#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on January 19, 2026; 11:40 GMT

TreeBASE (cc) 1994-2008

Study reference:
Tsutsumi C., Yatabe Y., Hirayama Y., Chiou W., & Kato M. 2021. Molecular analyses
of the disjunctly distributed Osmunda regalis and O. japonica (Osmundaceae), with
particular reference to introgression and hybridization. Plant Systematics and
Evolution, .

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S26988]

[ The following blocks are input data for analysis step 27365 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep27365;
        DIMENSIONS NTAX=14;
        TAXLABELS
            Osmunda_japonica_J1a
            Osmunda_japonica_J1b
            Osmunda_japonica_J2J3J4J5
            Osmunda_japonica_J6J7J8
            Osmunda_lancea_L1L2
            Osmunda_lancea_L3
            Osmunda_regalis_R10
            Osmunda_regalis_R11R12
            Osmunda_regalis_R16b
            Osmunda_regalis_R1R2R4
            Osmunda_regalis_R3
            Osmunda_regalis_R5R6R7R13R14R15R16aR17R18
            Osmunda_regalis_R8R9
            Plenasium_banksiifolium.seq
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M56338]

	TITLE Osmundaceae_L261;
	LINK TAXA = TaxaForAnalysisStep27365;
	DIMENSIONS NCHAR=322;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                                     10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320  ]
[                                                      .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .  ]
Osmunda_japonica_J1a                          GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCTCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATCTAGCAACTATTCCAATCACGGGCAC
Osmunda_japonica_J1b                          GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCTCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_japonica_J2J3J4J5                     GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCTCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTTTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATCTAGCAACTATTCCAATCACGGGCAC
Osmunda_japonica_J6J7J8                       GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCTCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACGATTCCAATCACGGGCAC
Osmunda_lancea_L1L2                           GGTCTAGCAGCTGAGATAATTGGGACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCTCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTGTGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_lancea_L3                             GGTCTAGCAGCTGAGATAATTGGGACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCTCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R10                           GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGTGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCGAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATCACTGAGCTTGTTATGTTTGCGCAGGTACTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R11R12                        GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATCACTGAGCTTGTTATGTTTGCGCAGGTACTAGCGCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R16b                          GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGGGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGGGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTAATGTCTGCGCAGGTACTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R1R2R4                        GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCTCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGTGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R3                            GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCTCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGTGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGACAAGTAAGGTCGTGTTGCGAGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R5R6R7R13R14R15R16aR17R18     GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGCGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGGGAGAGAGAG--TGAGTGAGAGGAGATGACTGAGCTTGTAATGTCTGCGCAGGTACTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Osmunda_regalis_R8R9                          GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTATACACCGTCTTCTCTGCTACAGACCCAAAGAGGAACGCTCGGGATTCTCACGTTCCAGTATGTTAATCTGCCCCTGCGAGCTGGTTTGAGTGGAGTGAGAAATTCTGCTCGGTCTTGCTGTGTAAAGAGCAAAGATGAGGGCAAGTAAGGTCGTGTTGCGAGAGAGAGAGAGTGAGTGAGAGGAGATGACTGAGCTTGTTATGTTTGCGCAGGTTCTAGCTCCACTCCCCATCGGCTTCGCAGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
Plenasium_banksiifolium.seq                   GGTCTAGCAGCTGAGATAATTGGAACCTTCGTTCTCGTGTACACCGTCTTCTCTGCTACCGACCCCAAGAGGAACGCCCGGGATTCTCACGTTCCAGTATGTCAATCTGCCCCTGCAAGCTGGTTTGTGTGGAGTGAA--------CTCGGTGTTGCTGTGTAAAGAGCAAAGATGAGGCCTAGTGAGGTCGTGTTGAGAGAGAGAGAGA-T-AGAGAGAG-AGATGACTGAGCTTGTTATGTTTGTGCAGGTACTAGCTCCACTTCCCATCGGCTTCGCCGTGTTCATGGTGCATTTAGCAACTATTCCAATCACGGGCAC
;
END;

[ The following blocks are output data for analysis step 27365 ]
BEGIN TREES;
      TITLE Osmundaceae_L261;
      LINK TAXA = TaxaForAnalysisStep27365;
      TREE con_50_majrule = [&R] (Plenasium_banksiifolium.seq:0.0303775,((Osmunda_regalis_R10:0.008759,Osmunda_regalis_R11R12:0.007151):0.006287,(Osmunda_regalis_R5R6R7R13R14R15R16aR17R18:0.003167,Osmunda_regalis_R16b:0.006228)1.00:0.01308,((Osmunda_regalis_R1R2R4:0.003094,Osmunda_regalis_R3:0.006229,Osmunda_regalis_R8R9:0.003112)0.97:0.008872,(Osmunda_japonica_J6J7J8:0.006292,Osmunda_japonica_J1b:0.003164,(Osmunda_japonica_J2J3J4J5:0.006357,Osmunda_japonica_J1a:0.003241)0.96:0.006227,(Osmunda_lancea_L1L2:0.006347,Osmunda_lancea_L3:0.003228)0.96:0.006248)0.94:0.007206)0.81:0.006491)0.83:0.0303775);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr126300]


END;
BEGIN TREES;
      TITLE Osmundaceae_L261;
      LINK TAXA = TaxaForAnalysisStep27365;
      TREE con_50_majrule = [&R] (Osmunda_regalis_R10:0.008759,Osmunda_regalis_R11R12:0.007151,(((Osmunda_japonica_J6J7J8:0.006292,(Osmunda_japonica_J2J3J4J5:0.006357,Osmunda_japonica_J1a:0.003241):0.006227,(Osmunda_lancea_L1L2:0.006347,Osmunda_lancea_L3:0.003228):0.006248,Osmunda_japonica_J1b:0.003164):0.007206,(Osmunda_regalis_R1R2R4:0.003094,Osmunda_regalis_R3:0.006229,Osmunda_regalis_R8R9:0.003112):0.008872):0.006491,(Osmunda_regalis_R5R6R7R13R14R15R16aR17R18:0.003167,Osmunda_regalis_R16b:0.006228):0.01308,Plenasium_banksiifolium.seq:0.060755):0.006287);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr126301]


END;