#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on May 14, 2026; 0:41 GMT TreeBASE (cc) 1994-2008 Study reference: Tsuihiji T., Watabe M., Tsogtbaatar K., Barsbold R., & Suzuki S. 2012. A tyrannosauroid frontal from the Upper Cretaceous (Cenomanian-Santonian) of the Gobi Desert, Mongolia. Vertebrata PalAsiatica, 50: 102-110. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S13016] [ The following blocks are input data for analysis step 9672 ] BEGIN TAXA; TITLE TaxaForAnalysisStep9672; DIMENSIONS NTAX=24; TAXLABELS Albertosaurus Alioramus Allosaurus Appalachiosaurus Bistahieversor Compsogs Daspletosaurus Dilong Dryptosaurus Eotyrannus Gorgosaurus Guanlong Kileskus Maniraptora Ornithomimos Proceratosaurus Raptorex Sinotyrannus Stokesosaurus Tarbosaurus Tyrannosauridae_MPC_D_102_4 Tyrannosaurus Utah Xiongguanlong ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M13677] TITLE Character_Matrix; LINK TAXA = TaxaForAnalysisStep9672; DIMENSIONS NCHAR=307; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 ] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ] Albertosaurus 01101011120000000000{01}12001{01}1001011101201201111002111{01}1{02}1?01?0010000001000102010020011100100101?11310111111111011012100111111111000{01}000110{01}{01}1111000010110111101111110111111110010111212111100?12?10?2121011????????11??????????????????01111110212101111111?010112111111111101211111012111??1???1???1111111111111201 Alioramus 001011??2?????0000011110111110?011101201101012012101010211101011100010000102000010000100101110011320111111110011012100111111111?101110010011101000110110111111011011111011110010111212211110??10000????00000100000111110?11011?21011111??????????????????????????1?111110110?????????21?11?10111121??00?11111?11?01 Allosaurus 01000000000001010000001000000100011000000000021021000100000010001????0000000001?0??100?00000?00000000000001100000100000000?0?10000000000????000?0001001000000001000000000000000000000200000000000000000000100000010000002001000{01}0000100000000010000000000001001010000000000100?000101000100000010201100000000000110 Appalachiosaurus ??1???????????000000?0100?01?010111?12011?1?1?0020?101?10??0?0??????????????????????????????????????????????????????????????????000000010100010?00?????????????????????11?110??01??????????0??1????2?2?0?????????????????????????????0??????????????????????????????????????1???????12111?010110???11111111111011?0 Bistahieversor ?0?0101112000000000?10210101?010111012011011010020110001001???10000??0000002000021100100100000??13111111?1?10011012100121111111?01000?010011111??????11???1?0101?1101111111100?0111212201100112?10?2?2?00????111?????????????????????0111?11??????????????????????????????????????????????010?1????0110111111?11001 Compsogs ?000000000000000000?00100?00?0?0000?000000??02000??00000?00??00?0????000000000010??010?10??0??0000?000???????100000000?000?0??0??????00?????????0??????????????????????0?000000000000000???0???????0000{01}00???????00000??00000?000?0?0??0000010000?0??00100?{01}0000000{01}0?00000000?00?0010000??0??00???0??00?000?0???00 Daspletosaurus 0110101122000110112011200211102111101201211012012101101{12}1111111111111011010200002110011011{01}11111132121111111001?112101221111111110111001101111{01}100111110111111011111111111110010111222211101112?100212101100011101111110211110????????1111111?212111?11111?111112111111??1??1?11111?12111??1?????2?11???1?111?11??? Dilong 0000001101001?01001?10000100?0?000000100000000103??00100?00??00010111100001100010??000?11000??0001000000010?????000000?00101000???????????????????020?0100?0010?0??????01001??100?010002?00?11?????101110000000100000001000000????????000000110000001???0??010000100?0?000010??00010??000??001?00000000010000??0100 Dryptosaurus ???????????????????????????????0???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????0??1212?1?1????????????????????????????????????????????????????10100?????1?????11????????????1???????12????0???1012?1?011111?????1?? Eotyrannus ???0??11?100?01110????????00?0?01????20110000?1010?0?1??0?00?00????????????????0????????????????????????01?1???0?????????????????????001??????0?00????????????????????????000?100002110100?????????2?2??00???00??0??00???1?0???????????00???011011011??0000???00?2???????????????????????????????????01???000?00100 Gorgosaurus 01101011120000000000{01}120010100101110120120111200211101{02}100100010000001000102010020011100100101011310111111111011012100111111111000100011011111??00110110101101111110111111110010111212111100112?10?2121011????????11????11101?????1?0??11111102121010111111010112111111101101211111012111?010?11?2?11111111111?1201 Guanlong 0001000101111001000?10100?00?10000010?1000000?003??00000??0??00?10001000001100010??010?10000?000010000000000?100000011?00100000?00000?????????????020?010000010000000000000????10?0000000010???????1010100???????0000001000001010000?0?000000100000010000000000001000000000001000000000000?0010000000010000000?0000 Kileskus ???1??110111100100???0?00?0000?0000?0?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????0100010??????????????00?????????????????????????????????????????????????????????????????????????????????????????????????000?????0 Maniraptora 00000?0000000000000?00100?00?00011100000001000000??000000000?01?00???0000000000?0??010?01000?000000001001000????0000000000?0?0?0000??001000?0?0?00000{01}000000{01}010000000000000000000000001{01}0000000?000000{01}0000000{01}0{01}00001?111100{01}1000?000000000000100000000000000000000000000001100??01100000000100000000010000000000 Ornithomimos 0000001000000?00000?00000?00?00010000000001?00000??00000?00??00?0000?0000000000?0??010?00?00??00000?01000000?000000000?000?0?0000??00001000?????0?00000100000000{01}00???000000???00000000000000000???000000000?00?000000000000100{12}00000000000000001000000{01}000{01}0101000100100{01}0000?0{01}00011001100000000000000100100?0000 Proceratosaurus 00?1001101111?01000?10100?00?1?000010?1???0?0?????????????????0?1000100000110001?????0??0??0?0???100000000??0??0???????????????0000??000?????0??0?0???????00?100???????0001???110?000000001000?????111000?????????????????????????????????????????????????????????????????????????????????????????????????????????? Raptorex ?1100?111200?0000010110001010010111012011010020010?10101?00?000000011000000100000??000?01000??0012?011???1?1??110111000011111?????????0?????????0????????????100?0????001000001011121211??0011??0??212100000?0000000??100110??1?????0??0110000212101111?11???111101111100010011???1112101??1??1?12?0?1??111?1?????? Sinotyrannus ???1???10?1?1?0110??10????0??1?00?????1???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????0????1????????????????????????0?????????????????????????????????????????????????????00?10?0??00001??????????????????????????????????????? Stokesosaurus ?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????0???????00?00?10000100?00100?0?????????????????????????120000000001?1000110010010??????1100001???????????? Tarbosaurus 1110111222000110112101211301102111101201211012010??1101211111110111110111102101021100210111111111321211111110010113121230111111111111101101111111111111011100201111111111111112011122212?1011121111212112111111?11111?102111111??1??1?11111110212111011111?11111211111111110121111111211111112111211111111111111211 Tyrannosauridae_MPC_D_102_4 ????????????????????????????????????????????????????????????????????????????????????????????????????????????????00111021??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Tyrannosaurus 11101112220001101121012{01}0{13}01102111101201211112010??110121011111011{01}1101111021010211002101111111113{12}1211111110011113121230111111111111101101111111111111011100201111111111111112011122222110111211112121121111110111111102111101211111111111110212111011111111111211111111110121111111211111112111211111111111111211 Utah ????????????????000?????0?01?????????????????2?021?100010?10??????????0???21?1??????0????01???0?13??21?111?10011112101211???????????????????????????011?????020?1?1???11??1100?011??????????11?????????10????????1????1????????????????????1?0211????1????????????????????????????????????010?11?1????????????????? Xiongguanlong 00100?1102000????0??1??00?01?0?01?0001001010??0010?00100??0??0??0?????0???01000?0???00?01000???0021?111110010?1????100?1?111??100?100????????????00100100000?10?1??????????????????????????????????2?2?000??000000000110000010??????????????????????????????????11?101??0011??????????????01010012????????????????? ; END; [ The following blocks are output data for analysis step 9672 ] BEGIN TREES; TITLE Tyrannosauridae; LINK TAXA = TaxaForAnalysisStep9672; TREE Fig._8 = [&R] (Allosaurus,((Compsogs,(Maniraptora,Ornithomimos)),((Kileskus,(Guanlong,Proceratosaurus,Sinotyrannus)),(Dilong,(Eotyrannus,Stokesosaurus,Xiongguanlong,Raptorex,Dryptosaurus,Appalachiosaurus,Alioramus,Bistahieversor,Tyrannosauridae_MPC_D_102_4,(Albertosaurus,Gorgosaurus),(Utah,(Daspletosaurus,(Tarbosaurus,Tyrannosaurus)))))))); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr55254] END;