#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on October 09, 2025; 8:03 GMT

TreeBASE (cc) 1994-2008

Study reference:
Bininda-emonds O., & Price S. 2008. A comprehensive phylogeny of extant horses, rhinos
and tapirs (Perissodactyla) through data combination. Zoosystematics and Evolution,
null.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S2218]

[ The following blocks are input data for analysis step 4657 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep4657;
        DIMENSIONS NTAX=19;
        TAXLABELS
            Ceratotherium_simum
            Dicerorhinus_sumatrensis
            Diceros_bicornis
            Equus_asinus
            Equus_burchellii
            Equus_caballus
            Equus_grevyi
            Equus_hemionus
            Equus_kiang
            Equus_onager
            Equus_quagga
            Equus_zebra
            Perissodactyla_MRP_outgroup
            Rhinoceros_sondaicus
            Rhinoceros_unicornis
            Tapirus_bairdii
            Tapirus_indicus
            Tapirus_pinchaque
            Tapirus_terrestris
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M3892]

	TITLE Perissodactyla_literature_MRP;
	LINK TAXA = TaxaForAnalysisStep4657;
	DIMENSIONS NCHAR=80;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1" MISSING=? GAP= -;

MATRIX
[                                       10        20        30        40        50        60        70        80]
[                                        .         .         .         .         .         .         .         .]
Ceratotherium_simum             01000101000111010001???????011001???????????????010010????0000???0011011??????10
Dicerorhinus_sumatrensis        ??????01000110?????????????011100??????????00101100010????????????????10????????
Diceros_bicornis                ??????01000111?????????????011001??????????00101010010???????????0011011????????
Equus_asinus                    100000100000001000001101000??????1000000100???????01000011111000011000??00100001
Equus_burchellii                ????????????????????0010110??????000100101110000??0100110?????110???????001110??
Equus_caballus                  100000100000001000000000101100000000111000011000??010000001000001100000010000001
Equus_grevyi                    ????????????????????10001101000000001001010???????01001101111110011000??001101??
Equus_hemionus                  ???????????????????????1000??????1100000??????????0100001???????????????????????
Equus_kiang                     ?????????????????????????????????1110000??????????0100??????????????????110000??
Equus_onager                    ????????????????????110??????????1110000100???????0100????1111001???????110000??
Equus_quagga                    ?????????????????????????????????0001110011???????0100??????????????????001110??
Equus_zebra                     ????????????????????0010101??????000110001011000??010010001100110???????001101??
Perissodactyla_MRP_outgroup     00000000000000000000???????000000??????????00000??0000???????????00000????????00
Rhinoceros_sondaicus            ???????????????????????????011110?????????????????0011??????????????????????????
Rhinoceros_unicornis            01000101000100010001???????011110???????????????100011???????????0010000??????10
Tapirus_bairdii                 01110001110000011100???????????????????????00110??1000??????????????????????????
Tapirus_indicus                 01110001100000011000???????????????????????00110001000????0000???00001??????????
Tapirus_pinchaque               01101001111000011110??????????????????????????????1000??????????????????????????
Tapirus_terrestris              01101001111000011110???????010000?????????????????1000???????????00001??????????
;
END;

BEGIN SETS;
	CHARSET NormanAshley2000_3 (CHARACTERS = Perissodactyla_literature_MRP) =  15-20;
	CHARSET Amato_etal1993 (CHARACTERS = Perissodactyla_literature_MRP) =  49-50;
	CHARSET HarrisPorter1980 (CHARACTERS = Perissodactyla_literature_MRP) =  73-78;
	CHARSET PitraVeits2000 (CHARACTERS = Perissodactyla_literature_MRP) =  66-70;
	CHARSET MoralesMelnick1994 (CHARACTERS = Perissodactyla_literature_MRP) =  71-72;
	CHARSET PerezBarberiaGordon1999 (CHARACTERS = Perissodactyla_literature_MRP) =  44-48;
	CHARSET NormanAshley2000_2 (CHARACTERS = Perissodactyla_literature_MRP) =  7-14;
	CHARSET ArnasonJanke2002 (CHARACTERS = Perissodactyla_literature_MRP) =  79-80;
	CHARSET NormanAshley2000_1 (CHARACTERS = Perissodactyla_literature_MRP) =  1-6;
	CHARSET Grubb1993 (CHARACTERS = Perissodactyla_literature_MRP) =  51-54;
	CHARSET GeorgeRyder1986 (CHARACTERS = Perissodactyla_literature_MRP) =  24-27;
	CHARSET Ishida_etal1995_1 (CHARACTERS = Perissodactyla_literature_MRP) =  58;
	CHARSET LowensteinRyder1985 (CHARACTERS = Perissodactyla_literature_MRP) =  41-43;
	CHARSET Ishida_etal1995_2 (CHARACTERS = Perissodactyla_literature_MRP) =  55-57;
	CHARSET Flint_etal1990 (CHARACTERS = Perissodactyla_literature_MRP) =  59-62;
	CHARSET OakenfullClegg1998 (CHARACTERS = Perissodactyla_literature_MRP) =  21-23;
	CHARSET Bennett1980 (CHARACTERS = Perissodactyla_literature_MRP) =  34-40;
	CHARSET Tougard_etal2001 (CHARACTERS = Perissodactyla_literature_MRP) =  28-33;
	CHARSET OakenfullClegg1998_mST (CHARACTERS = Perissodactyla_literature_MRP) =  63-65;
END;
[ The following blocks are output data for analysis step 4657 ]
BEGIN TREES;
      TITLE Tb9429;
      LINK TAXA = TaxaForAnalysisStep4657;
      TREE Fig._1a = [&R] ((((Ceratotherium_simum,Diceros_bicornis),Dicerorhinus_sumatrensis,(Rhinoceros_sondaicus,Rhinoceros_unicornis)),((Tapirus_bairdii,(Tapirus_pinchaque,Tapirus_terrestris)),Tapirus_indicus)),(((Equus_asinus,(Equus_hemionus,(Equus_kiang,Equus_onager))),(Equus_burchellii,Equus_grevyi)),Equus_caballus,Equus_quagga,Equus_zebra));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr3183]


END;