#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on January 19, 2026; 11:39 GMT

TreeBASE (cc) 1994-2008

Study reference:
Tsutsumi C., Yatabe Y., Hirayama Y., Chiou W., & Kato M. 2021. Molecular analyses
of the disjunctly distributed Osmunda regalis and O. japonica (Osmundaceae), with
particular reference to introgression and hybridization. Plant Systematics and
Evolution, .

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S26988]

[ The following blocks are input data for analysis step 27367 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep27367;
        DIMENSIONS NTAX=12;
        TAXLABELS
            Claytosmunda_claytoniana_CL2
            Osmunda_japonica_J1J2J3J4J5J6J7J8
            Osmunda_lancea_L1L2L3
            Osmunda_regalis_R10
            Osmunda_regalis_R111R12
            Osmunda_regalis_R13aR14a
            Osmunda_regalis_R13bR17R18
            Osmunda_regalis_R14b
            Osmunda_regalis_R15
            Osmunda_regalis_R16
            Osmunda_regalis_R1R2R3R4
            Osmunda_regalis_R5R6R7R8R9
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M56340]

	TITLE Osmundaceae_L418;
	LINK TAXA = TaxaForAnalysisStep27367;
	DIMENSIONS NCHAR=559;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                             10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510       520       530       540       550         ]
[                                              .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         ]
Claytosmunda_claytoniana_CL2          CCAGACTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATTGAAGACACTGGCTGTGATGCCCCCATTCCCCTCCCTAATGTTTCCAGCAAGATCCTTGCAAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAGGATGAAGTCAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGCTTGGTTGGGTACTTGCTAAGCTTTTTGGTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTCTGTACATTC-GTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGTTTTAAAGTTTGTACATTCGGGAGTCTTTGCTGACCTTTCCCTTCCTATTGTTATGCACAGGCGGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGTAATACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_japonica_J1J2J3J4J5J6J7J8     TCAGATTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTCAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGTTTTAAAGTTTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCGGCAAATTACCTCAACATCATGAATCTTTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGTAGTACTTTCAACATCAAGAACGATTTTACTCCAGAGGAAGAGGAA
Osmunda_lancea_L1L2L3                 TCAGATTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAAGAGGAAGTCAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGTTTTAAAGTTTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCGGCAAATTACCTCAACATCATGAATCTTTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGTAGTACTTTCAACATCAAGAACGATTTTACTCCAGAGGAAGAGGAA
Osmunda_regalis_R10                   TCAGATTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGCGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACGCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGCGTTAAAGTCTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGCAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R111R12               TCAAATTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGACCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGCGTTAAAGTCTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGCAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R13aR14a              TCAGACTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCTTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTGTTGTTTCTTGCTTGTGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGTTTTAAAGTTTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGCAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R13bR17R18            TCAGACTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGACGATGGTTGCGGTCCTTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTGTTGTTTCTTGCTTGTGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGTTTTAAAGTTTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGTAGTACTTTCAACATCAAAAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R14b                  TCAGACTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCTTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTGTTGTTTCTTGCTTGTGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGTTTTAAAGTTTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGTAGTACTTTCAACATCAAAAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R15                   TCAGACTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATTCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTCAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTGTTTTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGCTTTAAAGTCTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGCAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R16                   TCAGACTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTAGATTCTTTTCAAATCTTTTGCTCAGCCGTTGTCGTTCCATGTTTTAAAGTCTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCGGCAAATTACCTCAACATCATGAATCTTTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGTAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R1R2R3R4              TCAGATTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGTCTGGTGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGCGTTAAAGTCTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGCAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
Osmunda_regalis_R5R6R7R8R9            TCAGATTGTCGCCTTTGAATCCCAGACCATCAAGAACATGATCGAAGACACTGGCTGTGATGCCCCTATTCCCCTCCCTAACGTTTCCAGCAAGATCCTTGCCAAGGTCATTGAATTTTGTAAGTTCCATGTCAACTCTAAGAAGACTGGTGATGATGGTTGCGGTCCCTCCGAGGAGGAAGTTAAGTCCTTCGACGCTGAATTCGTGAAAGTTGATCAGCAAACCCTCTTCGACCTCATCCTGGTATGTTT------------------CTTGGTCTTCTTGTTTCTTGCTTGAGCATGATCTTTCGTTGCCTTTGTACATTCTTTTCAAATCTTTTGCTCAGCCGTTGTTGTTCCATGCGTTAAAGTCTGTAGATTCGGGAGTCTTTGCTGACTTTCCCCTT-CTATTGTTATGCACAGGCAGCAAATTACCTCAACATCATGAATCTGTTGGATCTGACCTGCCAGACCGTGGCAGACATGATCAAAGGCAAGACGCCAGAGGAGATTCGCAGTACTTTCAACATCAAGAACGATTTTACTCCAGAAGAAGAGGAA
;
END;

[ The following blocks are output data for analysis step 27367 ]
BEGIN TREES;
      TITLE Osmundaceae_L418;
      LINK TAXA = TaxaForAnalysisStep27367;
      TREE con_50_majrule = [&R] (Claytosmunda_claytoniana_CL2:0.0259805,(Osmunda_regalis_R16:0.00818,(Osmunda_japonica_J1J2J3J4J5J6J7J8:0.002155,Osmunda_lancea_L1L2L3:0.004306)0.98:0.008464,((Osmunda_regalis_R10:0.006556,Osmunda_regalis_R5R6R7R8R9:0.002157,Osmunda_regalis_R1R2R3R4:0.004376,Osmunda_regalis_R111R12:0.006534):0.007798,(Osmunda_regalis_R15:0.009861,(Osmunda_regalis_R13aR14a:0.002477,(Osmunda_regalis_R14b:0.002232,Osmunda_regalis_R13bR17R18:0.0043)0.99:0.006354)1.00:0.010326)0.58:0.005193)0.97:0.009273)0.96:0.0259805);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr126304]


END;
BEGIN TREES;
      TITLE Osmundaceae_L418;
      LINK TAXA = TaxaForAnalysisStep27367;
      TREE con_50_majrule = [&R] (Osmunda_regalis_R10:0.006556,Osmunda_regalis_R5R6R7R8R9:0.002157,Osmunda_regalis_R1R2R3R4:0.004376,Osmunda_regalis_R111R12:0.006534,(((Osmunda_japonica_J1J2J3J4J5J6J7J8:0.002155,Osmunda_lancea_L1L2L3:0.004306):0.008464,Osmunda_regalis_R16:0.00818,Claytosmunda_claytoniana_CL2:0.051961):0.009273,(Osmunda_regalis_R15:0.009861,(Osmunda_regalis_R13aR14a:0.002477,(Osmunda_regalis_R14b:0.002232,Osmunda_regalis_R13bR17R18:0.0043):0.006354):0.010326):0.005193):0.007798);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr126305]


END;