#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on July 12, 2025; 11:03 GMT TreeBASE (cc) 1994-2008 Study reference: Islam M., Simmons M., Archer R.H., & Islam M. 2006. Phylogeny of the Elaeodendron Group (Celastraceae) Inferred from Morphological Characters and Nuclear and Plastid Genes. Systematic Botany, null. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S1551] [ The following blocks are input data for analysis step 3018 ] BEGIN TAXA; TITLE TaxaForAnalysisStep3018; DIMENSIONS NTAX=35; TAXLABELS Allocassine_laurifolia Brexia_madagascariensis Canotia_holacantha Cassine_peragua Cassine_schinoides Catha_edulis Elaeodendron_australe Elaeodendron_australeSBG Elaeodendron_cunninghamii Elaeodendron_curtipendulum Elaeodendron_matabelicum Elaeodendron_melanocarpum Elaeodendron_orientale Elaeodendron_pininsulare Elaeodendron_vitiense Elaeodendron_xylocarpum Empleuridium_juniperinum Gymnosporia_mossambicensis Lauridia_reticulata Lauridia_tetragona Lepuropetalon_spathulatum Loeseneriella_africana Lydenburgia_abbottii Lydenburgia_cassinoides Maurocenia_frangula Mortonia_greggii Mystroxylon_aethiopicum Pleurostylia_capensis Pleurostylia_leucocarpa Pleurostylia_opposita Pseudosalacia_streyi Putterlickia_verrucosa Robsonodendron_eucleiforme Salacia_impressifolia Salaciopsis_glomerata ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M1185] TITLE Morphology; LINK TAXA = TaxaForAnalysisStep3018; DIMENSIONS NCHAR=65; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5" MISSING=? GAP= -; CHARSTATELABELS 1 thorn_presence / absent present, 2 stem_apices / terminating_bluntly terminating_in_sharp_points, 3 presence_of_glands_on_stems / absent present, 4 phyllotaxy_on_vegetative_shoots / alternate opposite_or_whorled, 5 phyllotaxy_on_plants_with_alternate_leaves / strictly_alternate alternateonvegetative_shoots_opposite_on_thorns_or_flowering_shoots, 6 phyllotaxy_on_plants_with_opposite_leaves / strictly_opposite opposite_or_whorled, 7 leaf_form / planar 'sessile delta-shaped scales' cataphylls 'needle-like', 8 leaf_pubescence / without_stellate_hairs with_stellate_hairs with_Malphigian_hairs, 9 leaf_venation / pinnate acrodromous, 10 leaf_position / more_or_less_evenly_spaced fascicled_on_short_branches, 11 domatia_in_axils_of_midrib_and_nerves / absent sometimes_present, 12 intercostal_venation / reticulate distinct_crossbars, 13 stipule_position_on_opposite_leaves / intrapetiolar interpetiolar, 14 inflorescence_position / axillary at_least_some_inflorescences_terminal epiphyllous_or_rarely_axillary, 15 inflorescence_type / cymose thyrsoid_to_racemose umbel fasciculate flowers_solitary, 16 flower_sexuality / unisexual bisexual, 17 'unisexual-flowered plants' / dioecious 'monoecious = polygamous', 18 perianth_merosity / '4-merous' '5-merous' '3-merous', 19 sepal_gland_presence / absent present, 20 petal_margin / 'entire, ciliate, or irregularly toothed' regularly_toothed, 21 corolla_symmetry / actinomorphic_or_weakly_zygomorphic zygomorphic_with_4_of_5_petals_arched, 22 petal_fleshiness / not_fleshy fleshy, 23 petal_connation / free 'free at base, connate above', 24 disk_presence / absent present, 25 disk_division / continuous discontinuous, 26 disk_shape / cupular 'annular, flat, or margins upturned', 27 disk_adnation / not_adnate_to_sepals adnate_to_sepals, 28 disk_pubescence / glabrous conspicuously_puberulent, 29 androgynophore_presence / absent present, 30 stamen_plus_staminode_number / three_or_generally_three same_as_petal_number two more_than_petal_number, 31 fertile_stamen_length / all_of_a_single_length of_two_different_lengths, 32 staminode_presence_in_same_flower_with_functional_stamens / absent present, 33 'stamen-petal arrangement' / alternate opposite, 34 numerous_stamen_arrangement / unicyclic bicyclic, 35 numerous_stamen_number / twice_the_number_of_petals more_than_twice_the_number_of_petals less_than_twice_the_number_of_petals, 36 filament_insertion_relative_to_disk / at_outer_disk_margin on_top_of_disk inside_inner_edge_of_disk, 37 anther_dehiscence_direction / 'introrse to introrse-latrorse' strictly_latrorse extrorse apical, 38 anther_dehiscence_plane / longitudinal oblique transverse, 39 anther_attachment / basifixed dorsifixed, 40 anther_versatility / fixed versatile, 41 connective_extension_shape / absent_or_apiculate bilobed large_ornamented_extension, 42 pollen_aggregation / monads tetrads_or_polyads, 43 pollen_annulus_presence / absent present, 44 ovary_pubescence / glabrous completely_pilose strigose tomentose dense_hirsute apex_pilose, 45 shape_of_ovary_apex / contiguous apical_hollow, 46 style_connation / connate not_connate, 47 style_position_on_fruit / absent_or_terminal lateral, 48 carpel_number / one two three '= perianth merosity' twice_perianth_merosity 'four, when not equal perianth merosity', 49 ovary_septa_walls / complete incomplete absent, 50 ovule_number_per_locule / one two_or_four variable_and_more_than_four, 51 placentation / axile parietal, 52 axile_ovule_attachment / 'basal to axile, erect to horizontal' pendulous, 53 obturator_presence / absent present, 54 fruit_type / dehiscent indehiscent cocci, 55 indehiscent_fruit_type / drupe berry samara nut_or_achene, 56 fruit_wing_form / at_apex at_side_along_each_locule, 57 capsular_fruit_shape / not_lobed_or_parted strongly_parted_among_locules lobed_but_not_parted_among_locules lobed_to_base_but_not_parted_among_locules flattened_along_each_locule_but_not_parted, 58 mericarp_connation / separate connate, 59 capsular_fruit_dehiscence / loculicidal one_side_laterally_split septicidal loculicidal_and_septicidal, 60 aril_presence / present absent, 61 aril_position_on_seed / entirely_enveloping_seed partly_enveloping_seed, 62 aril_form / fleshy basal_wing_with_vasculature_of_the_funiculus_along_wing mucilagenous_pulp wing_surrounding_seed_with_medial_or_basal_attachment_of_funiculus basal_wing_with_vasculature_of_the_funiculus_attached_above_wing, 63 basal_seed_wing_form / 'membranous, papyraceous, or thin coriaceous' narrow_stipe, 64 raphe_branching / unbranched branched, 65 endosperm_presence / present absent, ; MATRIX [ 10 20 30 40 50 60 70 ] [ . . . . . . . ] Allocassine_laurifolia 000{01}10000000?0{03}1-10000010?0001000--0000?0??00001?100011----1---?0 Brexia_madagascariensis 00000-000000-001-1000001010001000--000100??00003{01}200011----1---?0 Canotia_holacantha 01100-10000--001-1000001010001000--000110??000031{12}0000----30140?0 Cassine_peragua 0001-0000000000{01}1{01}000001010001000--000{01}000000001?100011----1---?0 Cassine_schinoides 0001-00000000001-0000001010001000--0000000000001?100011----1---?0 Catha_edulis 000010000000{012}001-1000001010001000--0001100000-02{01}10000--0-0014000 Elaeodendron_australe 0001-0000000?00000000001010001000--000?10??00001?100010----1---?? Elaeodendron_australeSBG 0001-0000000?00000000001010001000--000?10??00001?100010----1---?? Elaeodendron_cunninghamii 00000-000000?00{01}1{01}0000010?0001000--000??0???0002?10?010----1---?0 Elaeodendron_curtipendulum 0001-0000000?0000{01}0000010?0001000--000??0??000?2?10?010----1---?0 Elaeodendron_matabelicum 0001-0000000?001-1000001010001000--0001?0??000?2?10?010----1---?? Elaeodendron_melanocarpum 0001-0000000?0000{02}0000010?0001000--000?10??00002?100010----1---?? Elaeodendron_orientale 00000-000000000{01}1{01}000001010001000--0001{01}0000000{123}0100010----1---00 Elaeodendron_pininsulare 000{01}00000000?10{01}?{01}000001010001000--000??0??000?{12}?10?010----1---?0 Elaeodendron_vitiense 0001-0000000?001-{01}000001010001000--000?10??0000{135}?100010----1---?? Elaeodendron_xylocarpum 000{01}00000000000011000001010001000--000110000000{123}0100010----1---00 Empleuridium_juniperinum 00000-30-0---041-00000010?0001000--00?1?00?0000321?000--0-1010-00 Gymnosporia_mossambicensis 10001-000100-000{01}1000001010001000--0001100000002{01}10000--0-0000-0{01} Lauridia_reticulata 0001-0000000?0{01}1-00000010?0001000--000??0??000?1?100011----1---?0 Lauridia_tetragona 0001-0000000?0{01}1-{01}0000010?0001000--000??0??000?1?100011----1---?0 Lepuropetalon_spathulatum 00000-001-00-141-1000000----0301010-000000000002221-00--0-01---?0 Loeseneriella_africana 0001-0000000{012}001-100000100001000---2221000100002020000--100011001 Lydenburgia_abbottii 000010000000{012}001-1000001010001000--0000100000002{01}10000--0-01---00 Lydenburgia_cassinoides 000{01}?0000000?001-1000001010001000--000?10??000?{25}?10000--0-0{01}??-?? Maurocenia_frangula 0001-000000000{03}{01}110000010??001000--0000?00?0000{12}?101011----1---?0 Mortonia_greggii 00000-000000-111-1000001010001000--00011000000031100013----1---00 Mystroxylon_aethiopicum 00000-000000-0{023}1-1000001010001000--000?10???000{12}0100010----1---?0 Pleurostylia_capensis 0001-0000000?001-1000001010001000--?00?????0001?11??013----1---?? Pleurostylia_leucocarpa 0001-0000000000{01}1{01}000001010001000--0001000000011?{12}00013----1---?0 Pleurostylia_opposita 0001-00000000001-{01}000001010001000--0001000000011?100013----1---?0 Pseudosalacia_streyi 00000-000000-0{013}1-{01}000001010001000--1?0??0??00002?10000--0-01---?0 Putterlickia_verrucosa 10001-000100-001-1000001010001000--000010??00002?20000--0-0000-00 Robsonodendron_eucleiforme 00000-000000-0{01}1-10000010?0001000--010??0??00001?100010----1---?0 Salacia_impressifolia 000{01}000000000031-100000100000000---22210000000020100011----0?2-?1 Salaciopsis_glomerata 00000-000000-030010000010100-1000--0001100000002010000--0-0010-00 ; END; [ The following blocks are output data for analysis step 3018 ] BEGIN TREES; TITLE Tb7791; LINK TAXA = TaxaForAnalysisStep3018; TREE Morph_Strict = [&R] (Mortonia_greggii,(Allocassine_laurifolia,Cassine_peragua,Cassine_schinoides,Lauridia_reticulata,Lauridia_tetragona,Maurocenia_frangula,(Pleurostylia_capensis,Pleurostylia_leucocarpa,Pleurostylia_opposita)),Brexia_madagascariensis,Canotia_holacantha,Lydenburgia_abbottii,Catha_edulis,Elaeodendron_australe,Elaeodendron_australeSBG,Elaeodendron_cunninghamii,Elaeodendron_curtipendulum,Elaeodendron_matabelicum,Elaeodendron_melanocarpum,Elaeodendron_orientale,Elaeodendron_pininsulare,Elaeodendron_vitiense,Elaeodendron_xylocarpum,(Empleuridium_juniperinum,Lepuropetalon_spathulatum),(Gymnosporia_mossambicensis,Putterlickia_verrucosa),(Loeseneriella_africana,Salacia_impressifolia),Lydenburgia_cassinoides,Mystroxylon_aethiopicum,Pseudosalacia_streyi,Robsonodendron_eucleiforme,Salaciopsis_glomerata); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr2463] END;