#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 19, 2025; 4:55 GMT TreeBASE (cc) 1994-2008 Study reference: Pillon Y., Hopkins H., Munzinger J., & Chase M. 2008. A molecular and morphological survey of generic limits of Acsmithia and Spiraeanthemum (Cunoniaceae). Systematic Botany, null. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S2115] [ The following blocks are input data for analysis step 4400 ] BEGIN TAXA; TITLE TaxaForAnalysisStep4400; DIMENSIONS NTAX=11; TAXLABELS Brunellia_colombiana Hooglandia_ignambensis Platylophus_trifoliatus Spiraeanthemum_brongniartianum Spiraeanthemum_densiflorum_JM4575 Spiraeanthemum_densiflorum_YP667 Spiraeanthemum_ellipticum Spiraeanthemum_macgillivrayi Spiraeanthemum_meridionale Spiraeanthemum_pubescens Spiraeanthemum_samoense ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M3318] TITLE trnL; LINK TAXA = TaxaForAnalysisStep4400; DIMENSIONS NCHAR=952; FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 370 380 390 400 410 420 430 440 450 460 470 480 490 500 510 520 530 540 550 560 570 580 590 600 610 620 630 640 650 660 670 680 690 700 710 720 730 740 750 760 770 780 790 800 810 820 830 840 850 860 870 880 890 900 910 920 930 940 950 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Brunellia_colombiana CTTTCAAATTCAGAGAAACCCTGGAATTAAAAATGGTCAATCCTGAGCCAAATCCTGTTTTCCGAAAAAAAAAAAAAGGTTCAGAAAGCGAGAA------TAAAAAAAA--GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGATTGCGTTGCGTTAGTAAAGGAATCTTTCCGTCAAAACCCCAGAAAGGATAAAG---TAAAAAAA-GGATAAAAGTAAAAAGATAAACCCATATACATACGTATACATACTGAAATACTCTCTCAAATGATTAATGATGACCCGAATCCGTGTTTTTTATAAAACTAATTATAAAAATAATGAAAGATTTGTTGTGAATCGATTCCAAGTTGAAGAAAGAATCGAATATTAATTGATCAAATCATTCACTCCAGCGTCGGATAGATCTTTTGAAAACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCGTTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAA????????????????????????????????????????????????????????????????????????????TTCTCTCT-TTCGTTAACGAGTCAAAATTCGTTATCTTTCTCATTCATTTTACTCCTTCACAAACAAATGGATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGAGCAA---------------------------------------------------------AGTCTTCTTTTT---------GAAGATCCAAAAAATTTCAGTGCCTGGTAA-GACTTTGTAATACTTTTTTT-CGTCTTTTTAATT-GAC----------ATAGACCTAAGTCATCTAGTAAAATCAGGATGATACGTG Hooglandia_ignambensis CTTTCAAATTCAGAGAAACCCTGGAATTAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAA------TAAAAAAAA--GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAAAAAATGGGGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCCTCAAAACTCCAGAAAGGGTAAAG--------TAAAGGATAAA--------------CCCATATACGTACATATACGTACTGAACTACTATCTCAAATGATTAATGATGACCTGAATTCGTATTATTGATAAAAATAA------------TGAAAGGTTTGTTGTGAATGGATTCCAAGTTGAAGAAAGAATCGAATATTAATTGATCAAATCATTCACTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCGTTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGGGGAAAA?????????????????????????????????????????????????????????????CTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGGATCCTGGCGGAAATTTTTTTT-CTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATTTTTGAGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGCCTTCTTTTTTTTATTTTTGAAGATCCAAAAAATTTAAG-GACTGGTAT-TACTTTGTAATACTTTTTTT-CGTCTTTTTAATT-GAC----------ATAGACCCAAGTCATCTAGTAAAATGAGGATGATACGTC Platylophus_trifoliatus CTTTCAAATTCAGAGAAACCCTGGAATTAAAAATGGGCAATCCTGAGCCAAATCCCGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAA------TAAAAAAAA--GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCCTCAAAACTCCAGAAAGGGTAAAG--------TAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATTACCTGAATTCGTATTTTTTATAAAAATAA------------TGAAAGCTTTGTTGTGAATTGATTCCAAGTTGAAGAAAGAATCGAATATTAATTGATCAAATCATTCACTCCAGTGTCGGATAGATCTTTTGAATACCTGATTAATCAGAC?A?AATAAATATATAGTCCCATTCTACATGTCAATATTGACAA?AATGAAAT?TATA??AA????AAAA????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????AACTTACAAAGCCTTCTTTTT--AATTTTTGAAGATCCAAAAAATTTAAG-GCCTGGGAA-TACTTTGTAATACTTTTTTT-CGTCTTTTT-ATT-----TTAATTGACATAGACCCAAGTCATCTAGTAAAATGAGGATGATACGTC Spiraeanthemum_brongniartianum CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAA------TAAAAAAAAAAGGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTAAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCCCTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGTATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGGGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGTCCTCTTTTT--GATTTTTGAAGATCCAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTTTTT-CCTCTTTTTAATT-GACTTTAATTGACATAGACCCAAGTCATCTAGTAAAATGAGGATGATACGTC Spiraeanthemum_densiflorum_JM4575 CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAATAAAAATCAAAAAAA--GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAATAAATGGAGTTGACTGCGTTGCGTTAGTAAAGAAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTGAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCACTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGGATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGAGCAAGGAATCCCCATTTGAATGATTAACAATC----------CTCGTACTGAAACTTACAAAGGCCTCTTTTT--GATTTTTGAAGATCAAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTATTTTCGTCTTTTTAATT-GACTTTAATTGACATAGATCCAAGTCATCTAGTAAAATGAGGATGATACGTC Spiraeanthemum_densiflorum_YP667 CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAATAAAAATAAAAAAAA--GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAATAAATGGAGTTGACTGCGTTGCGTTAGTAACGAAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTGAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCACTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGGATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGAGCAAGGAATCCCCATTTGAATGATTAACAATC----------CTCGTACTGAAACTTACAAAG-CCTCTTTTTT-GATTTTTGAAGATCCAAAAAATT-CAG-GC-TGG??????????????????????????????????????????????????????????????????????????????????????????? Spiraeanthemum_ellipticum CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAA------TAAAAAAAAA-GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTAAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCCCTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGTATCCAGGCGGAAACTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGGGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGTCCTCTTTTT--GATTTTTGAAGATCCAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTTTTT-CGTCTTTTTAATT-GACTTTAATTGACATAGACCCAAGTCATCTAGTAAAATGAGGATGATACGTC Spiraeanthemum_macgillivrayi CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCCGAAAGCGAGAA------TAAAAAAAA-GGGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CTCATATACATACGTATACGTACTGAAATACTATTTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTAAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCACTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGTATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGGGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGTCCTCTTTTT--GATTTTTGAAGATCCAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTTTTT-CGTCTTTTTAATT-GACTTTAATTGACATAGACCCAAGTCATCTAGTAAAATGAGGATGATACGTC Spiraeanthemum_meridionale CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAA------TAAAAAAAAA-GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCGTCAAAACGCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTAAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCCCTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGTATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGGGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGTCCTCTTTTT--GATTTTTGAAGATCCAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTTTTTTCGTCTTTTTAATT-GACTTTAATTGACATAGACCCAAGTCATCTAGTAAAATGAGGATGATACGTC Spiraeanthemum_pubescens CTTTCAA-TTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCAGAAAGCGAGAA------TAAAAAAAAA-GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATATACATACGTATACGTACTGAAATACTATCTCAAATGATTAATGATGACCCGACTCCGTATTTTTTATAAAAATAA------------TGAAAGGTTTGTT--------ATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCCCTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGACGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAATGAAATTTATAGTAAGAGGAAAATCCGTCGACTTTAGAAATCGTGAGGGTTCAAGTCCCTCTATCCCCAAAAAAGCCCGTTTGACTCCCTAATTATTTATTCTCTCTGTTCGTTAACGATTCAAAATTCGTTATCTTTCTCATTCATTTTACTCTTTCACAAACAAATGTATCCAGGCGGAAATTTTTTTTTCTTATTCTTACTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGGGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGTCCTCTTTTT--GATTTTTGAAGATCCAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTTTTT-CGTCTTTTTAATT-GACTTTAATTGACATAGACCCAAGTCATCTAGTAAAATGAGGATGATAC??? Spiraeanthemum_samoense CTTTCAAATTCAGAGAAACCCTGGAATGAAAAATGGGCAATCCTGAGCCAAATCCTGTTTTCCGAAAACAAA-CAAAGGTTCCGAAAGCGAGAA------TAAAAAAAAA-GGATAGGTGCAGAGACTCAACGGAAGCTGTTCTAACACTAACAAATGGAGTTGACTGCGTTGCGTTAGTAAAGGAATCCTTCCGTCAAAACTCCAGAAAGGATAAAGTGATAAAGTAAAGGATAAA--------------CCCATAT??????GTA--CGTACTGAAATACTATTTCAAAT{GT}ATTAATGAT{GT}ACCCGAC?C-GTATTTTTTATAAAAATAA------------TGAAAGGTTTGTTGTAAATTGATTCCAAGTTGAAGAAAGAATTGAATATTAATTGATCAAATCATTCACTCCAGTGTCGGATAGATCTTTTGAAGACCTGATTAATCGGATGAGAATAAAGATAGAGTCCCATTCTACATGTCAATATTGACAACAAT????????????????????????????????????????????????????????????????????????????????????????????????????????????TCGTTAACGATTCAAAATTCGTTATCTTTCACATTCATTT-ACTCTTTCACAAACAAATGTATCCAGGCGGAAATTTTTTTTTCTTATTCTTATTACAAGTCTTGTGATAGATATGATACACGTACAAATGAACATCTTTGGGCAAGGAATCCCCATTTGAATGATTAACAATCCATATCATTACTCGTACTGAAACTTACAAAGTCCTCTTTTT--GATTTTTGAAGATCCAAAAAATTTCAG-GCCTGGTAC-TACTTTGTAATACTTTTTTT-CGT-TTTTTAATTCGACTTTAATTGACATAGACCCAAGTCATCTAATAAAATGAGGATGATAC??? ; END; BEGIN CODONS; CODONPOSSET * CodonPositions (CHARACTERS = trnL) = N: 1-952; CODONPOSSET CodonPositions (CHARACTERS = trnL) = N: 1-952; END; [ The following blocks are output data for analysis step 4400 ] BEGIN TREES; TITLE Tb9172; LINK TAXA = TaxaForAnalysisStep4400; TREE Fig._1 = [&R] (((Platylophus_trifoliatus,Hooglandia_ignambensis),(((((Spiraeanthemum_brongniartianum,Spiraeanthemum_meridionale),Spiraeanthemum_ellipticum),Spiraeanthemum_pubescens),(Spiraeanthemum_macgillivrayi,Spiraeanthemum_samoense)),(Spiraeanthemum_densiflorum_YP667,Spiraeanthemum_densiflorum_JM4575))),Brunellia_colombiana); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr5218] END;