#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on September 19, 2025; 10:51 GMT

TreeBASE (cc) 1994-2008

Study reference:
Mathews K., Dunne N., York E., & Struwe L. 2008. A phylogenetic analysis and taxonomic
revision of Bartonia (Gentianaceae-Gentianeae) based on molecular and morphological
evidence. Systematic Botany, null.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S2096]

[ The following blocks are input data for analysis step 4362 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep4362;
        DIMENSIONS NTAX=12;
        TAXLABELS
            Bartonia_paniculata_subsp._iodandra
            Bartonia_paniculata_subsp._paniculata
            Bartonia_texana
            Bartonia_verna
            Bartonia_virginica
            Bartonia_virginica_KM35
            Gentiana_decora
            Gentianella_quinquefolia
            Latouchea_fokienensis
            Megacodon_stylophorus
            Obolaria_virginica
            Swertia_tashiroi
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M3284]

	TITLE 'ITS1_5.8S_ITS2';
	LINK TAXA = TaxaForAnalysisStep4362;
	DIMENSIONS NCHAR=684;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                                 10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510       520       530       540       550       560       570       580       590       600       610       620       630       640       650       660       670       680    ]
[                                                  .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .    ]
Bartonia_paniculata_subsp._iodandra       TCGATA-CTGC-AATCAGATGAC--GAG--ACATGTTT--TTACAGA--GGTGG-CGGAAC-AGGGATACCACGAACCGGTGCCCTAAGGTGCGGAGGCGACCACAAGTCTCTCGTCGTGCTACACAA-CAACCCCGGGCGCAGTAAAGCGCCAAGGTAAACAATAAAAAGGATTGCCTGCCTCTCGATGTGCCGTACGCGGAGTTCATG-----------AGAGGACAACTGGCACCTGGAGAATCTAAA---CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAACTGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCAAGGGCACGTCTGCCTGGGCGTCACGCATCGCATCGCCCCTT--AACCCTGCTAGTCAACT-TGTACG-GGTGAAGTGAGGGG-CGGAAAATGGCTTCCCGTGCTTGGTCGCGGCTGGCATAAATATGAGTCTCTCTAGACAGGCGCGATGACACAGTGGTGGTTGATAACTTCAGCTAAGATGTTGTCACGCGACGGTC-GTCGTTTGAGGAGACTACCC-AACCCTGTTGCATGTGTCGTCATGACACATGCTACGACCGCACC??CCACCCAACAACAAC?AC?CACACAACCA
Bartonia_paniculata_subsp._paniculata     TCGAATCCTGCAAATCAGATGACC-GAG-AACATGTTT--TTACACAAGGGCTTCCGGAACAAGGGAAACCACGAACCGGTGCCCTAAGGTGCGGCGTCGACCACAAGTCTCTCGTCGTGCTACACAA-CAACCCCGGGCGCAGTAAAGCGCCAAGGTAAACAAGAAAAAGGATTGCCTGCCTCTCGATGTGCCGTACGCGGAGTTCATG-----------AGAGGGCAACTGGCACCTGGAGAATCTAAA---CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAACTGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCAAGGGCACGTCTGCCTGGGCGTCACGCATCGCATCGCCCCCT--AACCCTGCTAGTCAACT-TGTACG-GGTGAAGTGAGGGG-CGGAAAATGGCTTCCCGTGCTTGGTCGCGGCTGGCATAAATATGAGTCTCTCTAGACAGGCGCGATGACACAGTGGTGGTTGATAACTTCAGCTAAGATGTTGTCACGCGACGGTC-GTCGTTTGAGGAGACTACCC-GACCCTGTTGCACGTGTCGTCATGACACATGCTACGACCGCAAACCACAAAAAACAACAAC?AC?CACACAACCA
Bartonia_texana                           TCGAATCCTGCAAATCAGATGACC-GAG-AACATGTTT--TTACACAAGGGCTTCCGGAACAAGGGAAACCACGAACCGGTGCCCTAAGGTGCGGCGTCGACCACAAGTCTCTCGTCGTGCTACACAA-CAACCCCGGGCGCAGTAAAGCGCCAAGGTAAACAAGAAAAAGGATTGCCTGCCTCTCGATGTGCCGTACGCGGAGTTCATG-----------AGAGGGCAACTGGCACCTGGAGAATCTAAA---CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAACTGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCAAGGGCACGTCTGCCTGGGCGTCACGCATCGCATCGCCCCTT--AACCCTGCTAGTCAACT-TGTACG-GGTGAAGTGAGGGG-CGGAAAATGGCTTCCCGTGCTTGGTCGCGGCTGGCATAAATATGAGTCTCTCTAGACAGGCGCGATGACACAGTGGTGGTTGATAACTTCAGCTAAGATGTTGTCACGCGACGGTC-GTCGTTTGAGGAGACTACCC-GACCCTGTTGCACGTGTCGTCATGACACATGCTACGACCGCAAACCACAAAAAACAACAAC?AC?CACACAACCA
Bartonia_verna                            TC-TC?CCTACAAATCAGATGACC-GAG-AACATGTTTCTTAACATACGGGCTTCCGGAACTAGGGAAACCGCGGACCGGTGCCCTAAG-TGCGGCGTGGACCACAAGTCTCTCGTCGTGC-ACGCAA-CAACCCCGGGCGCAGTAAAGCGCCAAGGTAAACAAGAAAAAGGATTGCCGGCCTCTCGACGTGCCGTATGTGGATTTCACGAGAGGAAACCGGG?AGGAAACTGGCAACTGTAGCAACAAAAAAACGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCAAGGGCACGTCTGCCTGGGCGTCACGCATCGCGTCGTCCCCT--GACCCTGCATGTGAAAT-CATCCG-GGTGAAGTGAGGGG-CGGAAAATGGCTTCCCGTGCTTGGTCGCGGTTGGCATAAATATGAGTCTCTCTCGATGGACGCGAAGACA-AGTGGTGGTTGATAGCTCCGGCTAAGATTCTGTCATGCGACGCCC-GTCGTCTGAGGAGACAAACC-GACCCTATTGCACGTGTCGTCATGACATATGCTACGACCGCCAACCAAAAAACCCAAAAAAAAC?CACACACCCA
Bartonia_virginica                        TCGAATCCTACAAAACAGATGACC-GAG-AACATGTTTTTTAACACACGGGCTTTCGGAACGAGGGAAACCGCGGACCGGTGCCCCAAG-TGCGGCGTCGACCATAAGTCTCTCGTCATGC-ACGCAA-CAACCCCGGGCGCAGTAAAGCGCCAAGGTAAACAAGAAAAAGGATTGTCAGCCTCTCGGCGTGCCGTACGTGGATTTCACG-----------AGAGGGCAACTGGCAACTGCAGAAACA------????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????ATCGCATCGTCCCCC--GACCCCACATGTGAAAT-CATCCG-GGTGAAGTGAGGGG-CGGAAAATGGCTTCCCGTGCTTGGTCGCGGTTGGCATAAATATGAGTCTCTCTTGATGGACGTGAAGACA-AATGGTGGTTGATAACTTCGGCTAAGATTTTGTCACACGACGCCC-GTCGTTTGAGGAGACTGACC-GACCCTATTGCACGTGCCATCATGACACATGCTACGACCGCAAACCAAAAAACCCAACCAA?AC?CACACACCCA
Bartonia_virginica_KM35                   TCGAATCCTACAAAACAGATGACC-GAG-AACATGTTTTTTAACACACGGGCTTTCGGAACGAGGGAAACCGCGGACCGGTGCCCCAAG-TGCGGCGTCGACCACAAGTCTCTCGTCATGC-ACGCAA-CAACCCCGGGCGCAGTAAAGCGCCAAGGTAAACAAGAAAAAGGATTGTCAGCCTCTCGGCGTGCCGTACGTGGATTTCACG-----------AGAGGGCAACTGGCAACTGCAGAAACAAAAA--CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCAAGGGCACGTCTGCCTGGGCGTCACGCATCGCATCGTCCCCC--GACCCCACATGTGAAAT-CATCCG-GGTGAAGTGAGGGG-CGGAAAATGGCTTCCCGTGCTTGGTCGCGGTTGGCATAAATATGAGTCTCTCTTGATGGACGTGAAGACA-AATGGTGGTTGATAACTTCGGCTAAGATTTTGTCACACGACGCCC-GTCGTTTGAGGAGACTGACC-GACCCTATTGCACGTGCCATCATGACACATGCTACGACCGCAAACCAAAAAACCCAACAAACAC?CACACACCCA
Gentiana_decora                           TCGAATCCTGCTAAGCAGACGACCCGAG-AACATGTTT---AACACACGGGCGTTCGGGACGAGGGAAACCACGGACCGATGCCCCGAG-CATTACGTCGACCACCGGTCGCTCGTCGTGC-AAACAACCAACCCCGGGCGCAGAAACGCGCCAAGGAAAACGAAAAAA-GGATGGCCTGCCTCCCGTCGTGCCGTACGCGGTGTGCACG-----------GGAGGATCACGGGCGCCTAAAGAAACAAAAA--CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAACTGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCGCGTCGCCCCCCC-AACACCGTGCATGAAACA--TTGCCGGTTGTCGGAGGGG-CGGATATTGGCTTCCCGTGCTTCGGTGCGGCTGGCCTAAATGCAAGTCCCTTGCGACGGACACGACGACA-AGTGGTGGTTGATTGCCTCAACTAAGAAGCTGTCGCTCGTTGCCCCGTCGGATTAGGAGACTTCCATGACCCTAATGCACGCGTCGTCACGATGCCTGCCACGACCGCAAAACA?CAAACCAACCAAACAACACAACACAAA
Gentianella_quinquefolia                  TCGAAGCCTGCGAAGGAGACGACCCGAG-AACATGTTT---AACAAACGGACGTCCGGGACGAGGGAAACCCCAGACCGGCGCCCCGAG-CACGTCGTCGACCATAGGTTGCTCGTCGTGC-ACACAAACAACCC-GGGCGCATAAAAGCGCCAAGGAAAACAAGAAAG-GGATGGCTTGCCTCTCGACGCTCCGTCCGCGGAGTGCACG-----------GGAGGGCAACAGGCACCTGAAGAAACAAAAA--CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAACTGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCGCGTCGCCCCCCC-AACCCTGTGTGTTAACT-CATACG-GGTGACGTGAGGGGGCGGAAACTGGCTTCCCGTGCTTGGCCGCGGCTGGCCTAAATGCGAGTCCCTTGAGACGGACGCGACGACA-AGTGGTGGTTGATAGTCTCAACTAAGGTGCTGTCGCGCGACGCCC-GTCGAATTAGGAGACTCCCC-GACCCTGATGCACGCGTCGTTACGACGCTTGCTACGACCGCAAAACA?CAAACCACCCAAACAACCACAAACCCA
Latouchea_fokienensis                     TCGAATCCTGCGAAGCAGATGACCCGAG-AACATGTTT---ACATCACGGGTGTCCGGGACGGGGGAAACCACGGACCGGTGCCCCTAG-CACGGCGTCGACCATCGGTCGCTCGACGTGC-ACACAAACAACCCCGGGCGTATAAACGCGCCAAGGAAAACTATAAAG-GGATTGCCTGTCTCTCGACGCTCCGTACGCGGAGTGGACG-----------GGAGGGCAACAGGCGCCTGAAGAAACAA-----????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????-TCGCGTCGCCCCCC--AACCCCGTGCATTAACTATGTACT-GGTGATTTGAGGGG-CGGAAGCTGGCTTCCCGTGCTTACCCGCGGCTGGCCTAAAAGCGAGTCCTTTGCGACGGACGCGACGACA-AGTGGTGGTTGATTACCTCAACTAAGGTGCTGTCGCGCGACCTTC-GTCGAATGAGGAGACTCTCC-GACCCTGATGCATGCGTCGTCACGACGCATGCTACGACCGCAAAACA?CAAACCAACCACA?CC?AACACACCCA
Megacodon_stylophorus                     TCGAATCCTGCGAAGCAGACGACCCGAG-AACATGTTT---AACGCACGGGCGTCCGGGCCGGGGGAAACCACGGACCGGCGTCCCGAG-CGCGGCGCCGACCACCGGTCGCTCGTCGTGC-AAACAAACAACCCCGGGCGCAGAAAAGCGCCAAGGAAAACGAGAAAG-GGATGGCTTGCCTCTCGACGCGCCGTACGCGGAGTGCACG-----------GGAGGGCAACAGGCGCCTGAAGAAACAAAAA--CGACTCTCGGCAACGGATATCTCGGCTCTCGCATCGATGAAGAACGTAGCGAACTGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCGAGTCTTTGAACGCAAGTTGCGCCCGAAGCCATTAGGCCGAGGGCACGTCTGCCTGGGCGTCACGCATCGCGTCGCCCCCC--AACCTCGTGCGTTAACT-CGTACG-GGTGGCGTGAGGGGGCGGAAAATGGCTTCCCGTGCTTGGCCGCGGCTGGCCTAAATGCGAGTCCCTTGCGACGGACGCGACGACA-AGTGGTGGTTGATTGCCTCAACTAAGGTGCTGTCGCGCGACGCCC-GTCGATTGAGGAGACTCCCC-GACCCTGATGCATGCGTCGTCACGACGTCTGCTACGACCGCAAAACA?CAAACCAACCAAACAC?CACAAACCCA
Obolaria_virginica                        TCGAATCCTGCGAAGCAGACGACCCGAGTAACATGTTT---AACCCACGGGCATCCGGGACGGGGGAAACCACGGACCGGCGCTCCGAG-CACGGCGTCGACAACCTGTCGATTTCCGTGC-TCACAAACAACCCCGGGCGCAGAAAAGCGCCAAGGAAAACGTGAAAG-GGCTAGCCAACCACCCGACGCGCCGTACGCGGAGTGCACG-----------GGATGGAAACAGGCACCTTAAGAAACA------????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????ATCGCGTCGCCCCTC--AACCCCGTGTATTAACTACGTACGTGGTGAAGTGAGGG--CGGAAACTGGCTTCCCGTGGTTGCCCGCGGCTGGCCTAAATGTGAGTCCCTTGCGATGGACGCGACGACA-AGTGGTGGTTGATTGCCTCAACTAAGGTGCTGTCGCGCGACGTCT-GTCGAAAGAAGAGACTCTAC-GACCCTAATGCATG-GTCGTCACGACGCATGCTACGACCGCAAAAAA?CAAACCAAACCAA?AC?AAAC?ACCCC
Swertia_tashiroi                          TCGAATCCTGCGAAGCAGACGACCCGAG-AACATGTTT---ATCCCACGGGTGTCCGGGACGAGGGAAACCATGTACCGGTGCCCCGAG-CGCGGCGTCGACCACTGGTCGCTCGTCGTGC-ACACAAACAACCCCGGGCGTAGAAAAGCGCCAAGGAAAACAAGAAAG-GGAAGGTCTGCCTCTCGACGCGCCGTACGCGGAGTGCACG-----------GGAGGGCAACAGGCGCCTGAAGAAACA------????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????ATCGCGTCGCCCCCCCCAACCCCGTGTGTTAAAC-CGTACG-GGTGACGTGAGGGGGCGGAAAATGGCTTCCCGTGCTTG-TCGCGGCTGGCCTAAATGTGAGTCCCTTGCGATGGACGCGATGACA-AGTGGTGGTTGATTGCCTCAACTAAGGTGCTGTCGCGCGACGCCC-GTCGAGTGAGGAGACTCCCC-GACCCTGATGCACGTGTCGTCATGACGCCTGCTACGACCGCAAAACA?CAAACCAACCCAA?AAACACAACCCCA
;
END;

[ The following blocks are output data for analysis step 4362 ]
BEGIN TREES;
      TITLE Tb9134;
      LINK TAXA = TaxaForAnalysisStep4362;
      TREE Fig._3a = [&R] (Gentiana_decora,((Megacodon_stylophorus,((Swertia_tashiroi,((Bartonia_paniculata_subsp._paniculata,(Bartonia_texana,Bartonia_paniculata_subsp._iodandra)),(Bartonia_verna,(Bartonia_virginica,Bartonia_virginica_KM35)))Bartonia),Gentianella_quinquefolia)),(Latouchea_fokienensis,Obolaria_virginica))Swertiinae);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5206]


END;
BEGIN TREES;
      TITLE Tb9134;
      LINK TAXA = TaxaForAnalysisStep4362;
      TREE ITS_2 = [&R] (Gentiana_decora,(((Megacodon_stylophorus,(Swertia_tashiroi,((Bartonia_paniculata_subsp._paniculata,(Bartonia_texana,Bartonia_paniculata_subsp._iodandra)),(Bartonia_verna,(Bartonia_virginica,Bartonia_virginica_KM35)))Bartonia)),Gentianella_quinquefolia),(Latouchea_fokienensis,Obolaria_virginica))Swertiinae);
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5207]


END;