#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on September 19, 2025; 8:17 GMT TreeBASE (cc) 1994-2008 Study reference: Lehnert M., Kessler M., Schmidt-lebuhn A., Klimas S., Fehlberg S., & Ranker T. 2008. Phylogeny of the fern genus Melpomene (Polypodiaceae) inferred from morphology and chloroplast DNA analysis. Systematic Botany, null. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S2152] [ The following blocks are input data for analysis step 4492 ] BEGIN TAXA; TITLE TaxaForAnalysisStep4492; DIMENSIONS NTAX=74; TAXLABELS Lellingeria_apiculata_AY362343 Lellingeria_apiculata_AY460648 Lellingeria_hirsuta_AY460649 Lellingeria_limula_AY460650 Lellingeria_pseudomitchelliae_AY460652 Lellingeria_schenckii_AY460651 Lellingeria_subsessilis_AY460653 Melpomene_albicans_sp_nov_ined_ML514 Melpomene_albicans_sp_nov_ined_ML601 Melpomene_albicans_sp_nov_ined_ML717 Melpomene_caput_gorgonis_sp_nov_ined_ML372 Melpomene_erecta_ML1570 Melpomene_firma_ML1003 Melpomene_firma_ML516 Melpomene_flabelliformis_AF1 Melpomene_flabelliformis_AF2 Melpomene_flabelliformis_AY460656 Melpomene_flabelliformis_CO5 Melpomene_flabelliformis_IJ1874 Melpomene_flabelliformis_JK1189 Melpomene_flabelliformis_ML955 Melpomene_flabelliformis_TJ2927 Melpomene_flagellata_sp_nov_ined_ML555 Melpomene_gracilis_ML1554 Melpomene_jimenezii_sp_nov_ined_IJ1859 Melpomene_leptostoma_MK13515 Melpomene_leptostoma_MK13528 Melpomene_melanosticta_ML1533 Melpomene_michaelis_sp_nov_ined_ML519 Melpomene_moniliformis_AY460654 Melpomene_moniliformis_AY460655 Melpomene_moniliformis_ML1110 Melpomene_moniliformis_ML122 Melpomene_moniliformis_ML435 Melpomene_occidentalis_sp_nov_ined_ML1343 Melpomene_occidentalis_sp_nov_ined_ML1558a Melpomene_paradoxa_sp_nov_ined_ML388 Melpomene_paradoxa_sp_nov_ined_ML539 Melpomene_personata_sp_nov_ined_6946 Melpomene_personata_sp_nov_ined_K1126 Melpomene_personata_sp_nov_ined_ML145 Melpomene_peruviana_ML273 Melpomene_peruviana_ML571 Melpomene_peruviana_ML775 Melpomene_pilosissima_MK13514 Melpomene_pilosissima_TK2738 Melpomene_pseudonutans_AY460657 Melpomene_pseudonutans_ML119 Melpomene_pseudonutans_ML1558 Melpomene_sklenarii_sp_nov_ined_ML1465 Melpomene_sodiroi_ML1106 Melpomene_sodiroi_ML173 Melpomene_vernicosa_MK13502 Melpomene_vernicosa_ML1333 Melpomene_vulcanica_sp_nov_ined_ML174 Melpomene_vulcanica_sp_nov_ined_ML176 Melpomene_wolfii_ML1229 Melpomene_wolfii_ML985 Melpomene_xiphopteroides_MK13484 Melpomene_xiphopteroides_ML187 Melpomene_xiphopteroides_ML477 Melpomene_xiphopteroides_ML792 Micropolypodium_hyalinum_AY362344 Terpsichore_achilleifolia_AY460666 Terpsichore_alsopteris_AF468209 Terpsichore_anfractuosa_AY460668 Terpsichore_eggersii_AF468209 Terpsichore_lanigera_AY460672 Terpsichore_lehmannianaAY460673 Terpsichore_longisetosa_AY460674 Terpsichore_pichinchae_AY460675 Terpsichore_sp_AY460679 Terpsichore_subcabra_AY460677 Terpsichore_subtilis_AY460678 ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M3821] TITLE morpho_data_Melpomene; LINK TAXA = TaxaForAnalysisStep4492; DIMENSIONS NCHAR=39; FORMAT DATATYPE=Standard SYMBOLS= "A T C G 0 1 2 3 4" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 ] [ . . . . . . ] Lellingeria_apiculata_AY362343 0000211010012000122210102{12}2022{01}01100000 Lellingeria_apiculata_AY460648 3000211010012000122210102{12}2022{01}01100000 Lellingeria_hirsuta_AY460649 31002120100110001112101022000{12}{01}01100000 Lellingeria_limula_AY460650 300021200001100110031010213032100100000 Lellingeria_pseudomitchelliae_AY460652 300000000000000{01}1002111000322{23}100100000 Lellingeria_schenckii_AY460651 300021201001100{01}11021{12}102022321111{01}1000 Lellingeria_subsessilis_AY460653 3000212110013000101312102{13}00011{01}1100000 Melpomene_albicans_sp_nov_ined_ML514 {23}010222202{23}2301{12}1213111021112{02}00{12}000000 Melpomene_albicans_sp_nov_ined_ML601 {23}010222202{23}2301{12}1213111021112{02}00{12}000000 Melpomene_albicans_sp_nov_ined_ML717 {23}010222202{23}2301{12}1213111021112{02}00{12}000000 Melpomene_caput_gorgonis_sp_nov_ined_ML372 3110222203{23}2401{12}111321102{01}2202101{01}01000 Melpomene_erecta_ML1570 {12}121222202344111112102102{01}1100012002000 Melpomene_firma_ML1003 {123}{01}{02}0222201{13}24001122312102{13}2110{01}01200000 Melpomene_firma_ML516 {123}{01}{02}0222201{13}24001122312102{13}2110{01}01200000 Melpomene_flabelliformis_AF1 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_AF2 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_AY460656 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_CO5 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_IJ1874 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_JK1189 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_ML955 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flabelliformis_TJ2927 {0123}01122220232311212231{01}1020210{02}00{12}{01}01000 Melpomene_flagellata_sp_nov_ined_ML555 302022220222300{12}11122{01}1020210210{13}00{13}000 Melpomene_gracilis_ML1554 20{01}02{12}21011210001211001{12}21000{01}10{01}112000 Melpomene_jimenezii_sp_nov_ined_IJ1859 3{01}0022110222400012232010112121001001000 Melpomene_leptostoma_MK13515 300022220211200011131010210020102100000 Melpomene_leptostoma_MK13528 300022220211200011131010210020102100000 Melpomene_melanosticta_ML1533 3000222102312001111002102{01}000{02}00100{12}000 Melpomene_michaelis_sp_nov_ined_ML519 {23}0{02}02211021110001113201011222{23}101001000 Melpomene_moniliformis_AY460654 {0123}01122220222301211131{12}1020120300{01}011000 Melpomene_moniliformis_AY460655 {0123}01122220222301211131{12}1020120300{01}011000 Melpomene_moniliformis_ML1110 {0123}01122220222301211131{12}1020120300{01}011000 Melpomene_moniliformis_ML122 {0123}01122220222301211131{12}1020120300{01}011000 Melpomene_moniliformis_ML435 {0123}01122220222301211131{12}1020120300{01}011000 Melpomene_occidentalis_sp_nov_ined_ML1343 31{02}0222201{13}24001122{01}00102{12}2210001{02}00000 Melpomene_occidentalis_sp_nov_ined_ML1558a 31{02}0222201{13}24001122{01}00102{12}2210001{02}00000 Melpomene_paradoxa_sp_nov_ined_ML388 3011222{12}02223112111311102{01}110000{01}001000 Melpomene_paradoxa_sp_nov_ined_ML539 3011222{12}02223112111311102{01}110000{01}001000 Melpomene_personata_sp_nov_ined_6946 3011222102{12}130101213{12}01021212{01}10{12}{01}01000 Melpomene_personata_sp_nov_ined_K1126 3011222102{12}130101213{12}01021212{01}10{12}{01}01000 Melpomene_personata_sp_nov_ined_ML145 3011222102{12}130101213{12}01021212{01}10{12}{01}01000 Melpomene_peruviana_ML273 {02}011222202{23}1{23}01011131{01}102{02}211{02}{01}0{12}1{01}1000 Melpomene_peruviana_ML571 {02}011222202{23}1{23}01011131{01}102{02}211{02}{01}0{12}1{01}1000 Melpomene_peruviana_ML775 {02}011222202{23}1{23}01011131{01}102{02}211{02}{01}0{12}1{01}1000 Melpomene_pilosissima_MK13514 30{02}0222202{12}13{01}001113{12}01021223{02}10{12}101000 Melpomene_pilosissima_TK2738 30{02}0222202{12}13{01}001113{12}01021223{02}10{12}101000 Melpomene_pseudonutans_AY460657 {23}1{12}1222202{23}24012113{01}02102{13}2232{12}1{12}101000 Melpomene_pseudonutans_ML119 {23}1{12}1222202{23}24012113{01}02102{13}2232{12}1{12}101000 Melpomene_pseudonutans_ML1558 {23}1{12}1222202{23}24012113{01}02102{13}2232{12}1{12}101000 Melpomene_sklenarii_sp_nov_ined_ML1465 {01}011222102222112111{02}1{12}10202130{01}0{01}11{12}000 Melpomene_sodiroi_ML1106 {23}{01}10222102124{01}1012100{01}1021110{02}101{01}02000 Melpomene_sodiroi_ML173 {23}{01}10222102124{01}1012100{01}1021110{02}101{01}02000 Melpomene_vernicosa_MK13502 {23}110222202{23}2400112332{01}1021212{01}0{01}100{12}000 Melpomene_vernicosa_ML1333 {23}110222202{23}2400112332{01}1021212{01}0{01}100{12}000 Melpomene_vulcanica_sp_nov_ined_ML174 {03}011222102{12}3{34}1111223111022111{12}10{12}100000 Melpomene_vulcanica_sp_nov_ined_ML176 {03}011222102{12}3{34}1111223111022111{12}10{12}100000 Melpomene_wolfii_ML1229 31{02}0222202{12}{23}300{12}1120021020220{03}1{01}{13}003000 Melpomene_wolfii_ML985 31{02}0222202{12}{23}300{12}1120021020220{03}1{01}{13}003000 Melpomene_xiphopteroides_MK13484 30{01}0222102{12}1100{12}111321102{13}212{01}00{01}101000 Melpomene_xiphopteroides_ML187 30{01}0222102{12}1100{12}111321102{13}212{01}00{01}101000 Melpomene_xiphopteroides_ML477 30{01}0222102{12}1100{12}111321102{13}212{01}00{01}101000 Melpomene_xiphopteroides_ML792 30{01}0222102{12}1100{12}111321102{13}212{01}00{01}101000 Micropolypodium_hyalinum_AY362344 30002210100120001112221020312210{01}000000 Terpsichore_achilleifolia_AY460666 30002100000120011113112201322110{12}111000 Terpsichore_alsopteris_AF468209 311021001001201212321010203220{01}0{12}{01}00010 Terpsichore_anfractuosa_AY460668 {23}0002110{01}211100{12}1102121021213{02}{01}0{01}1{01}1111 Terpsichore_eggersii_AF468209 30002{12}001002300{01}121320102{12}322{02}00{12}100000 Terpsichore_lanigera_AY460672 300021001002200001032{01}102032221{01}{12}100000 Terpsichore_lehmannianaAY460673 31{01}021001001100003322010123122002100000 Terpsichore_longisetosa_AY460674 3000210000011001131320221{12}3220102100000 Terpsichore_pichinchae_AY460675 301021{01}0100120111122101021322{02}101000011 Terpsichore_sp_AY460679 3000210000011001131320221{12}3220102100000 Terpsichore_subcabra_AY460677 30002100000110000101001120000010{01}100000 Terpsichore_subtilis_AY460678 3000210000011000100220100032200{01}0100000 ; END; [ The following blocks are output data for analysis step 4492 ] BEGIN TREES; TITLE Tb9264; LINK TAXA = TaxaForAnalysisStep4492; TREE morph_tree = [&R] (Melpomene_gracilis_ML1554,(((((((((Melpomene_albicans_sp_nov_ined_ML514,Melpomene_albicans_sp_nov_ined_ML601),((Melpomene_caput_gorgonis_sp_nov_ined_ML372,Melpomene_flagellata_sp_nov_ined_ML555),(((((Melpomene_flabelliformis_AF2,Melpomene_flabelliformis_AY460656,Melpomene_flabelliformis_CO5,Melpomene_flabelliformis_ML955,Melpomene_flabelliformis_JK1189,Melpomene_flabelliformis_IJ1874,Melpomene_flabelliformis_TJ2927),(Melpomene_vulcanica_sp_nov_ined_ML176,Melpomene_vulcanica_sp_nov_ined_ML174)),Melpomene_sklenarii_sp_nov_ined_ML1465),(Melpomene_peruviana_ML273,Melpomene_peruviana_ML775)),((Melpomene_moniliformis_AY460654,Melpomene_moniliformis_AY460655,Melpomene_moniliformis_ML1110,Melpomene_moniliformis_ML435),(Melpomene_paradoxa_sp_nov_ined_ML388,Melpomene_paradoxa_sp_nov_ined_ML539))))),(Melpomene_vernicosa_ML1333,Melpomene_vernicosa_MK13502)),Melpomene_jimenezii_sp_nov_ined_IJ1859),(((Melpomene_erecta_ML1570,(Melpomene_pseudonutans_AY460657,Melpomene_pseudonutans_ML119,Melpomene_pseudonutans_ML1558)),(((Melpomene_firma_ML1003,Melpomene_firma_ML516),(Melpomene_occidentalis_sp_nov_ined_ML1343,Melpomene_occidentalis_sp_nov_ined_ML1558a)),(Melpomene_wolfii_ML1229,Melpomene_wolfii_ML985))),Melpomene_melanosticta_ML1533)),(Melpomene_xiphopteroides_ML187,Melpomene_xiphopteroides_ML477,Melpomene_xiphopteroides_ML792,Melpomene_xiphopteroides_MK13484)),(Melpomene_michaelis_sp_nov_ined_ML519,(((Melpomene_leptostoma_MK13515,Melpomene_leptostoma_MK13528),(((((((Terpsichore_achilleifolia_AY460666,Terpsichore_longisetosa_AY460674),(Terpsichore_subtilis_AY460678,Lellingeria_pseudomitchelliae_AY460652)),((Terpsichore_alsopteris_AF468209,Terpsichore_pichinchae_AY460675),(((Terpsichore_eggersii_AF468209,Terpsichore_lanigera_AY460672),(Terpsichore_lehmannianaAY460673,Micropolypodium_hyalinum_AY362344)),(Lellingeria_apiculata_AY460648,Lellingeria_apiculata_AY362343)))),(Terpsichore_anfractuosa_AY460668,Lellingeria_schenckii_AY460651)),Lellingeria_limula_AY460650),Terpsichore_subcabra_AY460677),(Lellingeria_hirsuta_AY460649,Lellingeria_subsessilis_AY460653))),(Melpomene_pilosissima_TK2738,Melpomene_pilosissima_MK13514)))),(Melpomene_personata_sp_nov_ined_6946,Melpomene_personata_sp_nov_ined_K1126,Melpomene_personata_sp_nov_ined_ML145)),(Melpomene_sodiroi_ML1106,Melpomene_sodiroi_ML173))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr3107] END;