#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on January 19, 2026; 11:39 GMT

TreeBASE (cc) 1994-2008

Study reference:
Tsutsumi C., Yatabe Y., Hirayama Y., Chiou W., & Kato M. 2021. Molecular analyses
of the disjunctly distributed Osmunda regalis and O. japonica (Osmundaceae), with
particular reference to introgression and hybridization. Plant Systematics and
Evolution, .

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S26988]

[ The following blocks are input data for analysis step 27368 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep27368;
        DIMENSIONS NTAX=8;
        TAXLABELS
            Osmunda_japonica_J1
            Osmunda_japonica_J2J3J4J5J6J7J8L1L2L3
            Osmunda_regalis_R13R16b
            Osmunda_regalis_R14R16aR17R18
            Osmunda_regalis_R15
            Osmunda_regalis_R1R5R6R7
            Osmunda_regalis_R8R9
            Plenasium_banksiifolium
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M56341]

	TITLE Osmundaceae_L459;
	LINK TAXA = TaxaForAnalysisStep27368;
	DIMENSIONS NCHAR=507;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                                 10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       ]
[                                                  .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .       ]
Osmunda_japonica_J1                       AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAAGCTGTGAGTCATTTTGTTTTGAGCATCGACTATGAGTCTCTTTCTGATTCCCTCTAGTGCTTTGTCCTCATTCTGTCCAAATTGTCCTAACTGTGCGGCAATGTTGAACATGAAGGTTTCCTTCTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAAGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTCGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCGGTTGCAACTGCTGCCATCTTGGAGTCTTCAACTCCTCTTATTGACGGCAAACAATATTATAATCTTTCTGTTCTTACTCGCACTGCTGATGGTGGTGAGGGCGGGAAGCAT
Osmunda_japonica_J2J3J4J5J6J7J8L1L2L3     AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAAGCTGTGAGTCATTCTGTTTTGAGCATCGACTATGAGTCTCTTTCTGATTCCCTCTAGTGCTTTGTCCTCATTCTGTCCAAATTTTCCTAACTGTGCGGCAATGTTGAACATGAAGGTTTCCTTCTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAAGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTCGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCGGTTGCAACTGCTGCCATCTTGGAGTCTTCAACTCCTCTTATTGACGGCAAACAATATTATAATCTTTCTGTTCTTACTCGCACTGCTGATGGTGGTGAGGGCGGGAAGCAT
Osmunda_regalis_R13R16b                   AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAAGCTGTGAGTCATTTTGTTTTGAGCATCGACTATGAGTTTCTTTCTGATTCCCTTTAGTGCTTTGTCCTCATTCTGTCCAAATTTTCCTAACTGTGTGGCAATGTTGAACATGAAGGTTTCCTTCTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAAGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTTGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCTGTTGCAACTGCTGCCATCTTGGAGTCTTCAACTCCGCTTATTGACGGCAAACAATATTATAATCTTTCTGTTCTTACTCGGACTGCTGATGGTGGTGAGGGCGGAAAGCAT
Osmunda_regalis_R14R16aR17R18             AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAAGCTGTGAGTCATTTTGTTTTGAGCATCGACTATGAGTCTCTTTCTGATTCCCTTTAGTGCTTTGTCCTCATTCTGTCCAAATTTTCCTAACTGTGTGGCAATGTTGAACATGAAGGTTTCCTTCTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAAGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTTGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCTGTTGCAACTGCTGCCATCTTGGAGTCTTCAACTCCGCTTATTGACGGCAAACAATATTATAATCTTTCTGTTCTTACTCGGACTGCTGATGGTGGTGAGGGCGGAAAGCAT
Osmunda_regalis_R15                       AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAATCCGTGAGTCATTTTGTTTTGAGCATCGATTATGAGTCTCTTTCTGATTCCCTTTAGTGCTTTGTCCTCATTCCGTCCGAATTTTCCTAACTGTGCCGCAATGTTGAACCTGAAGGTTTCCTACTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGCAAAGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTCGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCAGTTGCAACTGCTGCCATCTTGGAGTCTTCAACACCTGTTATTGACGGCAAACAATATTATAATCTTTCTGTTCTTACTCGGACTGCTGATGGTGGCGAGGGCGGGAAGCAT
Osmunda_regalis_R1R5R6R7                  AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAAGCTGTGAGTCATTTTGTTTTCAGCATCGACTATGAGTCTCTTTCTGATTCCCTCTAGTGCTTTGTCCTCATTCTGTCCAAATTTTCCTAACTGTGCGGCAATGTTGAACATGAAGGTTTCCTTCTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAGGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTCGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCAGTTGCAACTGCTGCCATCTTGGAGTCCTCAACTCCTCTTATTGACGGCAAACAATATTATAATCTGTCTGTTCTTACTCGGACTGCTGATGGTGGTGAGGGCGGGAAGCAT
Osmunda_regalis_R8R9                      AATTTCGACGCAATTAACAATATTGCTGTGATGATCACCCCTACATCCAAGTCCTCAATTAAGGACTATGGCTCCCCTGAGCAGTTCCTTGAAGCTGTGAGTCATTTTGTTTTGAGCATCGACTATGAGTCTCTTTCTGATTCCCTTTAGTGCTTTGTCCTCATTCTGTCCAAATTTTCCTAACTGTGTGGCAATGTTGAACATGAAGGTTTCCTTCTTGCTAGGGAAGCAAGCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAAGGTCCTGTTGGCTGCTAGTTTTACTGGAATGTTCATGCAGTTTGTCAATCGTAAGGTGAATGTAAACCGACTGAGCCTGAATTTCACTTGGCCACAGGGTGGGTTCGACCCCAACGCTGTTGCAACTGCTGCCATCTTGGAGTCTTCAACTCCGCTTATTGACGGCAAACAATATTATAATCTTTCTGTTCTTACTCGGACTGCCGATGGTGGTGAGGGCGGAAAGCAT
Plenasium_banksiifolium                   AATTTCGACGCACTTAACAATATTGCTGTGATGGTCACCCCTACATCCAAGTCCTCAATCAAGGACTATGGCTCCCCCGAGCAGTTCCTTGAAACTGTGAGTCGTTTTGTTTTGAGCATCAACTATGAGTCTCTTTCGGATTCTCTTTAGTGCTTTGTCCTCGCCCTGTCCAAATTTTCCTAACTGTGCGATTATGTGGAACAAGCAGGTTTCCTTCTTGCTAGGGAAGCAATCGTATGCGGGCAAGACCCAGTCTGAGGTATGTAATTGACTGAAAACGTCCTGTTGGCTGCTAGTTTTAGTGGAATGTTCATGCAGTTTATCAATCGTAAGGTGAATGCAAACTGACTGAAGCTGAATTTCACTTGGCCACAGGGTGGGTTCGACGCCAACGCAGTTGCAACTGCCGCCATCTTGGAGTCTTCAACTCCTCTTATTGACGGCAAACAGTATTATAATCTCTCTGTTCTTACTCGAACTGCTGATGGTGGTGAGGGCGGGAAACAT
;
END;

[ The following blocks are output data for analysis step 27368 ]
BEGIN TREES;
      TITLE Osmundaceae_L459;
      LINK TAXA = TaxaForAnalysisStep27368;
      TREE con_50_majrule = [&R] (Osmunda_japonica_J2J3J4J5J6J7J8L1L2L3:0.003872,Osmunda_japonica_J1:0.003902,((Osmunda_regalis_R15:0.025913,((Osmunda_regalis_R14R16aR17R18:0.001949,Osmunda_regalis_R13R16b:0.003888,Osmunda_regalis_R8R9:0.003875):0.010546,Plenasium_banksiifolium:0.067878):0.003794):0.003906,Osmunda_regalis_R1R5R6R7:0.009882):0.005929);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr126307]


END;
BEGIN TREES;
      TITLE Osmundaceae_L459;
      LINK TAXA = TaxaForAnalysisStep27368;
      TREE con_50_majrule = [&R] (Plenasium_banksiifolium:0.033939,((Osmunda_regalis_R14R16aR17R18:0.001949,Osmunda_regalis_R13R16b:0.003888,Osmunda_regalis_R8R9:0.003875)1.00:0.010546,(Osmunda_regalis_R15:0.025913,(Osmunda_regalis_R1R5R6R7:0.009882,(Osmunda_japonica_J2J3J4J5J6J7J8L1L2L3:0.003872,Osmunda_japonica_J1:0.003902):0.005929)1.00:0.003906)0.89:0.003794)0.69:0.033939);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr126306]


END;