#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on February 04, 2026; 4:24 GMT

TreeBASE (cc) 1994-2008

Study reference:
Schott R.K., Evans D.C., Williamson T.E., Carr T.D., & Goodwin M.B. 2009. The anatomy
and systematics of Colepiocephale lambei (Dinosauria: Pachycephalosauridae).
Journal of Vertebrate Paleontology, 29(3): 771-786.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S13241]

[ The following blocks are input data for analysis step 9931 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep9931;
        DIMENSIONS NTAX=16;
        TAXLABELS
            Ceratopsidae
            Colepiocephale
            Goyocephale
            Hanssuesia
            Homalocephale
            Ornithopoda
            Pachycephalosaurus
            Prenocephale_brevis
            Prenocephale_edmontonensis
            Prenocephale_goodwini
            Prenocephale_prenes
            Stegoceras
            Stenopelix
            Stygimoloch
            Tylocephale
            Wannanosaurus
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M14018]

	TITLE Character_Matrix_A;
	LINK TAXA = TaxaForAnalysisStep9931;
	DIMENSIONS NCHAR=49;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1" MISSING=? GAP= -;

MATRIX
[                                      10        20        30        40        50     ]
[                                       .         .         .         .         .     ]
Ceratopsidae                   0000000000000000000000000000000000000?00000000000
Colepiocephale                 ???????1???????????????1??????11100??????????0?1?
Goyocephale                    1?1111111111?11111111110???0??00000??000000000000
Hanssuesia                     ???1?1110????1?1?0?????1??????11111???0?000000001
Homalocephale                  1?11?1111???11111?1111111111110000000000000000000
Ornithopoda                    0000000000000000000000000000000000000?00000000000
Pachycephalosaurus             ???1?1110????1111??????11?????1111110111000001000
Prenocephale_brevis            ???1??111????1?1???????1??????11111???0?1??110000
Prenocephale_edmontonensis     ???1??111????1?1???????1??????11111???0?111111000
Prenocephale_goodwini          ???1?1111????1?1???????1??????11111???0?1110100??
Prenocephale_prenes            1?1111111????1111??1?11111111111111110001{01}0000000
Stegoceras                     ?11111111111111111111111111111111{01}000000000{01}00000
Stenopelix                     111??????00?0??????0?00????000???????????????????
Stygimoloch                    ???1?1110????1?1???????1??????11111??111000000000
Tylocephale                    ???1?1111????111?1?????11?????1111111000100000000
Wannanosaurus                  ???1111111111000?0?????0??????00000??00?000000000
;
END;

[ The following blocks are output data for analysis step 9931 ]
BEGIN TREES;
      TITLE Analysis_results_1;
      LINK TAXA = TaxaForAnalysisStep9931;
      TREE Figure_8A = [&R] (Ornithopoda,(Ceratopsidae,(Stenopelix,(Wannanosaurus,(Goyocephale,(Homalocephale,(Stegoceras,Colepiocephale,((Tylocephale,Prenocephale_prenes,(Prenocephale_goodwini,(Prenocephale_brevis,Prenocephale_edmontonensis))),(Hanssuesia,(Stygimoloch,Pachycephalosaurus))))))))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr55992]


END;