#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on November 13, 2026; 3:32 GMT TreeBASE (cc) 1994-2008 Study reference: Andújar C., Serrano J., & Gómez-zurita J. 2012. Winding up the molecular clock in the genus Carabus (Coleoptera: Carabidae): assessment of methodological decisions on rate and node age estimation. BMC Evolutionary Biology, 12: 40. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S12410] [ The following blocks are input data for analysis step 8967 ] BEGIN TAXA; TITLE TaxaForAnalysisStep8967; DIMENSIONS NTAX=27; TAXLABELS Carabus_abbreviatus_BABA44 Carabus_abbreviatus_TENE15007 Carabus_auronitens_ROMA93906 Carabus_cancellatus_ROMA94106 Carabus_cancellatus_TACHcc816 Carabus_clathratus_TURQ51901 Carabus_coriaceus_TURQ91306 Carabus_dufouri_ZUHE111 Carabus_famini_EURY1625 Carabus_famini_MORR54807 Carabus_irregularis_ROMA94006 Carabus_lusitanicus_VESC5 Carabus_macrocephalus_FORO157 Carabus_melancholicus_EALM42207 Carabus_melancholicus_ROBU37 Carabus_melancholicus_TIDI39104 Carabus_monilis_SAOU1538 Carabus_morbillosus_MAZA5908 Carabus_morbillosus_TUNA1622 Carabus_morbillosus_TUNB1623 Carabus_morbillosus_TUNC1624 Carabus_nemoralis_RONC1549 Carabus_riffensis_KETA569 Carabus_rugosus_KSAR65807 Carabus_rugosus_SMAR43907 Carabus_rutilans_SENY1548 Carabus_violaceus_POPO83000 ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M12272] TITLE Ingroup_ITS2_Q_INSI_200_nogaps; LINK TAXA = TaxaForAnalysisStep8967; DIMENSIONS NCHAR=302; FORMAT DATATYPE=DNA SYMBOLS= "A C G T" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Carabus_abbreviatus_BABA44 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGATCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCCAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATGAATGGGTCGTCCGTAGATACTTGGCTTGACGCGTATATGCATGCATACACACGCTTTACG Carabus_abbreviatus_TENE15007 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGATCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCCAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATGAATGGGTCGTCCGTAGATACTTGGCTTGACGCGTATATGCATGCATACACACGCTTTACG Carabus_auronitens_ROMA93906 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATAAGTTTATTCACACAGTGCACGACTCTCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAAATGGGACGTCCGTAGATACTTGGCTTAACGAACATTAGTGCGCATACACACGCTTTATG Carabus_cancellatus_ROMA94106 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGATCGATGATCTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATAACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACAAAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCACCATGATAAATGGATCGTTCGTAGATACTTGGCTTAATAAACATACCATTGCATACACACGCTTTACG Carabus_cancellatus_TACHcc816 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGATCGATGATCTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATAACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACAAAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCACCATGATAAATGGATCGTTCGTAGATACTTGGCTTAATAAACATACCATTGCATACACACGCTTTACG Carabus_clathratus_TURQ51901 CGGATACTGTTCCTGGACCCGCCTGTCTGAGGGTCGTGCTATCTTACCAGACTGCCGGTCGACGATGTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATTACACTGCCCGTTTCCTTAAATTACTGTTCACAATACCGAATAATGTTATTCACACAGTACACGTCTCCCGTATACCAATAAGTGCGGAGTTGTCGGACGTGTTTACATGATAAATGGCTCGTCCGTAGATACTTGGCTTGAGGAACGTACATACGTATACACACGCTTTATG Carabus_coriaceus_TURQ91306 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATAAGTTTATTCACACAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAGTTGGGTCGTCCGTAGATACTTGGCTTAACAAGCATATGTATACATACACACGTTTTACG Carabus_dufouri_ZUHE111 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCACTACACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACACAACCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCTACATGATGAATGGCTCGTCCGTAGATACTTGGCTTAACGCATATATGCTTGCATACACACGCTTTACG Carabus_famini_EURY1625 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGATCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATGAGTTTATTCACACAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAAATGGGTCGTCCGTAGATACTTGGCTTAACGCGAATATGCATGCATACACACGCTTTACG Carabus_famini_MORR54807 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGATCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATGAGTTTATTCACACAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAAATGGGTCGTCCGTAGATACTTGGCTTAACGCGATTATGCATGCATACACACGCTTTATG Carabus_irregularis_ROMA94006 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACGTGTTGTGTGCAATTCACATTACACTGCCCGTTTCCTTAAATCACTGTTCACATAAACGAATAATTTTATTCACACTGTACACGTCTCCCGTATATTAATAGGTGCGGAGTTGTCGGACGTGTCCAAATGGTATAGAGGTCGTCCGTAGATACTTGGCCTAACAGAGTCAAGCATACACAATTGCGCTTTACA Carabus_lusitanicus_VESC5 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCACTACACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACACAACCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCTACATGATGAATGGCTCGTCCGTAGATACTTGGCTTAACGCATATATGCTTGCGTACACACGCTTTACG Carabus_macrocephalus_FORO157 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCACTACACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACACAACCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCTACATGATGAATGGCTCGTCCGTAGATACTTGGCTTAACGCATATATGCTTGCATACACACGCTTTACG Carabus_melancholicus_EALM42207 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGTCGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAATCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGAATGTGTTTACATGATAGGTGACTCGTTCGTAGATACTTGGCTTAACAAACGTATGAATGCATACACACGCTTTATG Carabus_melancholicus_ROBU37 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGTCGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAATCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGAATGTGTTTACATGATAGGTGACTCGTTCGTAGATACTTGGCTTAACAAACGTATGTATGCATACACACGCTTTATG Carabus_melancholicus_TIDI39104 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGTCGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAATCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGAATGTGTTTACATGATAAGTGACTCGTTCGTAGATACTTGGCTTAACAAACGTATGTATGCATACACACGCTTTATG Carabus_monilis_SAOU1538 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATCACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAGATGGGTCGTCCGTAGATACTTGGCTTAACAAACATATGTATGCGTACACATCGCTTTAC Carabus_morbillosus_MAZA5908 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTTGATGATCCCGGCGACGATGGGTATCGGGGAACCAGTGGTGTCTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTGCACGACTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGACCACATAATAAATGGGTCGTCCGTAGATACTTGGCTTAATGAACATATGTATGCATACACACGCTTTATG Carabus_morbillosus_TUNA1622 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTTGATGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTCTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTGCACGACTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGACCACATAATAAATGGGTCGTCCGTAGATACTTGGCTTAATGAACATATGTATGCATACACACGCTTTATG Carabus_morbillosus_TUNB1623 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTTGATGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTCTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTGCACGACTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGACCACATAATAAATGGGTCGTCCGTAGATACTTGGCTTAATGAACATATGTATGCATACACACGCTTTATG Carabus_morbillosus_TUNC1624 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTTGATGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTCTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTGCACGACTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGACCACATAATAAATGGGTCGTCCGTAGATACTTGGCTTAATGAACATATGTATGCATACACACGCTTTATG Carabus_nemoralis_RONC1549 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGGCGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATATGTTTATTCACACAGTGCACGTCTCCTGTATGCTAATAAGTGCGGAGTTGTCGGGTGAATCAACATGATGGATGATTCATTCGTAGATACTTGGCTTAATGAACATATGTATGCATACGTTCGCTTTATG Carabus_riffensis_KETA569 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTTTCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCACTGCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTACACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCAACATGATGAATGACTCGTCCGTAGATACTTGGCTTAACGCATATATGTGTGCATACACACGCTTTACG Carabus_rugosus_KSAR65807 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTCTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCTCTGTTCACATAACCGAATAAGTTTATTCACACAGTGCACGACTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGACCACATAATAAATGGGTCGTCCGTAGATACTTGGCTTAACGAACATATGTATGCATACACACGCTTTATG Carabus_rugosus_SMAR43907 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTCTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACATAACCGAATAAGTTTATTCACACAGTGCACGACTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGACCACATAATAAATGGGTCGTCCGTAGATACTTGGCTTAACGAACATATGCATGCATACACACGCTTTATG Carabus_rutilans_SENY1548 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGATGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACGTTACACTGCCCGTTTCCTTAAATCACTGTTCACAAAACCGAATAAGTTTATTCACACAGTGCACGACTCTCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAAATGGGACGTCCGTAGATACTTGGCTTAACGAACATTAGTGCGCATACACACGTTTTACG Carabus_violaceus_POPO83000 CGGATACTGTTCCTGGACCCGCCTGGCTGAGGGTCGTGCTATATTACCAGACTGCCGGTCGACGATCTCGGCGACGATGGGTATCGGGGAACCAGTGGTGTTTTGTTACTTGTCGTGTGCAATTCACATCACACTGCCCGTTTCCTTAAATCACTGTTCACAATACCGAATAAGTTTATTCACACAGTGCACGTCTCCCGTATGCTAATAAGTGCGGAGTTGTCGGACGTGTCCACATGATAGATGGGTCGTCCGTAGATACTTGGCTTAACAAACATATGTATACATACACATCGCTTTTA ; END; [ The following blocks are output data for analysis step 8967 ] BEGIN TREES; TITLE Ingroup_Results_with_27_taxa_C; LINK TAXA = TaxaForAnalysisStep8967; TREE Ingroup_ITS2_Q_INSI_200_nogaps_ULN = [&R] (((((Carabus_lusitanicus_VESC5:2.551554800887601,(Carabus_macrocephalus_FORO157:0.6987287161759959,Carabus_dufouri_ZUHE111:0.6987287161759959):1.8528260847116016):7.742003272596101,Carabus_riffensis_KETA569:10.293558073483702):5.932644712465596,((Carabus_abbreviatus_BABA44:0.810310105472194,Carabus_abbreviatus_TENE15007:0.810310105472194):8.572723373225802,(Carabus_famini_MORR54807:2.6616694979279956,Carabus_famini_EURY1625:2.6616694979279956):6.721363980770004):6.843169307251298):6.531231020629374,((((Carabus_morbillosus_MAZA5908:2.3726468923410025,(Carabus_morbillosus_TUNC1624:1.1454637070714924,(Carabus_morbillosus_TUNA1622:0.5082567028539557,Carabus_morbillosus_TUNB1623:0.5082567028539557):0.6372070042175366):1.2271831852695065):5.088229655854329,(Carabus_rugosus_KSAR65807:2.9864268940333005,Carabus_rugosus_SMAR43907:2.9864268940333005):4.474449654162031):7.453620993626215,(Carabus_auronitens_ROMA93906:3.408230763689037,Carabus_rutilans_SENY1548:3.408230763689037):11.50626677813251):3.250853857785952,Carabus_nemoralis_RONC1549:18.1653513996075):4.592082406971173):7.775388895500328,(((Carabus_cancellatus_TACHcc816:0.6906517902820006,Carabus_cancellatus_ROMA94106:0.6906517902820006):17.7561139634285,(Carabus_irregularis_ROMA94006:17.0743681332789,(Carabus_coriaceus_TURQ91306:13.700828882413001,(Carabus_monilis_SAOU1538:3.871538634841201,Carabus_violaceus_POPO83000:3.871538634841201):9.8292902475718):3.373539250865898):1.3723976204316024):1.4028835775089483,(((Carabus_melancholicus_ROBU37:1.1262266146429987,Carabus_melancholicus_EALM42207:1.1262266146429987):1.7752035588965516,Carabus_melancholicus_TIDI39104:2.9014301735395502):11.861378223972551,Carabus_clathratus_TURQ51901:14.762808397512101):5.086840933707348):10.68317337085955); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr52201] END;