#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on September 19, 2025; 1:27 GMT

TreeBASE (cc) 1994-2008

Study reference:
Friar E., Prince L., Cruse-sanders J., Mcglaughlin M., Butterworth C., & Baldwin
B.G. 2008. Hybrid Origin and Genomic Mosaicism of Dubautia scabra (Hawaiian Silversword
Alliance; Asteraceae--Madiinae). Systematic Botany, null.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S1976]

[ The following blocks are input data for analysis step 4061 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep4061;
        DIMENSIONS NTAX=13;
        TAXLABELS
            Argyroxiphium_sandwicense_macrocephalum_ITS
            Argyroxiphium_sandwicense_sandwicense_ITS
            Dubautia_arborea_ITS
            Dubautia_ciliolata_ITS
            Dubautia_laxa_hirsuta_ITS
            Dubautia_linearis_ITS
            Dubautia_menziesii_ITS
            Dubautia_platyphylla_ITS
            Dubautia_reticulata_ITS
            Dubautia_scabra_HV_ITS
            Dubautia_scabra_OL_ITS
            Dubautia_scabra_PH_ITS
            Dubautia_waianapanapaensis_ITS
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M3005]

	TITLE nrITS;
	LINK TAXA = TaxaForAnalysisStep4061;
	DIMENSIONS NCHAR=644;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                                       10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510       520       530       540       550       560       570       580       590       600       610       620       630       640       ]
[                                                        .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .       ]
Argyroxiphium_sandwicense_macrocephalum_ITS     TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTCCTTGCGTGGCCATGTCGGCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCATGTAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTGGCTTCTTTTTAATCATAAATGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACG????AAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACGCATCACGTCGCCCCCACCATCCATCCCTGAC{CT}GGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCAGTTTGCGGTTGGCTTAAATACGAGTCTTGTCTAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGACGTGTTGTCTTTTGATGGCGCTTTGA
Argyroxiphium_sandwicense_sandwicense_ITS       TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTCCTTGCGTGGCCATGTCGGCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCATGTAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTGGCTTCTTTTTAATCATAAATGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACG?????????????TACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCAGTTTGCGGTTGGCTTAAATACGAGTCTTGTCTAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGACGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_arborea_ITS                            TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTTGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_ciliolata_ITS                          TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGAGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTTGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCATGTTGTCTTTTGATGGCGCTTTGA
Dubautia_laxa_hirsuta_ITS                       TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCACATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGGCCTGTGTCTGTGAGCTCCTCGGCGGGACTCATGGGCATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCA{CT}GTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGATTCCCAGTTTCTGGTGTGTTCATTGTTCGTGGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCATGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTGAGGGCACGTCTGCCTGGGCGTCACGCGTCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTTGTGCCTTCCTACTCTTGAAGCAGAAGACTCTTAAAATACCCTGACGTGTTGTCTTTGGATGGTGCTTTGA
Dubautia_linearis_ITS                           TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTTGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_menziesii_ITS                          TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTTTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_platyphylla_ITS                        TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTTTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_reticulata_ITS                         TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTTTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_scabra_HV_ITS                          TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATC{AG}TAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTTGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_scabra_OL_ITS                          TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CACTCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTTGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_scabra_PH_ITS                          TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTGTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATC{AG}TAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTTGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
Dubautia_waianapanapaensis_ITS                  TCGAATCCTGCACAGCAGAACGACCCGTGAACACGTACAACAACCTGGCCTCATGAGGACTGATCAATTTTGCTTCGGTTCTTGCGTGGCCATGTCGTCCTGTGTCTGTGAGCTCCTTGGCGGGACTCATGGACATGTTTTCGGCACAACAACAA-CCCCCGGCACGGCACGTGCCAAGGAAAACCAAACTTTAGAAGGTCCGTGCAATTGACTCCCAGTTTCTGGTGTGTTCATTGTTCGTTGCTTCTTTATAATCATAAACGACTCTCGGCAACGGATATCTCGGCTCACGCATCGATGAAGAACGTAGCAAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTTTTTGAACGCAAGTTGCGCCCGAAGCCTTCTGGTTAAGGGCACGTCTGCCTGGGCGTCATGCATCACGTCGCCCCCACCATCCATCCCTGACCGGATGTGTTGAT-GTGGGCGGAGATTGGTCTCCCGTGTCCATGATGCGGTTGGCCTAAATACGAGTCTTGTCGAGATGGACGCACGACTAGTGGTGGTTGATTACACAGTCGTCTCGTGTCGTGCCTTCCTATTCTTGAAGCAGAAGACTCTTAAAATACCCCGGCGTGTTGTCTTTTGATGGCGCTTTGA
;
END;

BEGIN CODONS;
CODONPOSSET * CodonPositions (CHARACTERS = nrITS) = 
		N: 1-644;
CODONPOSSET  CodonPositions (CHARACTERS = nrITS) = 
		N: 1-644;
END;
[ The following blocks are output data for analysis step 4061 ]
BEGIN TREES;
      TITLE Tb8833;
      LINK TAXA = TaxaForAnalysisStep4061;
      TREE Fig._3A = [&R] (Argyroxiphium_sandwicense_macrocephalum_ITS,(((Dubautia_arborea_ITS,Dubautia_ciliolata_ITS,Dubautia_linearis_ITS,Dubautia_scabra_HV_ITS,Dubautia_scabra_OL_ITS,Dubautia_scabra_PH_ITS),(Dubautia_menziesii_ITS,Dubautia_platyphylla_ITS,Dubautia_reticulata_ITS,Dubautia_waianapanapaensis_ITS)),Dubautia_laxa_hirsuta_ITS),Argyroxiphium_sandwicense_sandwicense_ITS);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5051]


END;