#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on April 25, 2024; 14:36 GMT

TreeBASE (cc) 1994-2008

Study reference:
Beutel R.G., Friedrich F., Hörnschemeyer T., Pohl H., Hünefeld F., Beckmann F., Meier
R., Misof B., Whiting M., & Vilhelmsen L. 2011. Morphological and molecular
evidence converge upon a robust phylogeny of the megadiverse Holometabola. Cladistics,
27: 341-355.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S11976]

[ The following blocks are input data for analysis step 8002 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep8002;
        DIMENSIONS NTAX=34;
        TAXLABELS
            Agathiphagidae
            Bibionidae
            Bittacidae
            Boreus
            Caeciliusidae
            Caurinus
            Chrysopidae
            Corydalidae
            Culicidae
            Cupedidae
            Diprionidae
            Eriocraniidae
            Helophoridae
            Inocelliidae
            Limnephilidae
            Mengenillidae
            Micropterigidae
            Nannochoristidae
            Nevrorthidae
            Ommatidae
            Osmylidae
            Panorpidae
            Pteronarcyidae
            Pulicidae
            Raphidiidae
            Rhyacophilidae
            Sialidae
            Tenthredinidae
            Tettigoniidae
            Tipulidae
            Trachypachidae
            Xenidae
            Xyelidae
            Zorotypidae
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M10442]

	TITLE CLA_338_sm_HolometAppIV;
	LINK TAXA = TaxaForAnalysisStep8002;
	DIMENSIONS NCHAR=356;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5" MISSING=? GAP= -;
	CHARSTATELABELS
		1  Orientation_of_head / orthognathous prognathous_or_slightly_inclined,
		2  'Shape of anterior (dorsal) side of head' / distinctly_convex 'very slightly convex or flat, nearly parallel to ventral (posterior) side',
		3  Neck_region / absent distinctly_developed,
		4  Split_cranial_setae / absent present,
		5  Lateral_eyes / 'well developed, with or without crystalline cones' 'distinctly reduced, with ca. 10 ommatidia and lacking a corn' well_developed_stemmata stemmata_closely_aggregated 'strongly reduced, 1-2 stemmata or single eyespot',
		6  Number_of_retinula_cells_in_ommatidia_or_stemmata / less_than_15 15_or_more,
		7  Paired_ocelli / absent present,
		8  'Transverse facial strengthening line (frontoclypeal suture)' / absent_or_only_vaguely_indicated present_and_distinct,
		9  Division_of_clpyeal_area_into_anteclyepus_and_postclypeus / absent present,
		10  Shape_of_clypeal_part_anterad_of_transverse_strengthening_line / broader_than_long 'longer than wide, trapezoid, anteriorly converging',
		11  Occipital_furrow / absent present,
		12  Gula / absent undivided_sclerotized_quadrangular_gula strongly_narrowed_gula,
		13  Hypostomal_bridge / absent 'present, not sperated from genal area' 'present, distinctly separated from genal area',
		14  Posterior_tentorial_grooves / close_to_hind_margin_of_head_capsule shifted_anterad,
		15  Dorsal_tentorial_arm / 'well developed, sclerotized' 'strongly reduced and ligament-like or absent',
		16  Shape_of_tentorium / 'not X-shaped, without constriction between ant and post  arms' 'X-shaped, with distinct constriction between ant and post  arms' largely_or_completely_reduced,
		17  Dorsoventral_muscle_spanning_foramen_occipitale / absent present,
		18  Articulation_of_labrum / free fused_with_clypeus,
		19  Exposure_of_anterior_epipharynx / not_exposed slightly_exposed distinctly_exposed,
		20  M._frontolabralis_with_attachment_to_external_wall_of_labrum / absent present,
		21  M._frontoepipharyngalis / absent 'present, inserted on tormae or posterolaterally on epipharynx' 'present, inserted close to median line' 'present, attached to messores',
		22  Antennal_segmentation / 'multisegmented, with antenommeres distinctly separated' 'pseudo-multisegmented, with mid-segment subdivided' '5-7 antennomeres' 4_antennomeres 3_antennomeres less_than_3_antennomeres,
		23  Sensorium_on_antepenultimate_antennomere / absent present,
		24  'Specialised terminal seta of flagellum (FITS)' / absent present,
		25  Number_of_extrinsic_antennal_muscles / 4. 3. 2. 1. 0.,
		26  'Intrinsic antennal muscles (Mm. scapopedicellaris)' / absent present,
		27  Mola / absent_or_strongly_reduced present,
		28  Articulated_lacinia_mobilis / absent present,
		29  'Mandibulo-maxillary sucking apparatus' / absent present,
		30  Poison_channel_of_maxillary_stylet / absent present,
		31  Position_of_maxilla / retracted 'protracted, cardines at level of prementum, max groove absen',
		32  Maxillolabial_complex / absent present,
		33  Shape_of_the_proximal_parts_of_the_maxilla / not_transverse transverse,
		34  'Posteromesal process of cardo/proximal element of the maxilla' / absent present,
		35  Galea_and_lacinia / present_as_discrete_structures extensively_or_completely_united,
		36  Stipital_flexor_muscles_of_lacinia_and_galea / both_present one_muscle absent,
		37  M._craniocardinalis / well_developed absent,
		38  Maxillary_palp / absent present,
		39  M._craniodististipitalis / absent present,
		40  M._submentopraementalis / absent present,
		41  M._praementoparaglossalis / absent present,
		42  M._praementoglossalis / absent present,
		43  Apical_prelabial_region / without_process_or_extension_or_with_short_unsclerotised_ligula 'with wedge-shaped, strongly sclerotised ligula' sclerotised_prelabial_extension_distinctly_developed together_with_hypopharynx_foming_compact_lobe_with_salivary_,
		44  Salivary_duct / absent well_developed 'strongly narrowed, without recognisable lumen',
		45  Cervix / absent present,
		46  Legs / absent present,
		47  Number_of_leg_segments / more_than_4 3.,
		48  Claws / double single absent,
		49  Larval_segment_XI / absent present,
		50  'Retractile prolegs on segments II-VII' / absent present,
		51  'Conical ventral protuberances on segments I-VIII' / absent present,
		52  Setiferous_lateral_filaments / absent present,
		53  Cerci / absent present,
		54  Urogomphi_on_segment_IX / absent present,
		55  Terminal_hooks / absent single_hook_on_fleshy_lobes_of_segment_X single_hook_on_subdivided_prolegs_with_scerotised_elements double_hooks_on_unsclerotised_proleg,
		56  Malpighian_tubules / all_free several_cryptonephric_malpighian_tubules,
		57  Larval_habitat / terrestrial semiaquatic aquatic,
		58  Endoparasitism_of_larvae / absent present,
		59  Movability_of_pupal_mandible / absent present,
		60  Size_of_pupal_mandible / not_hypertrophied hypertrophied,
		61  Exposure_of_posterior_head_region / fully_exposed at_least_partly_retracted,
		62  Orientation_of_head / orthognathous proganthous_or_slightly_inclined,
		63  Rostrum / absent_or_extremely_short 'elongated, not including maxillary and labial elements' 'elongated, including maxillary and labial elements',
		64  Frontal_apodeme / absent present,
		65  Shape_of_posterior_head_capsule / not_concave concave,
		66  Postgenal_bridge / absent 'present, not delimited laterally' 'present, laterally delimited',
		67  Dense_vestiture_of_microtrichia / absent present,
		68  Clypeus / not_inflected inflected,
		69  Shape_of_posterior_tentorium / 'not collar-like' 'collar-like' completely_absent,
		70  Movability_of_labrum / absent present,
		71  Mouthfield_sclerite / absent present,
		72  'Labro-epipharyngeal food canal' / absent present,
		73  M._frontolabralis / absent 'present, origin on frons' 'present, origin on clypeus',
		74  M._frontoepipharyngalis / absent present,
		75  Insertion_of_antenna / 'anteriorly between compound eyes, adjacent' 'anteriorly between compound eyes, not adjacent' laterally,
		76  Number_of_antennomeres / more_than_11 '11 or less, flagellomeres not forming clava' '11 or less, flagellomeres forming clava' '8 or less, flagellomeres flabellate',
		77  'Intercalary sclerite in lateral scapo-pedicellar membrane' / absent present,
		78  Shape_and_size_of_mandible / 'not elongated and blade-like or lamelliform' 'elongated, flattened or lamelliform' 'blade-like',
		79  Mandibular_mola / distinctly_developed strongly_reduced_or_absent,
		80  Haustellum / absent present,
		81  'Labio-maxillary complex' / absent present,
		82  Dorsal_concavitiy_of_anterior_labium_for_reception_of_elements_of_the_paired_mouthparts / absent present,
		83  Cardines / not_fused_to_labial_elements fused,
		84  Fusion_of_stipites_and_anterior_postmentum / absent present,
		85  Galea / vestigial_or_absent distinctly_developed,
		86  Galea_enfolds_lateral_part_of_labrum_and_lateral_mandibular_base / absent present,
		87  Shape_of_lacinia / 'not elongated and blade-like' 'elongated and blade-like',
		88  Salivary_channel_of_lacinia / absent present,
		89  Sensorium_on_3rd_maxillary_palpomere / absent present,
		90  M._craniocardinalis / absent present,
		91  Origin_of_M._tentoriocardinalis_and_M._tentoriostipitalis / tentorium frontoclypeal_region laterally_from_the_head_capsule,
		92  Flexion_points_between_maxillary_palpomeres_1_and_2_and_3_and_4 / absent present,
		93  Arched_sclerite_with_long_piliform_scales_formed_by_postlabium / absent present,
		94  Internal_ridge_of_maxillolabial_plate_or_postlabium / absent present,
		95  Number_of_labial_palpomeres / 3. 5. 2_or_less,
		96  Secondary_subdivision_of_apical_labial_palpomere / absent indistinct 'distinct, appearing as two separate palpomeres',
		97  Shape_of_apical_labial_palpomere / 'not spoon-shaped' 'spoon-shaped',
		98  Paraglossae / 'vestigial or absent, without muscles' present,
		99  M._praementoparagossalis / absent present,
		100  M._praementoglossalis / absent present,
		101  Size_of_Mm._praementopalpales / not_enlarged enlarged,
		102  Sclerotised_sitophore_plate / absent present,
		103  Epipharyngopharyngeal_lobe_reaching_into_pharynx_posteriorly / absent present,
		104  Size_of_M._clypeopalatalis / not_enlarged 'enlarged, two major subcomponents and multiple bundles' arranged_as_a_longitudinal_row_of_small_bundles,
		105  Transverse_muscle_of_the_epipharynx / absent present,
		106  Precerebral_pharyngeal_pumping_chamber_within_chitinous_clasps / absent present,
		107  Postcerebral_pharyngeal_pumping_apparatus / absent present,
		108  Strongly_developed_intrinsic_muscle_of_salivarium / absent present,
		109  Configuration_of_brain_and_suboesophageal_complex / distinctly_separated_by_the_circumoesophageal_connectives forming_a_compact_ganglionic_mass_around_the_pharynx,
		110  Membranes_interconnecting_thoracic_sclerites / exposed 'reduced, not exposed',
		111  Dorsal_cervical_sclerite / absent present,
		112  Lateral_cervical_sclerite / absent single_pair_of_sclerites_present two_pairs_of_sclerites_present,
		113  Posteromedian_corner_of_lateral_cervical_sclerite / not_protruding protruding_towards_prosternum 'fused in midline (forming a jugulare)',
		114  Lateral_cervical_sclerite_and_propleuron / not_fused partly_fused entirely_fused,
		115  Length_of_pronotum / shorter_than_pterothoracic_nota longer_than_pterothoracic_nota,
		116  Lateral_connection_of_pronotum_and_propleuron / absent partly_or_completely_connected,
		117  'Prothoracic-mesopleural connection' / absent pronotal_and_anterior_mesepisternal_process_connected posterior_propleuron_connected_to_anterior_mesepisternum pronotum_and_propleura_connected_by_intersegmental_sclerites,
		118  Externally_visible_part_of_prothoracic_basisternum / small_or_absent large,
		119  Prothoracic_precoxal_bridge / absent present 'basisternum and propleuron attached, but not fused',
		120  Profurcal_arm_and_propleura / not_connected 'not fused, connected by muscle' firmly_fused specifically_articulated,
		121  Profurca_with_free_dorsal_arms / absent present,
		122  Prospina / absent present,
		123  Protrochantin / absent distinctly_reduced well_developed fused_with_propleura,
		124  'Antenna-cleaning apparatus of fore leg' / absent 'formed by pre-apical tibial spur and the distal part of the pr' 'formed by an apical tibial spur and the inner side of the pro-basitar' formed_by_2_protibial_spurs_and_a_comb_of_smaller_spines_between_t,
		125  Movable_epiphysis_of_fore_tibia / absent present,
		126  Relative_size_of_pterothoracic_segments / 'th2=th3' 'th2>th3' 'th2<th3',
		127  Relative_size_of_wings / equally_sized fore_wings_distinctly_larger hind_wings_distinctly_larger,
		128  Median_mesonotal_suture / absent present,
		129  Membranous_area_between_mesoscutellum_and_mesopostnotum / absent present,
		130  Specific_locking_device_of_mesoscutellum / absent present,
		131  Mesothoracic_postalar_bridge / absent 'present, mesopostnotum and mesepimeron firmly fused',
		132  Size_of_mesothoracic_postalar_bridge / broad small,
		133  Lateral_part_of_mesopostnotum / 'small, pointed' 'enlarged, tongue-like',
		134  Mesothoracic_prealare / absent present,
		135  Mesothoracic_tegular_arm / absent present,
		136  Elongated_ventral_process_of_mesothoracic_pleural_ridge / absent present,
		137  Mesothoracic_meron_and_epimeron / separate broadly_fused,
		138  Mesosternum / externally_exposed invaginated,
		139  'Mesofurca and -epimeron' / disrete fused,
		140  'Ventral mesosternal process (below mesofurca) forming sterno-coxal joint' / absent present,
		141  Size_of_ventral_mesosternal_process / not_elongated elongated,
		142  Mesospina / absent present,
		143  Proximal_part_of_mesocoxae / recessed_into_coxal_cavities not_recessed_into_coxal_cavities,
		144  Mesocoxae / distinctly_sepapared_from_each_other closely_adjacent_medially,
		145  Mesothoracic_sternocoxale / 'continuous with anterior eucoxa, articulated with post. part' 'continuous with posterior eucoxa, articulated with ant. part' anteriorly_and_posteriorly_articulated,
		146  Mesocoxal_meron / absent present,
		147  Mesotrochantin / absent present,
		148  Size_of_mesotrochantin / well_developed small,
		149  Transverse_ridge_of_mesoscutum / absent present,
		150  Alacristae / absent present,
		151  Metathoracic_postalar_bridge / absent 'present, metapostnotum and metepimeron firmly fused',
		152  Metathoracic_tegular_arm / absent present,
		153  Metapostnotum / single_sclerite medially_divided,
		154  Size_of_metapostnotum / small_to_moderately_sized 'strongly elongated, shield-like',
		155  Metapostnotum_and_first_abdominal_tergum / not_fused firmly_fused,
		156  Metathoracic_katepisternum / absent present,
		157  Elongated_ventral_process_of_metathoracic_pleural_ridge / absent present,
		158  Metasternum / externally_exposed invaginated,
		159  'Fusion of metafurca and -epimeron' / absent present,
		160  Position_of_metafurca / posterior_end_of_discriminal_ridge anterior_end_of_discriminal_ridge,
		161  Anteromedian_metafurcal_process_closely_associated_with_metadiscrimen / absent present,
		162  'Ventral metasternal process (below metafurca) forming sterno-coxal joint' / absent present,
		163  Size_of_ventral_metasternal_process / not_elongated elongated,
		164  Connection_between_metafurcal_arm_and_epimeral_apophysis / no_connection muscular_connection connected_by_tendon completely_fused,
		165  Metatrochantin / absent 'present, externally visible' internalised,
		166  Orientation_of_metacoxae / not_transverse transverse,
		167  Metacoxae / distinctly_delimited_from_each_other adjacent_or_allmost_adjacent_medially,
		168  Metacoxal_meron / absent present,
		169  Metathoracic_meron_and_epimeron / separate broadly_fused,
		170  Metathoracic_sternocoxale / 'continuous with anterior eucoxal sclerotisation, articulated with po' 'continuous with posterior coxal sclerotisation, articulated with ante' articulated_with_anterior_and_posterior_part_of_metacoxa,
		171  Fore_wings / wings elytra halteres clasping_organs,
		172  Hind_wings / wings halteres 'peg-like sclerites',
		173  Transverse_hindwing_folding_mechanism / absent present,
		174  Number_of_costal_cross_veins / more_than_five less_than_five,
		175  Dense_vestiture_of_scales_on_wings / absent present,
		176  Hamuli / absent 'present, connecting fore wings and hind wings',
		177  'Insertion site of the anterior pronoto-cervical muscle' / lateral_cervical_sclerite_or_posterolateral_head_capsule corpotentorium,
		178  'M. profurca-phragmalis' / absent present,
		179  'Origin of M. profurca-phragmalis' / profurca posteriormost_part_of_propleura,
		180  'M. pronoto-coxalis posterior' / absent present,
		181  'M. pronoto-trochanteralis' / absent present,
		182  'Origin propleura-occipital muscle' / anterodorsal_edge_of_propleuron close_to_propleural_ridge propleural_apodeme profurca,
		183  'M. prospina-mesopleuralis' / absent present,
		184  'M. procoxa-cervicalis' / absent present,
		185  'Origin of M. procoxa-cervicalis' / lateral_cervical_sclerite corpotentorium,
		186  'Orientation of both bundles of M. procoxa-cervicalis' / parallel_to_each_other crossing_each_other,
		187  'Origin of M. profurca-tentorialis' / laterally_on_corpotentorium posteroventrally_on_head_capsule gula anteroventral_area_of_cervical_membrane,
		188  'M. profurca-spinalis' / absent 'present, connecting profurca and -spina' 'present, interconnecting profurcal arms',
		189  'M. profurca-mesospinalis' / absent present,
		190  'M. profurca-mesofurcalis' / strongly_reduced_or_absent present,
		191  'M. prospina-mesofurcalis' / absent present,
		192  'M. profurca-coxalis medialis' / absent present,
		193  'M. prospina-coxalis' / absent present,
		194  'M. mesoscutello-postnotalis' / absent present,
		195  'M. mesonoto-sternalis' / absent present,
		196  'M. mesonoto-coxalis posterior' / absent present,
		197  'M. mesonoto-trochanteralis' / absent present,
		198  'M. mesofurca-phragmalis' / absent present,
		199  'Insertion of M. mesofurca-phragmalis' / muscular by_long_thin_tendon,
		200  'Size of M. prophragma-mesanepisternalis' / strongly_developed moderately_sized_to_small,
		201  'Number of bundles of M. prophragma-mesanepisternalis' / 1. 2.,
		202  'M. mesonoto-basalaris' / absent present,
		203  'Insertion of M. mesonoto-basalaris' / immediately_mediad_of_basalar_stalk ventrolateral_part_of_basalar_disc,
		204  'M. mesonoto-pleuralis anterior' / absent present,
		205  'Number of distinct bundles of M. mesonoto-pleuralis anterior' / 1. 2.,
		206  'M. mesonoto-pleuralis medialis' / absent present,
		207  'Size of M. mesonoto-pleuralis medialis' / moderate large,
		208  'M. mesonoto-pleuralis posterior' / absent present,
		209  'Bundle of M. mesonoto-pleuralis posterior originating on dorsal third of pleural ridge' / absent present,
		210  'Bundle of M. mesonoto-pleuralis posterior originating on pleural arm' / absent present,
		211  'Insertion of M. mesonoto-pleuralis posterior' / posterolateral_mesoscutal_rim posterior_mesonotal_process fourth_axillary_sclerite,
		212  'M. mesanepisterno-axillaris' / absent present,
		213  'M. mesepimero-subalaris' / absent present,
		214  'M. mesopleura-subalaris' / absent present,
		215  'M. mesopleura-sternalis' / absent present,
		216  'M. mesofurca-pleuralis' / absent present,
		217  'Origin of M. mesofurca-pleuralis' / without_tendon with_a_tendon,
		218  'M. mesofurca-metanepisternalis' / absent present,
		219  'Insertion of M. mesobasalare-trochantinalis' / trochantin anterior_mesocoxal_rim,
		220  'M. mesanepisterno-coxalis posterior' / absent present,
		221  'M. mesospina-metafurcalis' / absent present,
		222  'M. metascutello-postnotalis' / absent present,
		223  'M. metanoto-sternalis' / absent present,
		224  'M. metanoto-coxalis posterior' / absent present,
		225  'M. metafurca-phragmalis' / absent present,
		226  'M. metanoto-basalaris' / absent present,
		227  'Insertion of M. metanoto-basalaris' / immediately_mediad_of_basalar_stalk ventrolateral_part_of_basalar_disc,
		228  'M. metanoto-pleuralis anterior' / absent present,
		229  'M. metanoto-pleuralis medialis' / absent present,
		230  'M. metanoto-pleuralis posterior' / absent present,
		231  'Insertion of M. metanoto-pleuralis posterior' / posterolateral_margin_of_metascutum posterior_metanotal_process,
		232  'M. metanepisterno-axillaris' / absent present,
		233  'M. metepimero-subalaris' / absent present,
		234  'M. metafurca-pleuralis' / absent present,
		235  'Insertion of M. metabasalare-trochantinalis' / metatrochantin anterior_metacoxal_rim,
		236  'Bundle of M. metafurca-trochanteralis with origin on discrimen' / absent present,
		237  Muscle_between_second_abdominal_sternum_and_metacoxa / absent present,
		238  Furcostigmal_muscle_between_metafurcal_arm_and_first_abdominal_stigma / absent present,
		239  Arolium / absent present,
		240  Ventral_hairy_soles_on_tarsomeres / absent 'present, microtrichia without pedestal' 'present, microtrichia with pedestal',
		241  Hairy_pulvilli / absent present,
		242  Tegula_in_the_fore_wing / absent present,
		243  Shape_of_anterior_notal_wing_process_of_the_mesonotum / nearly_triangular_in_shape elongate_triangular_tip_directed_anteriorly not_triangular_in_shape,
		244  Shape_of_the_posterior_notal_wing_process_of_the_mesonotum / 'long and slender process (at least two times longer than wide)' other_shape no_distinct_PNP_present,
		245  Shape_of_fore_wing_4Ax / 4Ax_absent 'approximately as wide as long, approximately triangular' 'clearly (more than 2 times) longer than wide' about_twice_as_long_as_wide,
		246  Distal_tip_of_fore_wing_4Ax / bent_anteriorly not_bent_anteriorly,
		247  Fore_wing_1Ax / individual_sclerite fused_to_notum_or_absent,
		248  Angle_between_distal_margins_of_body_and_neck_of_fore_wing_1Ax / less_than_120° between_120°_and_180° 180°_or_more,
		249  Shape_of_neck_of_fore_wing_1Ax / 'narrow, distal margin without projections' distal_margin_with_more_or_less_distinct_projection no_distinction_between_neck_and_body_possible,
		250  Transition_from_body_to_neck_in_fore_wing_1Ax / continuous 'neck and body distinctly separated (by incision or abrupt change o',
		251  Radial_plate_of_fore_wing / absent present,
		252  'Contact between base of radial vein (BR) and 2Ax of fore wing' / 'BR not fused to 2Ax, membranous intersection present' BR_approx_as_wide_as_2Ax_and_fused_to_it BR_approx_half_as_wide_as_2Ax_and_fused_to_it 'BR connected to 2Ax via a narrow sclerotized stripe (1\3 or less of the width of the 2Ax)',
		253  Contact_of_the_2Ax_to_the_1Ax_in_the_fore_wing / 'complete proximo-caudal part of 2Ax articulates with 1Ax' 'proximo-caudal part of 2Ax separated from 1Ax by a membranous ' 'articulation between 2Ax and 1Ax formed by proximo-cranial and di',
		254  Number_of_elements_of_fore_wing_3Ax / 1. 2.,
		255  Shape_of_fore_wing_3Ax / 'narrow, more than 4 times longer than wide' 'plate like, at most 3 times longer than wide' 'narrow, elongated, 3 to 4 times longer than wide, perpendicular pr',
		256  'Orientation of distal tip of fore wing 3Ax (wing extended)' / cranially_orientated distally_orientated,
		257  Caudal_arm_of_fore_wing_3Ax / 'no recognisable separation between caudal, distal and proximal ar' 'caudal arm recognisable, parallel-sided',
		258  Distance_of_the_articulation_of_the_distal_area_of_3Ax_from_the_notal_margin / at_least_2_times_the_maximum_width_of_the_1Ax less_than_2_times_the_maximum_width_of_the_1Ax,
		259  Median_plates_of_fore_wing / absent separate fused,
		260  'Length of mesothoracic posterior wing process (PWP)' / ends_on_the_same_level_as_basalare distinctly_overtops_basalare,
		261  Externally_visible_mesothoracic_subalare / with_process_in_the_middle_of_the_dorsal_margin without_process dorsally_deeply_emarginate absent,
		262  Shape_of_externally_visible_mesothoracic_subalare / aproximatly_as_long_as_high longer_than_high,
		263  Tegula_of_hind_wing / absent present,
		264  Subtegula_of_hind_wing / 'distinct, embracing anterior wing margin' small_sclerite_in_the_dorsal_wing_membrane absent,
		265  'Proportions of flat part of anterior notal wing process (ANP) of hind wing' / about_as_long_as_wide distinctly_wider_than_long clearly_longer_than_wide absent,
		266  Humeral_plate_of_hind_wing / separated_from_costal_vein_by_membrane formed_by_widened_end_of_costal_vein 'not recognisable, missing or indistinguishably fused to costal vein',
		267  Shape_of_anterior_notal_wing_process_of_hind_wing / nearly_triangular with_bifurcate_apex not_bifurcate_and_triangular,
		268  'Shape of posterior notal wing process (PNP) of hind wing' / 'long and slender process, at least 2 times longer than wide' long_and_widened_apically 'shorter, distinctly less than 2 times longer than wide' absent,
		269  4Ax_in_the_hind_wing / absent 'not U-shaped with the open side anteriorly directed' 'U-shaped with the open side anteriorly directed',
		270  Width_of_neck_of_hind_wing_1Ax / 'distinctly narrower than head region of 1Ax, with approximately stra' 'distinctly narrower than head region, distal margin concave' about_as_wide_as_indistinct_head_region very_slender_with_a_distinct_process_at_its_distal_margin extremely_short_or_absent,
		271  Orientation_of_head_and_neck_of_hind_wing_1Ax / head_and_neck_cranially_directed 'head and\or neck bent at about 90° distally' head_absent,
		272  'Proximo-caudal part of body of hind wing 1Ax' / 'distinctly longer than disto-caudal part' 'as long as disto-caudal part' 'part shorter than disto-caudal part',
		273  Distal_part_hind_wing_1Ax_body / 'curved, tip pointing caudally' tip_of_pointing_distally 'tip absent, distal corner blunt, rounded',
		274  Caudal_margin_of_body_of_hind_wing_1Ax / concave straight convex,
		275  Angle_alpha_between_metathoracic_1Ax_and_notal_margin / wider_than_50° between_25°_and_50° less_than_25°,
		276  Length_ratio_of_1Ax_and_notum / notum_more_than_3.8_times_longer 'notum 3-3.8 times longer' 'notum 2-3 times longer' 'notum 1.3-2 times longer',
		277  Angle_between_distal_margins_of_body_and_neck_of_hind_wing_1Ax / less_than_110° wider_than_110°,
		278  Shape_and_configuration_of_hind_wing_basiradiale / not_fused_with_2Ax at_least_half_as_wide_as_2Ax_and_fused_to_it less_than_half_as_wide_as_2Ax_and_fused_to_it,
		279  Position_of_contact_between_hind_wing_basiradiale_and_2Ax / middle_of_BR_cranial_margin 'proximo-cranial area of BR' 'disto-cranial area of 2Ax',
		280  Shape_of_hind_wing_2Ax_in_dorsal_view / triangular semicircular 'elongated and rod-shaped' irregular,
		281  Lateral_process_of_hind_wing_2Ax_extending_under_body_of_1Ax / absent present,
		282  Caudal_process_of_hind_wing_3Ax / absent present,
		283  'Contact between caudal process of hind wing 3Ax and posterior notal wing process (PNP) or 4Ax' / apical_parts_of_caudal_process_and_PNP_in_contact contact_extended_over_a_longer_distance '3Ax without notal contact area, articulates with caudo-distal tip of ',
		284  Shape_of_the_distal_process_of_hind_wing_3Ax / with_one_tip distal_process_absent 3Ax_fused_to_bases_of_anal_veins,
		285  Number_of_parts_of_hind_wing_3Ax / one_element more_than_one_element,
		286  Axillary_muscle_disc_of_hind_wing / absent present,
		287  'Shape of metathoracic posterior notal wing process (PNP) in ventro-lateral view' / large_sclerite_with_long_caudal_process 'not large and\or without caudal process',
		288  'Shape of metathoracic basalare (Ba)' / simple_plate ventral_plate_and_dorsal_extended_head extended_head_or_knob narrow_crescent_shaped_sclerite two_separate_simple_Ba_sclerites,
		289  Shape_of_dorsal_head_of_metathoracic_basalare / at_least_as_wide_as_the_widest_part_of_the_ventral_plate narrower,
		290  Process_of_anterior_margin_of_metathoracic_basalare / absent 'present, connected to the anterior wing margin',
		291  Shape_of_the_knob_of_the_metathoracic_basalare / simple_convexity very_small long_and_flat large_and_distended,
		292  Cavity_for_knob_of_metathoracic_basalare_at_ventral_side_of_hind_wing / 'ventral bases of subcosta (BSc) and humerus simple, pit absent' humerus_and_BSc_together_form_cavity_for_reception_of_basalar_kno cavity_formed_by_BSc_only,
		293  'Position of pleuro-axillary joint' / fulcrum_articulates_with_2Ax fulcrum_articulates_with_1Ax_and_2Ax fulcrum_articulates_with_1Ax 'fulcrum-2Ax joint is replaced by a joint formed by basalare (Ba) and',
		294  'Length of metathoracic pleural wing process (PWP)' / 'ends on same level with basalare (Ba)' distinctly_overtops_Ba,
		295  Size_of_externally_visible_metathoracic_subalare / between_one_sixth_and_one_third_of_length_of_notum size_exceeds_one_third_of_notal_length less_than_one_sixth_of_notal_length,
		296  Shape_of_externally_visible_metathoracic_subalare / longer_than_high about_as_long_as_high,
		297  Process_of_dorsal_margin_of_externally_visible_metathoracic_subalare / absent present,
		298  Field_of_sensilla_trichodea_in_membrane_between_subalare_and_dorsal_epimeral_margin / absent present,
		299  Close_association_between_metapostnotum_and_tergum_I / absent present,
		300  Fusion_of_abdominal_tergites_and_sternites_IX / not_fused partly_fused 'fused and forming a ring-like structure',
		301  Tergum_VIII / 'ventral margins normal, not enlarged' 'saddle-shaped, ventral margins enlarged' connected_with_ventral_sclerites_by_slender_sclerite_band fused_with_ventral_sclerites,
		302  'Ventral sclerites of segment VIII (gonocoxae and gonapophyses)' / distinct not_distinct,
		303  Connection_of_gonocoxae_VIII_of_right_and_left_body_half / absent fused_ventromedially,
		304  Connection_of_gonapophyses_VIII_of_right_and_left_body_half / separate fused,
		305  Length_of_ventral_sclerites_of_segment_VIII / not_distinctly_longer_than_tergum_VIII 'gonapophyses elongate, distinctly longer than tergum VIII' 'entire sclerotisation strongly prolonged caudally (gonocoxae and g',
		306  'Degree of fusion of terminal segments (IX - XI)' / clearly_separated terminal_segments_partially_fused 'completely fused, segmental margins not traceable',
		307  Sclerotisation_of_tergum_IX / evenly_sclerotised ventral_margins_of_sclerotised_area_enlarged all_parts_weakly_sclerotised 'sclerotisation not continuous, band-like' fusion_with_ventral_sclerites,
		308  Length_of_tergum_IX / about_as_long_as_preceding_tergites about_half_as_long_as_preceding_tergites,
		309  Ventral_sclerites_of_segment_IX / absent present,
		310  '310. Gonocoxae, gonapophyses and gonoplacs IX' / all_elements_well_developed gonapophyses_reduced gonapophyses_and_gonoplacs_reduced,
		311  Fusion_of_gonocoxae_IX_of_right_and_left_body_half / absent fused_ventromedially fused_dorsomedially,
		312  Fusion_of_gonapophyses_IX_of_right_and_left_body_half / absent basally_fused completely_fused,
		313  Lenght_of_ventral_sclerites_of_segment_IX / not_markedly_longer_than_tergum_IX only_gonocoxae_caudally_prolonged gonapophyses_and_gonoplacs_caudally_prolonged only_gonoplacs_caudally_prolonged,
		314  Median_closure_of_sclerotisation_of_tergum_X / absent present_with_tergite_X_distinctly_sclerotised present_with_tergite_X_weakly_sclerotised,
		315  Fusion_of_tergum_X_with_ventral_sclerotised_elements_of_segment_X / absent fused_to_form_sclerotized_ring,
		316  Cercus / absent_or_vestigial present socii,
		317  Retractability_of_postabdominal_segments / not_or_only_very_slightly_retractable 'segments posteriorly tapering, not fused, telescoping (tube-like)' 'segments IX + X (XI) fused, retractable into segment VIII' 'segment IX and gonocoxae VIII retractible (between tgVIII and stVII)',
		318  Anterior_apophyses_VIII / absent one_pair_on_gonocoxae_VIII two_pairs_on_tergum_VIII_and_gonocoxae_VIII one_pair_on_a_composite_formation_of_tergum_VIII_and_gonocoxae_V,
		319  Posterior_apophyses_IX / absent one_pair_originating_from_a_ventral_area one_pair_originating_from_a_dorsal_area,
		320  Extrusion_of_terminal_segments / by_hemolymph_pressure 'segment VIII and terminal unit (segments IX - XI) extruded by musc' gonocoxae_VIII_and_segment_IX_extruded_by_muscle_force,
		321  Spermathecal_process_of_genital_chamber / absent 'present, sclerotised' 'present, unsclerotised',
		322  Spermathecal_duct / absent present,
		323  Mesocuticle_in_the_spermathecal_duct / absent present,
		324  Muscularis_of_spermathecal_duct / absent mainly_longitudinal_fibres mainly_circular_fibres longitudinal_and_circular_fibres_mixed,
		325  Spermathecal_gland / absent present,
		326  Bursa_copulatrix / absent present,
		327  Size_of_bursa_copulatrix / moderate 'enlarged, balloon-like',
		328  'Muscle 06 (isVII-VIII06)' / absent present,
		329  'Muscle 08 (isVII-VIII08)' / absent present,
		330  'Muscle 11 (isVII-VIII11)' / absent present,
		331  'Muscle 12 (isVII-VIII12)' / absent present,
		332  'Muscle 24 (isVIII-IX11)' / absent present,
		333  'Muscle 29 (isX-XI02)' / absent present,
		334  'Muscle 34 (intraIX03)' / absent present,
		335  'Muscle 35 (intraIX04)' / absent present,
		336  'Muscle 48 (tVIII01)' / absent present,
		337  'Muscle 55 (seVIII-gc01)' / absent present,
		338  'Muscle 56 (seVIII-gc02)' / absent present,
		339  'Muscle 59 (seIX-gc02)' / absent present,
		340  Trichobothria_field_on_tergum_X / absent present,
		341  Shape_of_the_trichobothria_field_on_tergum_X / dense_circular_spot 'dense, band-like' loosely_arranged_trichobothria,
		342  Elongation_of_tergite_IX / absent present,
		343  Genitalic_capsule_of_males / absent 'present, basistyli not fused' 'present, basistyli fused',
		344  Stylar_organ_on_dististylus / absent present,
		345  Sperm_pump_with_genital_folds_enclosing_the_pumping_chamber / absent with_movable_chamber_and_longitudinal_tegimen_formed_by_roof_of_ with_movable_chamber_and_pistil_formed_by_median_lobe_of_fulcrum with_movable_pistil_formed_by_roof_of_endophallus,
		346  Aedeagus_apodeme_with_median_longitudinal_lamella / absent well_developed 'vestigial, serving as lever of the pistill levator',
		347  Sperm_pump_formed_by_modified_endophallus / absent present,
		348  Accumulated_trichobothria_on_anal_segment / absent 'present, forming a band' 'present, forming a rosette',
		349  'Acanthae of proventriculus close-set, prominently elongated' / absent present,
		350  Number_of_Malpighian_tubules / more_than_20 8_or_less,
		351  Ovarioles / panoistic meroistic,
		352  Telotrophic_ovarioles_of_specialised_type / absent present,
		353  Appearance_of_fully_developed_compound_eyes_including_external_apparatus / 'before penultimate (pupal) stage' at_pupal_stage,
		354  External_wing_buds_in_immature_stages / absent present,
		355  Male_sex_chromosomes / not_homogametic homogametic,
		356  'Haplo-diploidy' / absent present,
		;

MATRIX
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[                             .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .         .   ]
Agathiphagidae       10004?011-001010100115-04110000100121010003101010000000000110000000010002000101010000000000110000110?10??0000001100000021121111110?-01100111110111110001?0110110111?10110?00011001010{03}11101101101111110101010100---1101111111101110110-111100?1011000-001011202-10?1010211000001100110-3000010?1000101101100011-02--0------0222101121111111000001100-000000001101010
Bibionidae           10004?000-000010?11115-04100001010121010002100--00000000000-00000000110111000--0-1001--011000020011111011000100{12}{01}0012012000001101010000111010001{02}10-001000011100011000111-01010000-01-0100000001001000-011?110-1??2100?100-0001000-010-001-0000011213?0-2-00101-?1100111-0--020----0---30---00?110-03-000000011-0-011?1-0?0?1000010100-0000000011000-000001001??1000
Bittacidae           0000000111101011?00??400??00001111021010??0101110010000?00100010010011002100001010000100010000200111110211011001000010020010011010101001010110010111001000111100011010110000010000-10000--020101001010-0111100-10111111100100010011100-111100010012020001101001111210111312320111100111001021001000101001001010-00010----0010000010100-0000010011010-22?220011101000
Boreus               0000200101100011000005-0210000010012101000010111100000000010101001001100210000101011010001100120011101021101101100001002001000-0-0110--1010110010?0-0010001111000110001?0-32----01110{23}0100?20100000010---0------------00-01100-00--000---0-000000---------------------------------------------------------0{01}111-20000----0010000010?00-1000000010000-111000011001000
Caeciliusidae        00000011100000010000100010100000000?0001110101001000000000--00000000100000100000000000100000002000?0000100000002000100020120011110100100-000-000-01100100000001000-01000--000100010112110101011110111101010110-0---00000-1110011110100-00010000001210-0100011011111121023022020200101203010?0010-0-00101000000001001100020000000010?00-0000000000000-000000001100100
Caurinus             00002?01101??0???00??5-0??000001?????0????2?01111000000?0010100001001100212000001011010001000120011101001101110100001002001000-0-011---1010??001-?0-0010--1?110001?0001?0-32----01110{23}0101?20100000010---0------------00-01100-00--000---0-000000---------------------------------------------------------00111-00211?0000-11000010100-1000000000000-11?0000????10-?
Chrysopidae          10002100--000000010111010100110000111111000111000000000100101000000010000010001000000000000000000111000000000012100000020110000110?-010001010101-11000?010110100010?10110-00000001010-010100?111111111010110-0-0---110110101?111111010-111000?1001210-001000011001????02010?12111100020300020013-01221100000111-0001120-00000000010000-00000000000010000000201101000
Corydalidae          111021001-0101000000121011000000000000010002010000010030201011000000100000100010000000000000000001110000000000{01}1{12}0100112011000011011010001010101-1110010{01}0110100010?10110-000000?1010{23}01002{01}{01}11?0111110{01}{01}110-101100110110001{01}111111011011100010101200-0101010011?1211112112312021001110000021011100111100000011-0000110-00000000010?00-????????????12000000201001000
Culicidae            10014?000-00?11000{12}135-0310000100012101000?100--00000000200-00010000110111000220-1001-10?1000020011111011010100{12}10012012000001101010000111010001010-001000011100011000111-010100-0-01-00--000000001000-011?110-11121000100-0000000-010-000-000001122??0?000000110020??02010?0--------??-0001001110-0??-??000011-0-011?1-0-001000010?00-0000000001000-000000001101000
Cupedidae            1000-?010000001-0001130011100000000110000010110100000000000011000000100021210010000000000000000001??0000000-0100--01211001200220010-0000-100-111-01111000011010-00-011110-10110001000310--0001100011110--0-0-0-11001001101111011110001011010000200200-020003?01-0120110200010100100302000100011101310110000001?-0-201??002-03112010?00-000000000?0?0-000000001101000
Diprionidae          000000001-1000010000140001000001000100010031010101000000000?0000100120002?000010100000000000000200???11?00000001120020030112011110100000-1010101-00-00000011010101000010--00010101010301011101001?10010??1?0-1??---1001101-11?101??010-111101010012111010001000101201?12210214211011-123001200?2-0?10????010100012--10012--2000000--00-0000000000000-0000000?0??1001
Eriocraniidae        10004?010-00?0??100??5-0??00000100121010??3?010100000000001100000000100020001000100000000?0110000111010000000001100000021121111110?-01100111100111110001?0110110111?10110-000110010?03?11001011?11101101010100-0---1101111110100110100-111100?10?1200-010111202-10?111020000120110021103000200?1000101011100211-02--?------0232101121111001000001100-000000001??1010
Helophoridae         110020000-0201000101040011000000000200010000010100000100000011000000100020210000000000000000000001110000000-01010011011000200220010-0000-100-011-011110000?0010-00-00110--10110001000-00--0001000001010--0-0-0-0---1000100110011110000-11010000000??0-1?-?0??01-01--1102010202010012121011100111010121000000010-0-401?0000003112010?00-000000000?0?0-000000001101000
Inocelliidae         11102?001-0101???00??300??000010000000????0?01000000000?001?110000001000??000010000000000?00000001???????0??000?2010012???00000110?-000?01?10?01-11000?010110100010210110-000000????????????????????????????????????????????????????????????010101000-00100200000110111201031101110102100100?10110011010000010111010112-10000000010200-0000000000001100000010111100?
Limnephilidae        000030001-00101000001---4-00001100121010003101010000002020100000000011002?0000111000000000?0000001???10000000001100000020120111110110100011111011111001000110110011010110-0001001101030110110111111011110100-100---11011111011101101{01}0-11111001001??0-0???0??????0????12?1??0???????????0???00???????????100011-01310----101200?11021100000000001000-000000001101010
Mengenillidae        100020000-0000--01010---4-0000-000121000000001-2100010000110010000100-102103001000?01--00-0000-001---0001-0-0000--010010000002200011-000-100-001-00-00100110010000-00010--200100010?0-00--1001000010010--0-0-0-0---00010-0-0001010-000-1011-00000?120-1---0------10?3-?22222020121201?-3????00-2-031?1---000--------0------00--000--00-0000000000000-000000001--0100
Micropterigidae      10002?011-0010100000140021{01}0000100021010??3101010000000000100000000010002000100010000000000110200000010000000001100000021121111110?-01100111100111110001?0110110011?10110-0001100101031110010110111010-1010100-0---1101111110100110100-111100?1011000-001011202-102101021020030110010113010200?1000101101100011-00--0----?-0100021021111000100000000-000000001101010
Nannochoristidae     1001100110102010002024003101000100021010000101010000001020?-000102001101210001-011001-1111000020111111011010100100001002011001101010000101011001011100100011110001101011000001000111010101020111001010-0111101011111101100110010011110-111110010010020010001001001210111310321011100010301001001100101001002010-00001?0-00111000010100-1000010011010-021110011001000
Nevrorthidae         11102?00--01010001010100010011000011110100001100000000002010100000001000??100010000000000?000?0001??0000?00?00?20000000201100001100-010001010101-11000?0?0?10100010110110-00000001010-010100111111111101?110-1011001101101011111111010-11100011001200-01100?011001????02312213111100022300020013?0122110000010110011120-00000000010000-000000000000120000002?1??100?
Ommatidae            10002?000-0000???00??300??100000000??0????1?01000000000?000?11000000100021210010000000000000000001110000000-010?--{01}1211001200220010-0000-100-111-01111000011010-00-011110-10110001000310--0001000011110--0-0-0-11001001101111011110000-11010000200200-020003?01-01201102000101001003020001000111013101100000????????????0?-0?????????????????????????000000001??100?
Osmylidae            10002100--000000010111010100110000111111000111000000000110101000000010000?100010000000000000000001110000000?0012000000020110000110?-010001010101-11000?010110100010110110-00000001010-0101001111111111011110-100---1101101011111111000-11100011001200-01100?011001????02312213111100022300020013?01221100000111-0011110-11000000010?00-00000000000010000000201101000
Panorpidae           00000001111010110000040031100011110210100001011100100000001000100100110021000010100001000110002001110102110110110000100200100110101110010101100101110010001111000110101100000100011102010{01}020100011010-01111010111111111000100100111111111000010010120000101000001110111310322021200111000001001002101001002011-00001?1-00111000010?00-1000010000010-221220011101000
Pteronarcyidae       100000100-00000100001000001?000000000001110101000000100020--11000000100000200000000000000000000000000000?0000001000000?10120002110110100-000-1000010001000?1000000-01000--00000001011201010111111?111100011111??0??11011001?1?1111010??11110001001200-010001?01111211112?1220200101210-1010?00?0-0-0010000000----0000----0010000010?00-0000010000000-000000000000100
Pulicidae            1000--001-010110001025-011000010001210??002100--00000000000-10000101110121020--011001-1101000010111111011010100100003112000000-0-0-----001011001-00-0000--00010-01100010---------1000-00--120100000010---0------------00-0-000-00--000---0-000000---------------------------------------------------------00011-0--0?----1000000010?00-0000010001010-000310011001000
Raphidiidae          111021001-0101000000130001000010000000010001010000000000001011000000100000100010000000000000000001110000000?000?201001220100000110?-000001010101-11000?010110100010210110-00000001010310--210111011111010110-1011001101101011111111011011100010101000-00100200000110111201031101110102100100101110011010000010111010112-10000000010?00-00000000000011000000101111000
Rhyacophilidae       110030001-00101-00101---4-0000110012101000310101000000202010000000001100200000111000000000000000011101000000000110000002012011111011010001111?011111001000110110011010110-0001001101030111100111111011110100-100---1101110100110110100-1111?001001200-01100100111021011221200101100211-301020011010101101100311-01310----101--2111021100110100001000-000000001101010
Sialidae             110021011000001000001310210000000000000100020100000100002010110000001000002000100000000000000000011100000000000110000112011000011011010001010101-110001010?10100010110110-000000?101031100{02}10111111111001110-1011001101101011111111011011100010101000-000001001001?11110010210?111000023000200?110011110000010110000110-00000000010?00-00000000000012000000201111000
Tenthredinidae       000000001-10000100001{23}00310000010001000000310101010000000000000010012000{02}0010010100000000000000200000110000000{01}1120020030112011110100000-1010101-00-00000011010101000010--00010101010301011101001110010--100-110---1001101-1{01}110110010-111101010012111010001000101201?122102142110-1-??300120012-0?10????010100012--10012--2000000--00-0000001100000-000000000101001
Tettigoniidae        00000011000000000000100020-0000000000001110101001000100000--10000000100000100010000000000000000000000000000000010011001101200021100-0101-000-100-01100000010100000-01000--00000001011-0101011111101110-100-0-?-0---000110001100100-0-0-01100000001210-020?01101111111012212201?0001210-201020014-0-0010?000010001010110230010000010?00-0000000000000-000000000000100
Tipulidae            10004?000-0000--0{01}1115-0410{01}0010{01}012100000-100--00000000{012}00-00200100010111000--0-1001--011200020011111011010100210012002000001101010000111010001110-001000011100011000111-010100-0-01000--300001001000-011?11101112100?100-0001001?010-001?10010012?31001100101011100111201202001?1000-20120001110-03-000000011-20011?1-00010000010300-1000010011000-000001001101000
Trachypachidae       100020000-020100010103001100001000-200010000010000000100000011000000100021210010000000000000000001110000000-0100--?1?11000230220010-0000-100-011-011110000?0010-00-00110--101100?1000-00--0001000001110--0-0-0-0---1?00??0??0011110000-11010000200??0-0?--0??01-01--110200010200100300-011000111012121000000010-0-401?0000003112010?00-00000011010?0-000000001101000
Xenidae              100020000-0000--01010---4-0000-000121000000001-2100010000110000000100-102103001000?01--00-0000-001---0001-0-00{01}0--010010000002200011-000-100-001-00-00000110010000-00010--200100010?0-?0--1001000010010--0-0-0-0---00010-0-0001010-000-1011-?0010?--------0------10?3-1?3?2?0402012010--????00???????????000--------0------00--000--00-0000000000000-000000001--0100
Xyelidae             00000?0110100001000012000100000100011001003101010100000000100000100120002000000010000000000000010000011000000001110000030112011110100000-1010101-00-00000011110101001010--0001010101030101110110111111010100-110---1?11101111111110010-11110101001222101000100100120111211031421-2-0-??300020012-02100210010100012--10012--2000000--00-0000001100000-000000000??1001
Zorotypidae          000000101-000001000010001010000000000001110101000000100000--00000000100000100000000000000000000000000001000000120010000101200011100-0100-000-100-0100000--?0000000-01000--00010001011{02}10--01111100{01}1110110-100-10?0000{01}1001110{01}110-101000110000001210-0???01101111?1101202230401112210-0012000120000010100000----0010----0010000010000-0000000000000-000000001000100
;
END;

[ The following blocks are output data for analysis step 8002 ]
BEGIN TREES;
      TITLE CLA_338_sm_HolometAppIV;
      LINK TAXA = TaxaForAnalysisStep8002;
      TREE Fig._1 = [&R] (Pteronarcyidae,Tettigoniidae,(Zorotypidae,(Caeciliusidae,((Xyelidae,(Tenthredinidae,Diprionidae)),(((Chrysopidae,(Nevrorthidae,Osmylidae)),(Sialidae,(Corydalidae,(Raphidiidae,Inocelliidae)))),(((Rhyacophilidae,Limnephilidae),(Micropterigidae,(Agathiphagidae,Eriocraniidae))),(((Mengenillidae,Xenidae),((Ommatidae,Cupedidae),(Helophoridae,Trachypachidae))),((Pulicidae,(Culicidae,(Tipulidae,Bibionidae))),((Boreus,Caurinus),(Nannochoristidae,(Panorpidae,Bittacidae)))))))))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr48912]


END;