#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on June 15, 2021; 10:10 GMT TreeBASE (cc) 1994-2008 Study reference: Dikow T. 2009. A phylogenetic hypothesis for Asilidae based on a total evidence analysis of morphological and DNA sequence data (Insecta: Diptera: Brachycera: Asiloidea). Organisms Diversity & Evolution, 9(3): 165-188. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S9957] [ The following blocks are input data for analysis step 4922 ] BEGIN TAXA; TITLE TaxaForAnalysisStep4922; DIMENSIONS NTAX=88; TAXLABELS Ablautus_coquilletti Acnephalum_cylindricum Afroestricus_chiastoneurus Afroleptomydas_sp_Clanwilliam Apiocera_painteri Apsilocephala_longistyla Asilus_crabroniformis Asilus_sericeus Atomosia_puella Beameromyia_bifida Bombylius_major Ceraturgus_fasciatus Cerotainia_albipilosa Choerades_bella Clephydroneura_sp_Kepong Connomyia_varipennis Cyrtopogon_rattus Damalis_annulata Damalis_monochaetes Dasophrys_crenulatus Dasypogon_diadema Dioctria_atricapillus Dioctria_hyalipennis Dioctria_rufipes Diogmites_grossus Dysmachus_trigonus Emphysomera_conopsoides Emphysomera_pallidapex Eudioctria_albius Euscelidia_pulchra Gonioscelis_ventralis Hemigephyra_atra Holcocephala_abdominalis Holcocephala_calva Hoplistomerus_nobilis Lamyra_gulo Laphria_aktis Laphystia_tolandi Lasiocnemus_lugens Lasiopogon_aldrichii Lasiopogon_cinctus Laxenecera_albicincta Leptarthrus_brevirostris Leptogaster_arida Leptogaster_cylindrica Lestomyia_fraudiger Lycostommyia_albifacies Machimus_occidentalis Megaphorus_pulchrus Microstylum_sp_Karkloof Mitrodetus_dentitarsis Molobratia_teutonus Mydas_clavatus Nemomydas_brachyrhynchus Neoitamus_cyanurus Neolophonotus_bimaculatus Nicocles_politus Nusa_infumata Ommatius_tibialis Opomydas_townsendi Ospriocerus_aeacus Pegesimallus_laticornis Perasis_transvaalensis Philodicus_tenuipes Philonicus_albiceps Phycus_frommeri Pilica_formidolosa Plesiomma_sp_Guanacaste Pogonioefferia_pogonias Proctacanthus_philadelphicus Prolepsis_tristis Promachus_amastrus Prorates_sp_Escalante Rhabdogaster_pedion Rhipidocephala_sp_HaroldJohnson Saropogon_luteus Scylaticus_costalis Stenopogon_rufibarbis Stichopogon_elegantulus Stichopogon_punctum Stichopogon_trifasciatus Stiphrolamyra_angularis Tillobroma_punctipennis Tipulogaster_glabrata Tolmerus_atricapillus Trichardis_effrena Trichoura_sp_Tierberg Willistonina_bilineata ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M4175] TITLE Morphological_matrix; LINK TAXA = TaxaForAnalysisStep4922; DIMENSIONS NCHAR=220; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7 8" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 ] [ . . . . . . . . . . . . . . . . . . . . . . ] Ablautus_coquilletti 0011012020010110001000100110111011101111000031000011021002000201020122012201011000200002111111001200001100-0120202010000000-102000001000010000001010100000000000000000021402110001100000200000000000101-11011100000010000000 Acnephalum_cylindricum 0024013020110010001001101110111111101110000001001011001001000200220122011201011100200003111111011200001100-0120102000000010-101000001001010200001110100000000000000000011402110001101000000000000000101-00101200000010000000 Afroestricus_chiastoneurus 0023012020000110000000100110011011200111000001000011050002100200020121011401011010201202111111011210130110-012020201000001110000001000112100000010101000100000000111100000-0200001110000100011100000100-110112000000001011-0 Afroleptomydas_sp_Clanwilliam 001421302000000004000003010210020001100--01200000010030143-11010020111101012100001100100000111020001100100-00201010200000010002000114011320400302101100000100000000000011022211000101000300000010000002002000--00000011011-0 Apiocera_painteri 000000--20101011040000031110100000011011000000000001051002010100020122002201000000000003111011011210000100-00202110100011110002000100011210000000000100000000000000000021722210?011??000000000000000101-01011100000010001100 Apsilocephala_longistyla 000000--2111000004000003111210020001101110000000001100000100000002002-001000001000000001111111011210000000-002021100000111110000110000110000000011101000000000000000000000-0200001132000200000000000021-01001200000010000000 Asilus_crabroniformis 0022011020000010000000111110111011200111000000001011001001000201020121011101011011202202111011011210100110-012020201000001110000001020112100100010101000000000000000000202-0200001100010100000000000100-100110000000021011-0 Asilus_sericeus 0022011020000010000000100110011011200111000000001011000002000101020121011101001011202102111011011210100110-01{01}020201000111110010001020112100100010101000000000000000000202-0200001111010100000000000100-100110000000021011-0 Atomosia_puella 0022010020100010000000211-10111011101112010000000011010053-00200020121011011011102101001111111111210030110-012010100020001110000011010114003000010101110000001022000011000-5200001121000022220006010001-01001100000012001100 Beameromyia_bifida 0011010020010010000000200110111021202111000001000011000002000000010100021011022000100001110111011110010010-0101101000001110-1010020000000000002010101000200000100000000000-4200200112001000000000000112022101210000020001100 Bombylius_major 010000--1001000000000003011010120001100--0000000001100000100011001001-101200011000101100100000011110001000-002010101000111110010000011-0-11000011010000100000000000000003812110001132000000000000010001-01001201000020000100 Ceraturgus_fasciatus 0024012020110110021001111011111011101111000001000011011031000200020111011201011000200002111011011110000110-00201010000000111001000001000000000001010100001000000000000021402110?011??000100000000010101-00011200001110000100 Cerotainia_albipilosa 0022011020000010000000221-10111011101112010001010111110053-00100020121011011011002101001111011111110030110-012010100020001110010011000114103000010101110000001000000011000-5200001120001000000006010001-01001100000022001100 Choerades_bella 0022011020100110020010221-10111011101112010011000011120043-00200010111011111011100201001111111031211010110-002010100000111110000001010112000000010101000000000020000000001-5200001121000020000005010002101001200000032101110 Clephydroneura_sp_Kepong 0022011020000010000000100110011011200111000001000011001001000100020121011101011011201102111111011110110110-012020201000111110010001020112100000010101000000000011110000203-0200001122010111000000000100-100110001000021011-0 Connomyia_varipennis 0012013020110110001000111111111011101110100000001011010002000200020122012301011000200002111111011210000100-01202020100011111001000001001000000001010100000000000000000021702110001121000300000000010000-00011200000010000000 Cyrtopogon_rattus 0024013020110110000000100111111011101111000001000011010011000100020111011301011000200002111111011210000110-01201010000010111000001001000000000001010100000000000000000021402110001121000200000000000100-10011100001110000000 Damalis_annulata 0013012020000010020000211-10111011101111000101000011040023-00100010111010111011000200000101111011100010110-012010212000000110010000010012102000010101000000000000000000000-5200001121100100000001010002012001200000010000000 Damalis_monochaetes 0012011020000010020000211-10111011101111000101000011040023-00100020111010111011000201000100111011100110110-012010211000000110000000010012102000010101000000000000000000000-5200001101100100000001010202012001100000020000000 Dasophrys_crenulatus 0024013020000210000000100110011011200111000000000011051001000101021121011201011000201102111111011210100110-012010201000011110000001000112100000010101000000000011100000203-0200001010010111100000000100-100110000000001011-0 Dasypogon_diadema 0014011020110010002001111111111111101111100000000011011012000200220122011301001000200003111011001200000112121202020000011111000000001011210010001010100000000000000000010602110001111000200000000010001-10001100100110000000 Dioctria_atricapillus 0022010020000210031100211-10111011101111000000000011111032000100020111011311022000200001110010111110000110-012012200020001110010010010000000000010101000010000000000000000-0200001101001000000004000001-00011200001010000100 Dioctria_hyalipennis 0022011020000210031100211-10111011101111000000000011111032000100020111011311022000200001110010111110000110-012012200020001110010010010000000001010101000110000000000000000-0200001101001000000004000001-00011200001010000100 Dioctria_rufipes 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0022013020000110000000100110011011200111000001000011050002100200021121011101011010202202111111011210130110-012020201000001110010001000112100000010101000100100000111100000-4200001110000200000000000100-100111000000001011-0 Eudioctria_albius 0021010020100210001000101110111011101111000001000011121032000100010111011301002000200001110111011000000110-012012100020101110000010000000000000010101000010000000000000004-0200001101001000000004010000-01011200001010000100 Euscelidia_pulchra 0011010020010010000000200110111021200111000001000011000002000000010100021011022000100001110011121110020010-0101001000001010-1010020000000000001010101000200000100000000000-4200100111000000000000000112022101210000020001100 Gonioscelis_ventralis 1013012020110210000000101110111011001110000000001011010001000200021121012201011000000003111111011200000110-01202020100011111001000001010200011001010100000000000000000011402110001101000200000000000100-11111200000010000000 Hemigephyra_atra 000000--21110200042000030100100200011011000000000001010012000000020121002100000000100001111000011200000000-002021200000111110000100010110001000010101000000000000000000037-0201001131001000000000000000-00011200000010101110 Holcocephala_abdominalis 0020110020000010020000211-10111011101111000101000011011002000100010111011111011000200000100010021010000110-012010100000000110010010000010000000010101001000000000000000000-4200001101100200000004000202022111201000012001000 Holcocephala_calva 0020110020000000020000211-10111011101111000101000011011002000100010111011111011000200000100010021010000110-012010100000000110010010000010000000010101000100000000000000000-4200001101100200000004000202022111201000012001000 Hoplistomerus_nobilis 0024010020110110001010201-10111011101111010001001011011012000200020111012011011000201000111010111111000110-002010102001001110000002011112002110110101010000000022222000000-5200001121000022222003010002100101200000022101110 Lamyra_gulo 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0013012020110210000001100111111011100111000000010011001002000102021111011001000000200002111111011200000110-01201010000000011000002001001000000101010100000000000000000020602110001100001200000001010101-11011200010010001001 Lasiopogon_cinctus 001301202011021000000010011111101110011100000001001101100200010202111101100100000020000{23}111111011200000110-01201010000000011000002001001000000101010100000000000000000020602110001100000200000001010101-11011200010010001000 Laxenecera_albicincta 0024013020100110000000100010111011101111010001001011021043-0020002011101111101110020110{23}111111011211010110-002010110001000110000001011112001100010101000000000020000000000-5200001121000022000003010002000101100000022101110 Leptarthrus_brevirostris 00140130201100100000011101111111111011110000010000110100010001000201110113010110001000031001111111000001110012010200000111110010010010010100001010101010000000000000021101-2200001120100000000000000101-20011200000010000000 Leptogaster_arida 0011010020010010000000200110111021200111000001000011000002000000020100021011022000100001110011021100010010-0101101000001100-1010020000001000002010101000200000100000000000-4200010112000000000000000110-11101210000020001100 Leptogaster_cylindrica 0011010020010010000000200110111021200111000001000011000002000000020100021011022000100001111011021110010010-0101101000000000-1010020000001000002010101000200000100000000000-4200010111001000000000000111-11101210000020001100 Lestomyia_fraudiger 0023012020110010000001111111111111101111100001000011021012000201220122012301001000200002111111011200000112011202010100011111100000000000210000001010100000000000000000021402110001100000200000000010001-10011200000110000100 Lycostommyia_albifacies 0014010020110110000001211-1111111110111010000000001101001100010022011101120101100020110{23}111011011210000110-01201020200010111000000001000200000001010100000000000000000011402110001121000200000000010101-11011100100120010000 Machimus_occidentalis 0023012020000110000000100110011011200111000001000011001001000101020121011101011011202102111111011210100110-012020201000001110000001020112100000010101000000000011110000203-0200001111010111110000000100-100110001000021011-0 Megaphorus_pulchrus 0022013020000110000000100110011011200111000001000011000002000101010111011201011010101102111111011110110110-010010101000000110100001010112001000012101001000000000000000002-1200001110010100000000000100-100211000000021011-0 Microstylum_sp_Karkloof 0012011020000110021001211110111011101110100000001011020012000100020112012301011001201103111111011210000100-01201020100011111000000001011210010001010120000000000000000020402110001112000300000000010101-10011200001010010000 Mitrodetus_dentitarsis 001421300000020004000003010210020001100--01200000010030143-11110010111101012100001000100000111010011100100-00201010100000110002000114111320420302001100000100000000000011022212--0012000100000010000102002010--0001020101110 Molobratia_teutonus 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0022011020000010000000100110011011200111000001001011051002000101020121011201001010202102111011011210110110-011020201000001110000001010112000100012101001000000000000000202-0220010011010111110000000100-100210001000021010-0 Philonicus_albiceps 0022011020000110000000100110011011200111000001001011001001000201020121011101011011202102111111011210100110-012020201000111110000001020112100100010101000000000011100000202-0220001111010111111000000100-100110000000021011-0 Phycus_frommeri 000000--21110200042000031112100200011011000000000001010001000100020121001100010000200002111011011210000000-00201110000011111000010001011000010001010100000000000000000000712210001131000000000000000002000011200000010101110 Pilica_formidolosa 0023012020001010110010211-11111011101111010001001011020043-00200020121011111011101201101111111011211110110-002010200001111110000001011112000100010101000000000022220000001-3200000122010122220005010002101001200100022101110 Plesiomma_sp_Guanacaste 0024010020110010001000111110111021101112000200020011011011000200020121012301001000200003111011010000000100-01202020000011111000000001001210000001010100010000000000000021402110011121100200000000010100-10011200000020000100 Pogonioefferia_pogonias 0024013020000110000000100110011011200111000001001011051002000201020121011201011000201102111111011210100110-010020201000101110000001030112000000001101001000000000000000203-0200001111000100000000001100-200010001000021010-0 Proctacanthus_philadelphicus 0023012020000010000001101111011011200111000000001011051001000201020122012201011000201102111011011210100110-012020201000001110000001030112000000000101001000000000000000202-0221001132010100000000000100-100110000000011010-0 Prolepsis_tristis 0013012020110110001001111111111011101110100000001011010043-00200020122012301001000201003111011011210000110-01202020100011111000000001011210000001010100000000000000000020402110001132000200000000010101-11011100001110000100 Promachus_amastrus 0022011020000010000000100110011011200111000001001011051002000201020121011201011010202102111111011210110110-012020201000011110010001010112000000012101001000000000000000002-1200011000010111111000000100-100210000000021011-0 Prorates_sp_Escalante 010000--2111000004000003010010020001101100000000000100001200000000002-112001000000000000101000001100000000-000001100000111110020100010012101000010100001000000000000000000-221????????00000000000000102001021200000010101110 Rhabdogaster_pedion 0021011020110210001000110010111111101111000001000011010001000100020111010301011000100002111111011200030110-01201020100011111000001000001010200101010100010000000000000021402110001101100000000000000101-10001200000010010000 Rhipidocephala_sp_HaroldJohnson 0010011021100010000000221-10111011101111000101000011011032000100010111011111011000200000100111021010000110-012010101000011110010010010000000000010101001000010000000000005-0200001101100200000002000101-11011200000010001000 Saropogon_luteus 0011010020110010002001110011111111101111000001001011121012000200220122012301001000200003111011011200000112121202010000011111000000001000210000001010100000000000000000021402110001121000200000004010100-10011200001020000000 Scylaticus_costalis 001201102011021000100110111011111110111010000000101101001200010002012101220100100020000{23}111111011200000100-01202020100011111000000001000210000001010100000000000000000021402110001121100200000000010101-10011200000130000000 Stenopogon_rufibarbis 1024013020110210000001101110111011200110100000000011001001000100021122011201001000000003111111011200000110-01202020100011111000000001000200001011010100010000000000000001402110001101000200000000000100-10011101000020000100 Stichopogon_elegantulus 0011010120110210000000110111111011100111000001010011000002000102020111011011001000200001111101021100010100-002011201000000110000020010010000000010101000000000000000000206021100011010002000000040000020310112002001001010-1 Stichopogon_punctum 0011010120110210001000110111111011100111000001010011000002000202020111011011011000200001111111021110010100-002011100000000110000020010010000001010101000000000000000000106021100011010002000000040000020310112002001001010-1 Stichopogon_trifasciatus 0011010120110210000000100111111011100111000001010011000002000102020111011011001000200001101010021100010100-002011200000000110000020010010000001010101000000000000000000006021100011211002000000040000020310112002001001010-1 Stiphrolamyra_angularis 0024012020000010000010221-11111011001111010001101011020043-00200020111011111011100201101111011111111000110-002010200000111110100001010112000100110101010000000022220000001-5200001121000022200003010002111111100000022101110 Tillobroma_punctipennis 0024013020110110000001101110111111101110100001000011020012000200010111011301011000200002111111011210000110-00201120100011111000000001000200000001010100010000000000000021402110001121100200000000010101-10011201000110000000 Tipulogaster_glabrata 0021010020010010000000200110111021200111000001000011000002000000020100021301022000100001110000021110010010-0101001000001110-1010020000001000002010101000200000100000000000-4200010112000000000000000110-11101210000020001100 Tolmerus_atricapillus 0023012020000110000000100110011011200111000000000011001001000101020121011101011011202102111111011210100110-012020201000001110000001020112000000010101000000000011111000203-0200001100010111110000000100-100110000000021011-0 Trichardis_effrena 0024010020110110000000100010111011101111010001000011021012000200020121012011011100201102111010111011010110-002010102001000110000001011112001010110101010000000022222000000-5200001121000022222003010002000101100000022101110 Trichoura_sp_Tierberg 0011010120110210001000100110111021101110000001011011000002000101020122011201001000200002111111011200001100-01202120000000011000001001011210200001010100000000000000000021602110001100001000000000000102022011200000010000000 Willistonina_bilineata 0011010120110210001001100110011011001110000001011011021012000201020121011301001000200003111011011200000100-01202120000000111000001001000010000001010100100000000000000021402110001121000200000000000200-10011100000010000101 ; END; [ The following blocks are output data for analysis step 4922 ] BEGIN TREES; TITLE Tb10330; LINK TAXA = TaxaForAnalysisStep4922; TREE Fig._4 = [&R] (Bombylius_major,(((Mitrodetus_dentitarsis,(Opomydas_townsendi,Afroleptomydas_sp_Clanwilliam)),(Nemomydas_brachyrhynchus,Mydas_clavatus))Mydidae,(Prorates_sp_Escalante,((Hemigephyra_atra,Phycus_frommeri)Therevidae,(Apsilocephala_longistyla,(Apiocera_painteri,((((Atomosia_puella,Cerotainia_albipilosa),(Perasis_transvaalensis,((Laphria_aktis,Choerades_bella),(Pilica_formidolosa,(Nusa_infumata,(Laxenecera_albicincta,(Laphystia_tolandi,(Hoplistomerus_nobilis,Trichardis_effrena)))),(Lamyra_gulo,Stiphrolamyra_angularis)))))Laphriinae,(((Rhipidocephala_sp_HaroldJohnson,((Holcocephala_abdominalis,Holcocephala_calva),(Damalis_monochaetes,Damalis_annulata)))Trigonomiminae,(Eudioctria_albius,(Dioctria_atricapillus,(Dioctria_hyalipennis,Dioctria_rufipes)))Dioctriinae),(Ceraturgus_fasciatus,((Cyrtopogon_rattus,(Nicocles_politus,(Leptarthrus_brevirostris,Rhabdogaster_pedion))),(Lycostommyia_albifacies,(Tillobroma_punctipennis,((Diogmites_grossus,(Molobratia_teutonus,Pegesimallus_laticornis)),(Lestomyia_fraudiger,(Dasypogon_diadema,Saropogon_luteus,(Plesiomma_sp_Guanacaste,(Willistonina_bilineata,(Acnephalum_cylindricum,(Ablautus_coquilletti,(Trichoura_sp_Tierberg,((Lasiopogon_aldrichii,Lasiopogon_cinctus),(Stichopogon_trifasciatus,(Stichopogon_elegantulus,Stichopogon_punctum)))Stichopogoninae)))),(Connomyia_varipennis,(Scylaticus_costalis,(Ospriocerus_aeacus,(Stenopogon_rufibarbis,Gonioscelis_ventralis))),(Microstylum_sp_Karkloof,Prolepsis_tristis)))))))))))),((Lasiocnemus_lugens,(Leptogaster_cylindrica,(Leptogaster_arida,(Tipulogaster_glabrata,(Beameromyia_bifida,Euscelidia_pulchra)))))Leptogastrinae,(((Emphysomera_conopsoides,Emphysomera_pallidapex),(Afroestricus_chiastoneurus,Ommatius_tibialis))Ommatiinae,(Pogonioefferia_pogonias,Proctacanthus_philadelphicus,(Asilus_crabroniformis,Asilus_sericeus,(Philonicus_albiceps,(Neoitamus_cyanurus,(Clephydroneura_sp_Kepong,(Machimus_occidentalis,(Dysmachus_trigonus,Tolmerus_atricapillus)))))),(Dasophrys_crenulatus,Neolophonotus_bimaculatus),(Megaphorus_pulchrus,(Philodicus_tenuipes,Promachus_amastrus)))Asilinae)))Asilidae)))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr6056] END;