--0000--00-000---010011001000200000010210010?1010 Tetraponera_punctulata 100000001102{12}020--0000--10-000---010011001011110101011101100?0110 Zigrasimecia_tonsora 1011010010-1{12}021--0000-100-000---110110001012011101111100100?0110 ; END; [ The following blocks are output data for analysis step 26811 ] BEGIN TREES; TITLE 'Formicidae Parsimony_ImpliedWeights_ConstrainedTopology'; LINK TAXA = TaxaForAnalysisStep26811; TREE tagged_tree = [&R] (Metapolybia_cingulata,((Scolia_verticalis,(Chalybion_californicum,Heterogyna)),((Camelomecia_janovitzi,(Brownimecia_clavata,((Martialis_heureka,(Opamyrma_hungvuong,(Leptanilla_swani,Anomalomyrma_sp))),(((Apomyrma_stygia,Amblyopone_pallipes),(Proceratium_stictum,((Tatuidris_tatusia,Paraponera_clavata),(Platythyrea_punctata,(Leptogenys_diminuta,Hypoponera_opacior))))),((Cerapachys_sexspinus,Leptanilloides_nomada),((Aneuretus,(Tetraponera_punctulata,(Myrmecia_nigriceps,Nothomyrmecia_macrops))),((Lasius_californicus,Formica_fusca,Kyromyrma_neffi),(Acanthoponera_minor,(Pogonomyrmex_californicus,Myrmica_americana)))))))),(Sphecomyrma_freyi,((Gerontoformica_magnus,Gerontoformica_gracilis,Gerontoformica_pilosus,Gerontoformica_spiralis),(Zigrasimecia_tonsora,Myanmyrma_gracilis)))),(((Haidomyrmodes_mammuthus,(Haidomyrmex_scimitarus,Haidoterminus_cippus)),(Protoceratomyrmex,(Ceratomyrmex,Linguamyrmex_vladi))),(Aquilomyrmex,(Dhagnathos,Chonidris_insolita)))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr124451] END; BEGIN TREES; TITLE Formicidae_Parsimony_ImpliedWeights_Unconstrained_Topology; LINK TAXA = TaxaForAnalysisStep26811; TREE tnt_1 = [&R] (Metapolybia_cingulata,(Scolia_verticalis,((Chalybion_californicum,Heterogyna),(((Camelomecia_janovitzi,(Brownimecia_clavata,(((Paraponera_clavata,(Acanthoponera_minor,((Amblyopone_pallipes,(Martialis_heureka,(Proceratium_stictum,((Cerapachys_sexspinus,((Leptanilla_swani,Anomalomyrma_sp),(Tatuidris_tatusia,Leptanilloides_nomada))),(Apomyrma_stygia,Opamyrma_hungvuong))))),((Aneuretus,(Lasius_californicus,Formica_fusca,Kyromyrma_neffi)),(Pogonomyrmex_californicus,Myrmica_americana))))),(Hypoponera_opacior,(Platythyrea_punctata,Leptogenys_diminuta))),(Tetraponera_punctulata,(Myrmecia_nigriceps,Nothomyrmecia_macrops))))),(Sphecomyrma_freyi,((Gerontoformica_magnus,Gerontoformica_gracilis,Gerontoformica_pilosus,Gerontoformica_spiralis),(Zigrasimecia_tonsora,Myanmyrma_gracilis)))),(((Haidomyrmodes_mammuthus,(Haidomyrmex_scimitarus,Haidoterminus_cippus)),(Protoceratomyrmex,(Ceratomyrmex,Linguamyrmex_vladi))),(Aquilomyrmex,(Dhagnathos,Chonidris_insolita))))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr124450] END; BEGIN TREES; TITLE 'Formicidae Parsimony_ImpliedWeights_ConstrainedTopology'; LINK TAXA = TaxaForAnalysisStep26811; TREE tnt_1 = [&R] (Metapolybia_cingulata,((Scolia_verticalis,(Chalybion_californicum,Heterogyna)),((Camelomecia_janovitzi,(Brownimecia_clavata,((Martialis_heureka,(Opamyrma_hungvuong,(Leptanilla_swani,Anomalomyrma_sp))),(((Apomyrma_stygia,Amblyopone_pallipes),(Proceratium_stictum,((Tatuidris_tatusia,Paraponera_clavata),(Platythyrea_punctata,(Leptogenys_diminuta,Hypoponera_opacior))))),((Cerapachys_sexspinus,Leptanilloides_nomada),((Aneuretus,(Tetraponera_punctulata,(Myrmecia_nigriceps,Nothomyrmecia_macrops))),((Lasius_californicus,Formica_fusca,Kyromyrma_neffi),(Acanthoponera_minor,(Pogonomyrmex_californicus,Myrmica_americana)))))))),(Sphecomyrma_freyi,((Gerontoformica_magnus,Gerontoformica_gracilis,Gerontoformica_pilosus,Gerontoformica_spiralis),(Zigrasimecia_tonsora,Myanmyrma_gracilis)))),(((Haidomyrmodes_mammuthus,(Haidomyrmex_scimitarus,Haidoterminus_cippus)),(Protoceratomyrmex,(Ceratomyrmex,Linguamyrmex_vladi))),(Aquilomyrmex,(Dhagnathos,Chonidris_insolita)))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr124452] END; BEGIN TREES; TITLE Formicidae_Parsimony_ImpliedWeights_Unconstrained_Topology; LINK TAXA = TaxaForAnalysisStep26811; TREE tagged_tree = [&R] (Metapolybia_cingulata,(Scolia_verticalis,((Chalybion_californicum,Heterogyna),(((Camelomecia_janovitzi,(Brownimecia_clavata,(((Paraponera_clavata,(Acanthoponera_minor,((Amblyopone_pallipes,(Martialis_heureka,(Proceratium_stictum,((Cerapachys_sexspinus,((Leptanilla_swani,Anomalomyrma_sp),(Tatuidris_tatusia,Leptanilloides_nomada))),(Apomyrma_stygia,Opamyrma_hungvuong))))),((Aneuretus,(Lasius_californicus,Formica_fusca,Kyromyrma_neffi)),(Pogonomyrmex_californicus,Myrmica_americana))))),(Hypoponera_opacior,(Platythyrea_punctata,Leptogenys_diminuta))),(Tetraponera_punctulata,(Myrmecia_nigriceps,Nothomyrmecia_macrops))))),(Sphecomyrma_freyi,((Gerontoformica_magnus,Gerontoformica_gracilis,Gerontoformica_pilosus,Gerontoformica_spiralis),(Zigrasimecia_tonsora,Myanmyrma_gracilis)))),(((Haidomyrmodes_mammuthus,(Haidomyrmex_scimitarus,Haidoterminus_cippus)),(Protoceratomyrmex,(Ceratomyrmex,Linguamyrmex_vladi))),(Aquilomyrmex,(Dhagnathos,Chonidris_insolita))))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr124449] END;