#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on April 17, 2026; 4:12 GMT

TreeBASE (cc) 1994-2008

Study reference:
Hart M., Jeffrey abt C., & Emlet R. 2011. Molecular phylogeny of echinometrid sea
urchins: new species of ‘Heliocidaris’ with derived modes of reproduction. Invertebrate
Biology, .

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S10972]

[ The following blocks are input data for analysis step 6616 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep6616;
        DIMENSIONS NTAX=12;
        TAXLABELS
            Echinometra_mathaei_1
            Echinometra_mathaei_2
            Echinometra_oblonga_1
            Echinometra_oblonga_2
            Echinometra_species_A_1
            Echinometra_species_A_2
            Heliocidaris_erythrogramma_1
            Heliocidaris_erythrogramma_2
            Heliocidaris_tuberculata_1
            Pachycentrotus_australiae_1
            Pachycentrotus_bajulus_1
            Pachyentrotus_bajulus_2
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M7006]

	TITLE COI;
	LINK TAXA = TaxaForAnalysisStep6616;
	DIMENSIONS NCHAR=510;
	FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -;

MATRIX
[                                        10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510]
[                                         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .]
Echinometra_mathaei_1            AAAGATGACCAAATCTACAAAG?GGTCGTTACAGCACACGCATTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGTAACTGGTTAATCCCTCTAATGATTGGAGCACCAGACATGGCCTTTCCCCGAATGAAAAACATGAGCTTTTGACTAGTTCCTCCTTCCTTCATTTTATTACTAGCCTCGGCAGGAGTAGAAAGTGGAGCAGGAACAGGCTGAACAATCTATCCTCCACTATCAAGAAAAATTGCCCATGCTGGAGGATCCGTAGACCTAGCAATTTTCTCCCTTCATCTTGCCGGTGCCTCATCTATTCTAGCTTCTATAAATTTTATTACAACTATCATAAATATGCGAACACCAGGAATGTCCTTTGACCGGCTCCCACTGTTCGTATGATCCGTATTTGTAACTGCATTTCTGCTACTTTTATCTCTTCCAGTATTAGCTGGAGCAATAACAATGCTTCTTACAGATCGAAATATTAACACAACA
Echinometra_mathaei_2            AAAGACGACCAAATCTACAAAGTGGTCGTTACAGCACACGCATTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGTAACTGGTTAATCCCTCTAATGATTGGAGCACCAGACATGGCCTTTCCCCGAATGAAAAACATGAGCTTTTGACTAGTTCCTCCTTCCTTCATTCTATTACTAGCCTCGGCAGGAGTAGAAAGTGGAGCAGGAACAGGCTGAACAATCTATCCTCCACTATCAAGAAAAATTGCCCATGCTGGAGGATCCGTGGACCTAGCAATTTTCTCCCTTCATCTTGCCGGTGCCTCATCTATTCTAGCTTCTATAAATTTTATTACAACTATCATAAATATGCGAACACCAGGAATGTCTTTTGACCGACTCCCACTGTTCGTATGATCCGTATTTGTAACTGCATTTCTGCTACTTTTATCTCTTCCAGTATTAGCTGGAGCAATAACAATGCTTCTTACAGATCGAAATATTAACACAACA
Echinometra_oblonga_1            AAAGACGACCAAATCTACAAAGTGGTCGTTACAGCACACGCATTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGAAACTGGTTAATCCCTCTGATGATTGGAGCACCAGACATGGCCTTTCCCCGAATGAAAAACATGAGCTTTTGACTAGTTCCTCCTTCCTTCATTTTATTACTAGCCTCGGCAGGAGTAGAAAGCGGAGCAGGAACAGGCTGAACAATCTATCCTCCACTATCAAGAAAAATTGCCCATGCTGGAGGATCTGTAGACCTAGCAATTTTCTCCCTTCATCTTGCCGGTGCCTCATCTATTCTAGCCTCTATAAATTTTATTACAACTATTATAAATATGCGGACACCAGGAATGTCCTTTGACCGGCTCCCACTGTTCGTATGATCTGTATTTGTAACTGCATTTCTACTACTTTTATCTCTTCCAGTATTAGCTGGGGCAATAACAATGCTTCTCACAGATCGAAATATTAACACAACA
Echinometra_oblonga_2            AAAGACGACCAAATCTACAAAGTGGTCGTTACAGCACACGCATTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGAAACTGGTTAATCCCTCTGATGATTGGAGCACCAGACATGGCCTTTCCCCGAATGAAAAACATGAGCTTTTGACTAGTTCCTCCTTCCTTCATTTTATTACTAGCCTCGGCAGGAGTAGAAAGTGGAGCAGGAACAGGCTGAACAATCTATCCTCCACTATCAAGAAAAATTGCCCATGCTGGAGGATCTGTAGACCTAGCAATTTTCTCCCTTCATCTTGCCGGTGCCTCATCTATTCTAGCTTCTATAAATTTTATTACAACTATTATAAATATGCGGACGCCAGGAATGTCCTTTGACCGGCTCCCACTGTTCGTATGATCTGTATTTGTAACTGCATTTCTACTACTTTTATCTCTTC?AGTATTAGCTGGGGCAATAACAATGCTTCTCACAGATCGAAATATTAACACAACA
Echinometra_species_A_1          AAAGACGACCAAATCTACAAAGTGGTCGTTACAGCACACGCATTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGTAACTGATTAATCCCTCTGATGATTGGAGCACCAGACATGGCCTTTCCCCGAATGAAAAATATGAGCTTTTGACTAGTTCCTCCTTCTTTCATTTTATTACTAGCCTCAGCAGGAGTAGAGAGTGGAGCAGGAACAGGCTGAACAATCTATCCTCCACTATCAAGGAAAATTGCCCATGCTGGAGGGTCTGTAGACCTAGCAATTTTCTCCCTTCATCTTGCCGGTGCCTCATCTATTCTAGCTTCTATAAATTTTATTACAACTATTATAAATATGCGAACACCAGGAATGTCCTTTGACCGACTCCCATTGTTCGTTTGGTCCGTATTTGTGACTGCATTTCTACTACTTTTATCTCTTCCGGTACTAGCTGGAGCAATAACAATGCTTCTTACAGATCGAAATATTAACACAACA
Echinometra_species_A_2          AAAGACGACCAAATCTACAAAGTGGTCGTTACAGCACACGCATTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGTAACTGATTAATCCCTCTGATGATTGGAGCACCAGACATGGCCTTTCCCCGAATGAAAAATATGAGCTTTTGACTAGTTCCTCCTTCTTTCATTTTATTACTAGCCTCAGCAGGAGTAGAAAGTGGAGCAGGAACAGGCTGAACAATCTATCCTCCACTATCAAGGAAAATTGCCCATGCTGGAGGGTCTGTAGACCTAGCAATTTTCTCCCTTCATCTTGCCGGTGCCTCATCTATTCTAGCTTCTATAAATTTTATTACAACTATTATAAATATGCGAACACCAGGAATGTCCTTTGACCGACTCCCATTGTTCGTTTGGTCCGTATTTGTGACTGCATTTCTACTACTTTTATCTCTTCCGGTACTAGCTGGAGCAATAACAATGCTTCTTACAGATCGAAATATTAACACAACA
Heliocidaris_erythrogramma_1     AAAGATGACCAAATTTACAAAGTAGTTGTTACAGCACACGCACTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGGTTTGGAAACTGACTTATACCACTAATGATTGGTGCACCAGACATGGCCTTTCCACGAATGAAAAATATGAGTTTTTGGCTAATCCCACCATCCTTCATTTTACTCTTAGCTTCCGCAGGAGTAGAAAGAGGAGCGGGGACTGGTTGAACTATTTATCCACCCCTATCTAGAAAAATAGCACACGCCGGTGGATCAGTCGATCTAGCAATCTTTTCTCTCCATCTTGCTGGTGCCTCCTCAATCCTAGCCTCCATAAACTTTATAACGACAATTATTAACATGCGAACACCAGGAATGTCTTTCGACCGACTTCCCCTATTTGTTTGATCAGTCTTTGTAACAGCCTTTTTACTTCTCCTATCTCTACCAGTTCTTGCAGGTGCAATTACTATGCTCCTAACAGACCGAAATATAAATACAACC
Heliocidaris_erythrogramma_2     AAAGATGACCAAATTTACAAAGTAGTTGTTACAGCACACGCACTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGGTTTGGAAACTGACTTATACCACTAATGATTGGTGCACCAGACATGGCCTTTCCACGAATGAAAAATATGAGTTTTTGGCTAATCCCACCATCCTTCATTTTACTCTTAGCTTCCGCAGGAGTAGAAAGAGGAGCGGGGACTGGTTGAACTATTTATCCGCCCCTATCTAGAAAAATAGCACACGCCGGTGGATCAGTCGATCTAGCAATCTTTTCTCTCCATCTTGCTGGTGCCTCCTCAATCCTAGCCTCCATAAACTTTATAACGACAATCATTAACATGCGAACACCAGGTATGTCTTTCGACCGACTTCCCCTATTTGTTTGATCAGTCTTTGTAACAGCCTTTTTACTTCTCCTATCTCTACCAGTTCTTGCAGGTGCAATTACTATGCTCCTAACAGACCGAAATATAAATACAACC
Heliocidaris_tuberculata_1       AAAGATGACCAAATTTACAAAGTAGTCGTTACCGCACACGCACTGGTTATGATTTTCTTCATGGTAATGCCAATAATGATTGGAGGGTTTGGAAACTGACTTATTCCACTAATGATTGGTGCCCCAGACATGGCCTTCCCCCGTATGAAAAAAATGAGTTTCTGGCTAATCCCCCCTTCTTTTATCTTACTATTAGCTTCAGCAGGAGTAGAAAGAGGAGCAGGAACAGGCTGGACTATTTACCCACCTCTGTCAAGAAAAATAGCACATGCTGGTGGATCTGTTGACCTAGCGATCTTTTCTCTCCACCTTGCCGGTGCATCCTCAATTCTGGCCTCCATCAACTTCATAACAACAATTATTAACATGCGAACACCAGGAATGTCTTTTGACCGACTACCCCTATTCGTATGATCAGTCTTTGTAACAGCCTTTTTGCTGCTCCTTTCCCTACCGGTCCTAGCAGGTGCAATTACTATGCTCTTAACAGACCGAAACATTAACACAACC
Pachycentrotus_australiae_1      AAAGATGACCAAATTTACAAAGTAATTGTTACAGCACATGCACTAGTTATGATCTTTTTTATGGTAATGCCAATAATGATTGGAGGATTTGGAAACTGACTCATACCACTAATGATTGGTGCACCAGACATGGCCTTTCCACGAATGAAAAATATGAGTTTTTGGCTAATTCCACCATCTTTCATTTTACTCTTAGCTTCCGCAGGAGTAGAAAGAGGAGCAGGGACTGGTTGAACTATTTATCCACCCCTATCTAGAAAAATAGCACATGCCGGAGGATCGGTCGATCTAGCAATCTTTTCTCTCCATCTTGCTGGGGCCTCTTCAATCCTAGCCTCCATAAACTTTATAACAACAATTATTAACATGCGAACACCAGGAATGTCTTTTGACCGACTTCCACTATTTGTCTGATCAGTCTTTGTAACAGCCTTTTTACTTCTTCTATCTCTGCCAGTTCTTGCCGGTGCGATTACTATGCTCCTAACAGACCGAAATATAAATACAACC
Pachycentrotus_bajulus_1         AAAGATGACCAAATTTATAAAGTAATTGTTACAGCACATGCACTAGTTATGATATTTTTTATGGTAATGCCAATAATGATTGGAGGGTTTGGAAACTGACTTATTCCACTAATGATTGGTGCACCAGACATGGCCTTTCCACGAATGAAAAACATGAGTTTTTGACTAATTCCACCCTCTTTTATTTTACTCTTAGCTTCCGCAGGAGTAGAAAGAGGAGCAGGGACTGGTTGAACTATTTATCCTCCTCTATCCAGAAAAATAGCACATGCCGGTGGATCAGTTGATCTAGCAATCTTTTCTCTCCATCTTGCTGGTGCCTCCTCAATCCTTGCCTCCATAAACTTTATAACAACAATTATTAACATGCGAACACCAGGAATGTCTTTCGATCGACTTCCCCTATTTGTTTGATCAGTCTTTGTAACAGCCTTTTTACTTCTATTATCTCTACCAGTTCTTGCCGGTGCAATTAATATGCTTCTAACAGACCGAAACATAAATACAACT
Pachyentrotus_bajulus_2          AAAGATGACCAAATTTATAAAGTAATTGTTACAGCACATGCACTAGTTATGATATTTTTTATGGTAATGCCAATAATGATTGGAGGGTTTGGAAACTGACTTATTCCACTAATGATTGGTGCACCAGACATGGCCTTTCCACGAATGAAAAACATGAGTTTTTGACTAATTCCACCCTCTTTTATTTTACTCTTAGCTTCCGCAGGAGTAGAAAGAGGAGCAGGGACTGGTTGAACTATTTATCCTCCTCTATCCAGAAAAATAGCACATGCCGGTGGATCAGTTGATCTAGCAATCTTTTCTCTCCATCTTGCTGGTGCCTCCTCAATCCTTGCCTCCATAAACTTTATAACAACAATTATTAACATGCGAACACCAGGAATGTCTTTCGATCGACTTCCCCTATTTGTTTGATCAGTCTTTGTAACAGCCTTTTTACTTCTATTATCTCTACCAGTTCTTGCCGGTGCAATTACTATGCTTCTAACAGACCGAAACATAAATACAACT
;
END;

[ The following blocks are output data for analysis step 6616 ]
BEGIN TREES;
      TITLE COI_TB;
      LINK TAXA = TaxaForAnalysisStep6616;
      TREE ML = [&R] ((((Pachycentrotus_australiae_1,(Pachycentrotus_bajulus_1,Pachyentrotus_bajulus_2)),(Heliocidaris_erythrogramma_1,Heliocidaris_erythrogramma_2)),Heliocidaris_tuberculata_1),(Echinometra_mathaei_1,(Echinometra_mathaei_2,((Echinometra_oblonga_1,Echinometra_oblonga_2),(Echinometra_species_A_2,Echinometra_species_A_1)))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr25492]


END;