#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on November 11, 2026; 15:52 GMT TreeBASE (cc) 1994-2008 Study reference: Page R., Price R., & Hellenthal R. 1995. Phylogeny of Geomydoecus and Thomomydoecus pocket gopher lice (Phthiraptera:Trichodectidae) inferred from cladistic analysis of adult and first instar morphology. Systematic Entomology, 20(2): 129-143. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S573] [ The following blocks are input data for analysis step 824 ] BEGIN TAXA; TITLE TaxaForAnalysisStep824; DIMENSIONS NTAX=122; TAXLABELS Geomydoecus_actuosi Geomydoecus_albati Geomydoecus_alcorni Geomydoecus_alleni Geomydoecus_angularis Geomydoecus_aurei_aurei Geomydoecus_aurei_grahamensis Geomydoecus_bajaiensis Geomydoecus_betleyae Geomydoecus_biagiae Geomydoecus_bulleri_bulleri Geomydoecus_bulleri_intermedius Geomydoecus_bulleri_melanuri Geomydoecus_burti Geomydoecus_californicus Geomydoecus_centralis Geomydoecus_chapini Geomydoecus_cherrei Geomydoecus_chiapensis Geomydoecus_chihuahuae_chihuahuae Geomydoecus_chihuahuae_emersoni Geomydoecus_clausonae Geomydoecus_cliftoni Geomydoecus_copei Geomydoecus_coronadoi_coronadoi Geomydoecus_coronadoi_saccharalis Geomydoecus_coronadoi_submerriami Geomydoecus_costaricensis Geomydoecus_craigi Geomydoecus_crovelloi Geomydoecus_dakotensis Geomydoecus_dalgleishi Geomydoecus_davidhafneri Geomydoecus_duchesnensis Geomydoecus_ewingi Geomydoecus_expansus Geomydoecus_extimi Geomydoecus_fuchsi Geomydoecus_fulvescens Geomydoecus_fulvi Geomydoecus_geomydis Geomydoecus_guadalupensis Geomydoecus_heaneyi Geomydoecus_hoffmani Geomydoecus_hueyi Geomydoecus_idahoensis Geomydoecus_illinoensis Geomydoecus_jaliscoensis Geomydoecus_jonesi Geomydoecus_limitaris_bakeri Geomydoecus_limitaris_halli Geomydoecus_limitaris_limitaris Geomydoecus_limitaris_tolteci Geomydoecus_martini Geomydoecus_mcgregori Geomydoecus_merriami Geomydoecus_mexicanus Geomydoecus_mobilensis Geomydoecus_musculi Geomydoecus_nadleri Geomydoecus_nayaritensis Geomydoecus_nebrathkensis Geomydoecus_neotruncatus Geomydoecus_oklahomensis Geomydoecus_oregonus Geomydoecus_panamensis_dariensi Geomydoecus_panamensis_panamensis Geomydoecus_pattoni Geomydoecus_perotensis_irolonis Geomydoecus_perotensis_perotensis Geomydoecus_polydentatus_angustirostris Geomydoecus_polydentatus_polydentatus Geomydoecus_pygacanthi Geomydoecus_quadridentatus Geomydoecus_scleritus Geomydoecus_setzeri Geomydoecus_shastensis Geomydoecus_sinolae Geomydoecus_spickai Geomydoecus_subcalifornicus Geomydoecus_subgeomydis Geomydoecus_subnubili Geomydoecus_tamaulipensis Geomydoecus_telli Geomydoecus_texanus_texanus Geomydoecus_texanus_tropicalis Geomydoecus_thaeleri Geomydoecus_thomomyus Geomydoecus_tolucae Geomydoecus_traubi Geomydoecus_trichopi Geomydoecus_truncatus Geomydoecus_umbrini Geomydoecus_ustulati_clarkii Geomydoecus_ustulati_ustulati Geomydoecus_veracruzensis Geomydoecus_warmanae Geomydoecus_welleri_multilineatus Geomydoecus_welleri_welleri Geomydoecus_wernecki_planiceps Geomydoecus_wernecki_wernecki Geomydoecus_yucatanensis Thomomydoecus_arleneae Thomomydoecus_asymmetricus Thomomydoecus_barbarae Thomomydoecus_birneyi Thomomydoecus_byersi Thomomydoecus_dickermani Thomomydoecus_genowaysi Thomomydoecus_greeri Thomomydoecus_jamesbeeri Thomomydoecus_johnhafneri Thomomydoecus_markhafneri Thomomydoecus_minor Thomomydoecus_neocopei Thomomydoecus_orizabae Thomomydoecus_peregrini Thomomydoecus_potteri Thomomydoecus_timmi Thomomydoecus_wardi Thomomydoecus_williamsi Thomomydoecus_zacatecae ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2195] TITLE ; LINK TAXA = TaxaForAnalysisStep824; DIMENSIONS NCHAR=58; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7 8 9" MISSING=? GAP= -; CHARSTATELABELS 1 Lateral_seta_VI / 0 present absent, 2 Marginal_terminal_setae / 0 none one_v._long two_short, 3 Lateral_metanotal_seta / 0 'very long, > 0.2 mm' 'short, < 0.09 mm', 4 Temple_sculpturing / 0 none many_dark_bumps few_dark_bumps sparse_bumps_nr_margin small_bumps_in_rows, 5 SMT_4 / 0 present absent, 6 Int._sternal_setae / 0 '<=1' '>=2', 7 Seta_on_coxa_III / 0 'large, thick, > 0.020' 'medium, 0.010-0.020' 'short, <= 0.010' 'slender, >= 0.025', 8 MT_seta_4 / 0 prominent small, 9 'Med. tergal setae I-VI' / 0 'all med-long' all_short 'ant. long, post v. short' all_v._short, 10 Med._tergal_seta_VII / 0 absent present, 11 Med._tergal_seta_VII / 0 absent present, 12 Med._metanotal_seta / 0 'long, > 0.2 mm' 'short, < 0.11 mm', 13 Int._sternal_setae_II / absent present, 14 'Med length sternal setae IV-V' / 0 '5-15' '17-25' '27-35' '37-45', 15 Relative_size_med_sternal_setae / 0 '= int. setae' larger_than_int._setae, 16 Med_sternal_setae_on_IX / 0 absent present, 17 'Relative size sternal setae II-' / 0 similar VII_shorter 'VI-VII shorter' 'II, II-III, II-IV shorter' V_much_longer, 18 Submarginal_seta_on_VII / 0 long medium 'short-minute', 19 Mean_temple_width / 0 '240-262' '264-310' '311-340' '344-360' '381-395' '433-482', 20 Med._length_head_seta_1 / 0 '5-25' '27-50' '52-75' '77-140', 21 Med_length_head_seta_2 / 0 '5-10' '12-20' '22-32' '52-62', 22 Med._length_pronotal_seta / 0 '7-25' '27-50' '52-77' '82-112', 23 Marginal_temple_seta_5 / 0 'short-minute, < 10' '15-20 long', 24 Length_submarg._head_seta_3 / 0 '10-22' '25-40' '42-62', 25 Length_dorsal_ant._pedical_seta / 0 '27-45' '47-65' '67-85' '87-112', 26 Lat_seta_IV / 0 short long, 27 Rel._length_submarg._VI_vs_VII / 0 comparable much_shorter, 28 'Submarg setae II-V' / 0 all_long all_short all_very_long 'II-III or II-IV very long, rest', 29 Int._tergal_setae / 0 long medium short, 30 Very_long_lat._seta_VII / absent present, 31 Length_int._sternal_seta_VII / 0 '5-17' '20-37' '42-57' '67-120', 32 Int._sternal_setae_III / absent present, 33 Lateral_seta_V / short long, 34 'LNS TG8 / HD WDTH' / '< 0.09' '0.09-0.20' '0.20-0.27' '>0.27', 35 NGS_LPS / none '1-2' '3-5' '6-9' '10-11' '12+', 36 'LNS TG7 / HD WDTH' / '<= 0.18' '0.18-0.25' '0.25-0.35' '0.35+', 37 'PSLTG IN / HD WDTH' / '<= 0.07' '0.07-0.15' '0.15-0.225' '0.225-0.27' '0.27+', 38 'PSLTG OT / HD WDTH' / '<= 0.10' '0.10-0.23' '0.23+', 39 SET_STN2 / '<= 7' '7-9.5' '9.5-13' '13+', 40 'GS LGTH / TOT LGTH' / 0 '<= 0.09' '0.09-0.12' '0.12+', 41 Last_tergite_setal_formula / '2+2+2' '3+3' '1+4+1', 42 Ant._margin_genital_sac / smooth prominent_central_notch ant._protrusion, 43 'male MG TMP / fem MG TMP' / '< 0.7' '0.7-1.15' '> 1.15', 44 SMG_TMP_position / lateral medial, 45 Male_scape_margin / straight weak_convexity distinct_protuberance, 46 'MG TMP/ HD WDTH' / '<= 0.035' '0.035-0.076' '>0.076', 47 'SMG TMP / HD WDTH' / '< 0.09' '0.09-0.15' '0.15-0.21' '0.21-0.28' '> 0.28', 48 'HD LGTH/TOT LGTH' / '< 0.25' '> 0.25', 49 'Male scape length (SCP LGTH/HD' / '< 0.31' '0.31-0.45' '>0.45', 50 SET_TG7 / '<=13.5' '>13.5', 51 SET_STN8 / '<5' '5-7' '>7', 52 No._genital_sac_spines / none two three four five six eight_or_more, 53 'GPA WDTH / HD WDTH' / '<0.19' '0.19-0.24' '0.24-0.31' '>0.31', 54 Grouped_setae_on_II_and_III / absent present, 55 Genital_parameral_arch_shape / semicircular medial_post._apical_proj. marked_medial_thickening post._margin_truncated lateral_shoulders triagular_apical_proj. med._apical_proj._bulbous long_pical_proj. irregular no_apical_proj., 56 Genital_endomeral_plate_shape / ovoid diamond elongated_diamond bluntly_triangular irregular triangular deep_terminal_bifurcation rounded_post_margin elongated_triangle long, 57 Genital_spine_shape / stout 'slender, curved' 'small, triangular', 58 'Outer mid-dorsal head seta' / short long, ; MATRIX [ 10 20 30 40 50 ] [ . . . . . ] Geomydoecus_actuosi 1122121112111221122233111122311111312133100021301115305501 Geomydoecus_albati 1122121111111411112333121111113111312133100021301115305501 Geomydoecus_alcorni 112111211211031111221?223114112111412123001001201115305300 Geomydoecus_alleni 1121113221110311116431134114214010012103101100011113305700 Geomydoecus_angularis 1122121111111411212333122111113111312133100021301115305501 Geomydoecus_aurei_aurei 1122121112111221122222111122312111212133100021301115305501 Geomydoecus_aurei_grahamensis 1122121112111221122232112022311101212133100021301115305501 Geomydoecus_bajaiensis 1122121111111411212344122111113111212133100021301115305501 Geomydoecus_betleyae 212121311222012143211111202232100201322100?0?????????????0 Geomydoecus_biagiae 212121311222012243211112102232100201322100?0?????????????0 Geomydoecus_bulleri_bulleri 2122112121111121132122222022311101512123001001201115305300 Geomydoecus_bulleri_intermedius 2122112121111121132212224022311101512113001001201115305300 Geomydoecus_bulleri_melanuri 2122112121111111133111211022311101511123001001201115305300 Geomydoecus_burti 2122112141111111133222234022311101512113001001201115305300 Geomydoecus_californicus 1122121111111411112444132111114111312133100021301115305501 Geomydoecus_centralis 1122121111111411112333131111113111312133100021301115305501 Geomydoecus_chapini 11211132422103111352211240223120001331021021000101100078?0 Geomydoecus_cherrei 1121113242110211133111114022312000012113112101011112305600 Geomydoecus_chiapensis 2121113241110311116421124112213011112103102101?11115305500 Geomydoecus_chihuahuae_chihuahuae 1121113112211122432111133022311103034123101002301115302500 Geomydoecus_chihuahuae_emersoni 1121113112211122432112134022311103034123101002201115305500 Geomydoecus_clausonae 1122121111111411212344121111112111212133100021301115305501 Geomydoecus_cliftoni 1122121112111221122222111122311111412123100021101115305501 Geomydoecus_copei 11211132422103111141211240123120010032021011000110000178?0 Geomydoecus_coronadoi_coronadoi 1123111141111211232221112022311101210023100001102115305501 Geomydoecus_coronadoi_saccharalis 1123111141111211132232114022312101210023100001102115305501 Geomydoecus_coronadoi_submerriami 1123111141111211432232113022312101210023100001102115305501 Geomydoecus_costaricensis 1121113242110211134121114022312000012113112101011113205600 Geomydoecus_craigi 2121213112220122432111121022321002014221001000401113309510 Geomydoecus_crovelloi 1122121112111321112223111111311111212133100021301115305501 Geomydoecus_dakotensis 2121213112220122432111121022321002013222001000401125309510 Geomydoecus_dalgleishi 1124111112111121132232121011311101012123100021111115305501 Geomydoecus_davidhafneri ?????????????????????????????????0012113112101011115305500 Geomydoecus_duchesnensis 21212131122201224321111110223210020132210010004111103095?0 Geomydoecus_ewingi 1122111122111111332222111022311101212123100011101113305501 Geomydoecus_expansus 1123112141111321112322123014312111212123101001101115305500 Geomydoecus_extimi 1122121112111411113433112113112111312123100021101115305501 Geomydoecus_fuchsi 2121213112220122432111121022321002013221001000401115309510 Geomydoecus_fulvescens 11211121121102111232232341141121110011110010012111201041?0 Geomydoecus_fulvi 1121113112210122532121134022311002023123100002401115302500 Geomydoecus_geomydis 1124111112111121132332122022311101412123100011101115305501 Geomydoecus_guadalupensis 1122121111111221112223111122311111312133100021301113305501 Geomydoecus_heaneyi 1124111112111121122332131011311101112123100021101115305501 Geomydoecus_hoffmani 11211132422103111351211240223120001331021021000101100078?0 Geomydoecus_hueyi 1125124111111312212334122111212111412133100001301115305501 Geomydoecus_idahoensis 1121112112111122132232123121311101312133100001301115305501 Geomydoecus_illinoensis 1121111112111121122322121012311101212123100001101115305501 Geomydoecus_jaliscoensis 1122121112111211113333111121212111412123100021101115305501 Geomydoecus_jonesi 112111322111031111642113?114214010012103101100011123305700 Geomydoecus_limitaris_bakeri 1122121112211221122222111022311101212133100021301115305501 Geomydoecus_limitaris_halli 1122121112111221122223111122312111212133100021301115305501 Geomydoecus_limitaris_limitaris 1122121112111221122223111122311111312133100021301115305501 Geomydoecus_limitaris_tolteci 1122121112111221122223111122312111412133100021301115305501 Geomydoecus_martini 1123112141111321112222124114312111312123100001101115305500 Geomydoecus_mcgregori 1121114121110311113122234114112111112123001001201124305300 Geomydoecus_merriami 1123112141111121132221122022311101511023100001102115305501 Geomydoecus_mexicanus 112111321221041112221?2241111131110022011010012111202000?0 Geomydoecus_mobilensis 112111314211111133222113402231110131211210?0?????????????0 Geomydoecus_musculi 1122121112111211112232111121211111212123100021101115305501 Geomydoecus_nadleri 2122112121111121132121223022311101512113001001101115305300 Geomydoecus_nayaritensis 1122121111111311113333111114112111412123100021101115305501 Geomydoecus_nebrathkensis 1124111112111122122222111022311101112123100011101115305501 Geomydoecus_neotruncatus 1121111132111111332121113022311101112122100001011115305500 Geomydoecus_oklahomensis 1124111112111121132332121022311101112123100021111115305501 Geomydoecus_oregonus 1121112112111122123232123121311101312123100001201115305501 Geomydoecus_panamensis_dariensi 1121113242110211135221124022312000012113112101011115305500 Geomydoecus_panamensis_panamensis 2121113242110211134221134022311000012113112101011115305500 Geomydoecus_pattoni 112111311221012243222?134022311002023113101002301115306500 Geomydoecus_perotensis_irolonis 11211121121102111321122241211121110011110010011111201000?0 Geomydoecus_perotensis_perotensis 11211121122104111122222240211131110011110010011111201000?0 Geomydoecus_polydentatus_angustirostris 1123112141111221432122123022311101211023100001102116305501 Geomydoecus_polydentatus_polydentatus 1123112141111121432121111022311101211023100001102116305501 Geomydoecus_pygacanthi 2121113241110411116431124112213011102103102101011115305500 Geomydoecus_quadridentatus 112212111211131121223311101231210111212310002110111330550? Geomydoecus_scleritus 1121113142111111332221134022311101322113101001111115305500 Geomydoecus_setzeri 1121113242110211133111114022311000012113112101011115205500 Geomydoecus_shastensis 1125124112111322112323133111112111312133100001301115305501 Geomydoecus_sinolae 1122121112111311122223111122312111412133100021301115305501 Geomydoecus_spickai 1124111112111121132332122022311101112123100021111115305501 Geomydoecus_subcalifornicus 1122121112111411112334111111212111512133100021301113305501 Geomydoecus_subgeomydis 1122111122111111332222111022311101212123100011101115305501 Geomydoecus_subnubili 1211113121110211112112134012212000102113101001001103305500 Geomydoecus_tamaulipensis 1111113121110311113124134012213001123113101001001113305500 Geomydoecus_telli 1123111141111222433222122022312101011023120001102115305501 Geomydoecus_texanus_texanus 1111113121110111133121124022311101112123101001001113305500 Geomydoecus_texanus_tropicalis 1111113121110111232111124022311101112123101001001113305500 Geomydoecus_thaeleri 2121213112220121432111111022321002013221001000401115309510 Geomydoecus_thomomyus 2121213112220122432111111022321002013221001000401113309510 Geomydoecus_tolucae 1121113112211122132121134022311102013133100001301115302500 Geomydoecus_traubi 1121113212110311123112224122212110001201001001211121201220 Geomydoecus_trichopi 2221113112210121432111124022311101211123001001201115305300 Geomydoecus_truncatus 1121111132111111332121124022311101112122100001011115303700 Geomydoecus_umbrini 1122121111111211112333111121212111312123100021101115305501 Geomydoecus_ustulati_clarkii 1111113121110211112122134012212001112113101001001113305500 Geomydoecus_ustulati_ustulati 1111113121110211113112134012212001112113101001001113305500 Geomydoecus_veracruzensis 1123111141111311422121111111112111311023100001101115305501 Geomydoecus_warmanae 1122121112111121112123111111311111312133100021301115305501 Geomydoecus_welleri_multilineatus 1122121112111321112123111121112111313133100021101115305501 Geomydoecus_welleri_welleri 1122121112111311122122111121212111213133100021101115305501 Geomydoecus_wernecki_planiceps 1121114122110311124113234114113111302123001001202125305300 Geomydoecus_wernecki_wernecki 1121114122110311113122234114112111212123001001202125305300 Geomydoecus_yucatanensis 11211132422103111351211140223120001331021021000111100078?0 Thomomydoecus_arleneae 232111314221011243111?1220223210020221210010020001102159?0 Thomomydoecus_asymmetricus 132111313221012243111112202231110202222?0010020000100184?0 Thomomydoecus_barbarae 23211131422101124311121220223210020221210010020001102159?0 Thomomydoecus_birneyi 132111313221012243111112102231110202222?0010020000100184?0 Thomomydoecus_byersi ?????????????????????????????????20221210010020011201159?0 Thomomydoecus_dickermani 132111313221012243111112102231110202222?0010020000100184?0 Thomomydoecus_genowaysi 132111313221012243111112102231110202222?0010020000100184?0 Thomomydoecus_greeri 13211131322101224311111210223111020222210010020000100184?0 Thomomydoecus_jamesbeeri 132111311221012243121?12202231111202222?0010020000100184?0 Thomomydoecus_johnhafneri 132111314221012243111111102231110202222?0010020100000184?0 Thomomydoecus_markhafneri 132111313221012243111112102231110202222?0010020000000184?0 Thomomydoecus_minor 132111314221012243111112101231110202222?0010020000100184?0 Thomomydoecus_neocopei 132111313221012243111112102231110202222?0010020000101184?0 Thomomydoecus_orizabae 132111311221012243111112202231110202212?2010020000000184?0 Thomomydoecus_peregrini 132111313221012243111212202231111202222?0010020000000184?0 Thomomydoecus_potteri ?????????????????????????????????20222210010020000000184?0 Thomomydoecus_timmi 132111313221012243111212102231110202222?0010020000100184?0 Thomomydoecus_wardi 23211131422101124311121220223210020221210010020000102159?0 Thomomydoecus_williamsi 132111313221012243111213202231110202222?0010020000100184?0 Thomomydoecus_zacatecae 132111311221012241111213102231111202222?0010020000100184?0 ; END; BEGIN ASSUMPTIONS; WTSET * Hennig86 SAW = 3: 7 32 34 38 45 49 , 2: 13 17 37 39-40 43 46-47 53 , 10: 5-6 12 23 42 44 55 57-58 , 0: 10 15 18 20-22 24-25 27 35 , 4: 3 8 30 41 50 54 , 1: 1 9 11 14 16 19 26 28-29 31 33 36 48 51-52 , 6: 2 4 , 8: 56 ; WTSET Hennig86 SAW = 1: 1 9 11 14 16 19 26 28-29 31 33 36 48 51-52 , 8: 56 , 6: 2 4 , 2: 13 17 37 39-40 43 46-47 53 , 4: 3 8 30 41 50 54 , 10: 5-6 12 23 42 44 55 57-58 , 0: 10 15 18 20-22 24-25 27 35 , 3: 7 32 34 38 45 49 ; END; [ The following blocks are output data for analysis step 824 ] BEGIN TREES; TITLE Tb5601; LINK TAXA = TaxaForAnalysisStep824; TREE Fig._5 = [&R] ((((((((((((((((((Geomydoecus_californicus,Geomydoecus_centralis,Geomydoecus_albati),(Geomydoecus_bajaiensis,Geomydoecus_clausonae,Geomydoecus_angularis)),Geomydoecus_subcalifornicus),(Geomydoecus_limitaris_limitaris,(Geomydoecus_limitaris_bakeri,Geomydoecus_aurei_grahamensis),Geomydoecus_crovelloi,Geomydoecus_warmanae,Geomydoecus_guadalupensis,Geomydoecus_limitaris_tolteci,Geomydoecus_sinolae,Geomydoecus_limitaris_halli,Geomydoecus_aurei_aurei,Geomydoecus_actuosi),(Geomydoecus_shastensis,Geomydoecus_hueyi)),(Geomydoecus_welleri_welleri,Geomydoecus_welleri_multilineatus)),Geomydoecus_jaliscoensis,Geomydoecus_umbrini,(Geomydoecus_extimi,Geomydoecus_nayaritensis)),Geomydoecus_musculi),Geomydoecus_cliftoni),Geomydoecus_quadridentatus),(Geomydoecus_subgeomydis,Geomydoecus_ewingi),Geomydoecus_nebrathkensis,Geomydoecus_oklahomensis,Geomydoecus_spickai,Geomydoecus_heaneyi,Geomydoecus_dalgleishi,Geomydoecus_geomydis),Geomydoecus_illinoensis),(Geomydoecus_oregonus,Geomydoecus_idahoensis)),((Geomydoecus_telli,(Geomydoecus_coronadoi_coronadoi,Geomydoecus_coronadoi_saccharalis,Geomydoecus_coronadoi_submerriami),Geomydoecus_veracruzensis),Geomydoecus_polydentatus_polydentatus,Geomydoecus_polydentatus_angustirostris,Geomydoecus_merriami)),(Geomydoecus_expansus,Geomydoecus_martini)),((Geomydoecus_truncatus,Geomydoecus_neotruncatus),(Geomydoecus_scleritus,Geomydoecus_mobilensis))),(Geomydoecus_trichopi,((((Geomydoecus_nadleri,Geomydoecus_bulleri_intermedius),Geomydoecus_burti),Geomydoecus_bulleri_melanuri,Geomydoecus_bulleri_bulleri),((((Geomydoecus_mexicanus,Geomydoecus_traubi),((Geomydoecus_perotensis_perotensis,Geomydoecus_perotensis_irolonis),Geomydoecus_fulvescens)),(Geomydoecus_mcgregori,(Geomydoecus_wernecki_wernecki,Geomydoecus_wernecki_planiceps))),Geomydoecus_alcorni))),((((((Geomydoecus_copei,(Geomydoecus_yucatanensis,(Geomydoecus_chapini,Geomydoecus_hoffmani))),((Geomydoecus_chiapensis,Geomydoecus_pygacanthi),(Geomydoecus_alleni,Geomydoecus_jonesi))),((Geomydoecus_setzeri,Geomydoecus_panamensis_dariensi,Geomydoecus_panamensis_panamensis,Geomydoecus_davidhafneri),(Geomydoecus_costaricensis,Geomydoecus_cherrei))),Geomydoecus_subnubili),Geomydoecus_ustulati_ustulati,Geomydoecus_tamaulipensis,Geomydoecus_ustulati_clarkii),Geomydoecus_texanus_texanus,Geomydoecus_texanus_tropicalis)),Geomydoecus_tolucae,Geomydoecus_fulvi,Geomydoecus_pattoni,Geomydoecus_chihuahuae_chihuahuae,Geomydoecus_chihuahuae_emersoni,(Geomydoecus_thomomyus,Geomydoecus_duchesnensis,Geomydoecus_craigi,Geomydoecus_biagiae,(Geomydoecus_betleyae,Geomydoecus_thaeleri),Geomydoecus_fuchsi,Geomydoecus_dakotensis))Geomydoecus,(Thomomydoecus_byersi,((Thomomydoecus_neocopei,Thomomydoecus_genowaysi,Thomomydoecus_greeri,Thomomydoecus_potteri,Thomomydoecus_johnhafneri,Thomomydoecus_markhafneri,Thomomydoecus_minor,Thomomydoecus_birneyi,Thomomydoecus_timmi,Thomomydoecus_dickermani,Thomomydoecus_asymmetricus,(Thomomydoecus_jamesbeeri,Thomomydoecus_zacatecae),Thomomydoecus_peregrini,Thomomydoecus_williamsi,Thomomydoecus_orizabae),Thomomydoecus_wardi,Thomomydoecus_barbarae,Thomomydoecus_arleneae))Thomomydoecus); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr3504] END;