#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on November 10, 2026; 3:12 GMT

TreeBASE (cc) 1994-2008

Study reference:
Huyse T., & Volckaert F. 2005. Comparing Host and Parasite Phylogenies: Gyrodactylus
Flatworms Jumping from Goby to Goby. Systematic Biology, 54(5): 710-718.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S1470]

[ The following blocks are input data for analysis step 2851 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep2851;
        DIMENSIONS NTAX=8;
        TAXLABELS
            Gobius_paganellus
            Gobiusculus_flavescens
            Pomatoschistus_lozanoi
            Pomatoschistus_marmoratus
            Pomatoschistus_microps
            Pomatoschistus_minutus
            Pomatoschistus_norvegicus
            Pomatoschistus_pictus
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M1032]

	TITLE Host_12S_16S;
	LINK TAXA = TaxaForAnalysisStep2851;
	DIMENSIONS NCHAR=786;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                     10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510       520       530       540       550       560       570       580       590       600       610       620       630       640       650       660       670       680       690       700       710       720       730       740       750       760       770       780      ]
[                                      .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .      ]
Gobius_paganellus             CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAATCGATATTCCCCGTTCAACCTCACCCCCTCTTGTTTATACCGCCTATATACCGCCGTCGTCAGCTTACCCTGTGAAGGATTAATAGTAAGCAAAATTGGCAGAGCCTAAAACGTCAGGTCAAGGTGTAGCGTAC-AGGGGGGAAGAAATGGGCTACATTTGCTGACCCAGCAAACACGGACGTTGCACTGAAACGTGCAGCTAAAGGAGGATTTAGCAGTAAGGAGAGAATAGAGCGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCGAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCCGTATGAATGGCATAACGAGGGCTGTACTGTCTCCTTTCTCCAGTCAATGAAATTGATCTACTCGTGCAGAAGCGGGTATTAACACATAAGACGAGAAGACCCTATGGAGCTTTAGACACCAAAACAGACCACGTTAAAATCCCTGGACCAACAGGACTAACCAAGTGGGCGCCTGTTTTTTTGTCTTTGGTTGGGGCGACCGCGGAGAAACACAAATCCCCCGAGTGGAACGGGATCAACTCCCTGAGA-CAAGC?CCACAGGCCTAGCACCCAGAAAATCTGACCAAAAAGATCCGGCAATGCCGATCAACGGACCCAGTTACCCTAGGGATAACAGCGCAATCCCCTTCTAGAGCCCACATCAACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACTTCCTAATGGTGCAGCCGCTATTA
Gobiusculus_flavescens        CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTTAACCTCACCCTCTCTTGTTCTCCCCGCCTATATACCGCCGTCGTCAGCTTACCCTGTGAAGGACTAATAGTGAGCGTAATTGGTACAACCCAAAACGTCAGGTCGAGGTGTAGCGTATGAGAGGGGAAGAAATGGGCTACATTCACTGAGTCAGTGAAAACGAACGATGCCTTGAAACACTTATCTGAAGGAGGATTTAGCAGTAAGAGGGCAGTAGAGCGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTCTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATGAACACATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCAAACGTTAAAACCCCTGAGTAAACAGACATAACCAAGTTGATGCCTGCCCTTTTGTCTTTGGTTGGGGCGACCGCGGAGGAACAAAAATCCCCCACGTGGAACAGGAC-CTGCTCCTCAAAACCAGAGCCTCAGCTCTAGTAAACAGAACATCTGACCAAAAAGATCCGGTACTACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTCAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
Pomatoschistus_lozanoi        CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTAAACCTCACCCCCTCTTGTTCTCCCCGCCTATATACCGCCGTCGTCAGCTTACCCTCTGAAGGACTAATAGTGAGCATAATTGGTACAACCTAAAACGTCAGGTCGAGGTGTAGCGTATGAGGGGGGAAGAAATGGGCTACATTCACTGGCCCAGTGAAAACGAACGATGCTTTGAAACACCTATCCGAAGGAGGATTTAGCAGTAAGAGGGCAATAGAGTGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTTTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATAAAACCATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCAAACGTTAAGACCCCTGGGCCAACAGACACAACCAAGTTAATGCCTGCCCTTTTGTCTTTGGTTGGGGCGACCGCGGGGGAACAAAAATCCCCCACGTGGAACAGGAC-CTGCTCCTCAAAGCCAGAGCCACAGCTCTAGTAAACAGAACATCTGACCAAAAAGATCCGGTATTACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTCAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
Pomatoschistus_marmoratus     CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTTAACCTCACCCCCTCTTGTTCCCCCCGCCTATATACCGCCGTCGTCAGCTTACCCTGTGAAGGACTAACAGTGAGCATAATTGGCACAGCCTAAAACGTCAGGTCGAGGTGTAGCGTATGAGGGGGGAAGAAATGGGCTACATTCACTGAGCCAGTGAAAACGAACGATGCCTTGAAACAACCATCTGAAGGAGGATTTAGCAGTAAGAGGGCAGTAGAGCGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTCTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATGAAAACATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCGAACGTTAAGACCCCTGGGCCAACAGCTACAACCAAGTTAGCGCCTGCCCTTTTGTCTTTGGTTGGGGCGACCGCGGGGGAACGAAAATCCCCCACGTGGAACAGGAC-CTGCTCCTTAAGACCAGAGCCTCAGCTCTAGTAAACAGAACATCTGACCATAAAGATCCGGTAATACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTTAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
Pomatoschistus_microps        CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTTAACCTCACCCTCTCTTGTTCCTCCCGCCTATATACCGCCGTCGTCAGCTTACCCTGTGAAGGACTAACAGTGAGCATAATTGGCACAGCCTAAAACGTCAGGTCGAGGTGTAGCGTATGAGGGGGGAAGAAATGGGCTACATTCACTGAGCCAGTGAAAACGAACGATGCCTTGAAACACTTATCTGAAGGAGGATTTAGCAGTAAGAGGGCAATAGAGCGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTCTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATGAGAACATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCGAACGTTAAGACACCTGGACGAACAGCTACAACCAAGTTAGTGCCTGCCCTCTTGTCTTTGGTTGGGGCGACCGCGGGGGAACGAAAATCCCCCACGTGGAACAGGAC-CTGATCCTTAAAACCAGAGCCTCAGCTCTAGTAAACAGAACATCTGACCAAAAAGATCCGGTAATACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTTAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
Pomatoschistus_minutus        CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTAAACCTCACCCCCTCTTGTTCTCCCCGCCTATATACCGCCGTCGTCAGCTTACCCTATGAAGGACTAATAGTGAGCGCAATTGGTACAACCTAAAACGTCAGGTCGAGGTGTAGCGTATGAGGGGGGAAGAAATGGGCTACATTCACTGGCCCAGTGAAAACGAACGATGATTTGAAACACCTATCCGAAGGAGGATTTAGCAGTAAGAGGGCAATAGAGTGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTTTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATAAAACCATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCAAACGTTAAGACCCCTGGATAAACAGACACAACCTAGTTAGTGCCTGCCCTTTTGTCTTTGGTTGGGGCGACCGCGGGGGAACAAAAATCCCCCACGTGGAACAGGAC-CTGCTCCTTAAAACCAGAGCCACAGCTCTAGTAAACAGAACATCTGACCAAAAAGATCCGGTATTACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTCAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
Pomatoschistus_norvegicus     CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTAAACCTCACCCCCTCTTGTTCTCCCCGCCTATATACCGCCGTCGTCAGCTTACCCTGTGAAGGACTAATAGTGAGCATAATTGGTACAACCTAAAACGTCAGGTCGAGGTGTAGCGTATGAGGGGGGAAGAAATGGGCTACATTCACTGGCCCAGTGAAAACGAACGATGCTTTGAAACACCTATCCGAAGGAGGATTTAGCAGTAAGAGGGAAATAGAGTGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTTTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATAAAACCATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCAAACGTTAAGACCCCTGAATCAACAGACACAACCAAGTTAGTGCCTGCCCTTTTGTCTTTGGTTGGGGCGACCGCGGGGGAACAAAAATCCCCCACGTGGAACAGGAC-CTGCTCCTTAAAGCCAGAGCCACAGCTCTAGTGAACAGAACATCTGACCAAAAAGATCCGGTATTACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTCAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
Pomatoschistus_pictus         CCCAAAGGACTTGGCGGTGCTTTAGACCCCCCTAGAGGAGCCTGTTCTAGAACCGATAACCCCCGTTTAACCTCACCCCCTCTTGTTCTCCCCGCCTATATACCGCCGTCGTCAGCTTACCCTGTGAAGGACTAATAGTGAGCGTAATTGGTACAACCTAAAACGTCAGGTCGAGGTGTAGCGTATGAGGGGGGAAGAAATGGGCTACATTTACTGAATCAGTGAAAACGAACGATGCCTTGAAACACCTATCTCAAGGAGGATTTAGCAGTAAGAGGGCAGTAGAGCGCCCCTCTTTAACGGCCGCGGTATTTTGACCGTGCAAAGGTAGCGTAATCACTTGTCTTTTAAATGAAGACCTGTATGAATGGCATAACGAGGGCTAAGCTGTCTCCTCTCCCCAGTCAATGAAATTGATCTTTCCGTGCAGAAGCGGGAATGAAAACATAAGACGAGAAGACCCTATGGAGCTTAAGACAC-AAGGCAGCAAACGTTAAAACCCCTGGGCAAACAGATACAACCAAGTTAATGCCTGCCCTTTTGTCTTTTGTTGGGGCGACCGCGGGGGAACACAAATCCCCCACGTGGAACAGGAC-CTGCTCCTCAAAATCAGAGCCTCAGCTCTAGTAAACAGAACATCTGACCAAAAAGATCCGGTATTACCGATCAACGGACCGAGTTACCCTAGGGATAACAGCGCAATCCCCTTTCAGAGCCCATATCGACAAGGGGGTTTACGACCTCGATGTTGGATCAGGACCCCCTAATGGTGCAGCCGCTATTA
;
END;

BEGIN CODONS;
CODONPOSSET * CodonPositions (CHARACTERS = Host_12S_16S) = 
		N: 1-786;
CODONPOSSET  CodonPositions (CHARACTERS = Host_12S_16S) = 
		N: 1-786;
END;
[ The following blocks are output data for analysis step 2851 ]
BEGIN TREES;
      TITLE Tb7624;
      LINK TAXA = TaxaForAnalysisStep2851;
      TREE Fig._1.1 = [&R] (Gobius_paganellus,((Pomatoschistus_pictus,Gobiusculus_flavescens),((Pomatoschistus_minutus,(Pomatoschistus_norvegicus,Pomatoschistus_lozanoi)),(Pomatoschistus_microps,Pomatoschistus_marmoratus))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr500]


END;
BEGIN TREES;
      TITLE Tb7624;
      LINK TAXA = TaxaForAnalysisStep2851;
      TREE Fig._1.3 = [&R] (Gobius_paganellus,((Pomatoschistus_pictus,Gobiusculus_flavescens),((Pomatoschistus_lozanoi,(Pomatoschistus_norvegicus,Pomatoschistus_minutus)),(Pomatoschistus_microps,Pomatoschistus_marmoratus))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr499]


END;
BEGIN TREES;
      TITLE Tb7624;
      LINK TAXA = TaxaForAnalysisStep2851;
      TREE Fig._1.2 = [&R] (Gobius_paganellus,((Pomatoschistus_pictus,Gobiusculus_flavescens),((Pomatoschistus_norvegicus,(Pomatoschistus_minutus,Pomatoschistus_lozanoi)),(Pomatoschistus_microps,Pomatoschistus_marmoratus))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr498]


END;