#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on February 04, 2026; 4:24 GMT

TreeBASE (cc) 1994-2008

Study reference:
Schott R.K., Evans D.C., Williamson T.E., Carr T.D., & Goodwin M.B. 2009. The anatomy
and systematics of Colepiocephale lambei (Dinosauria: Pachycephalosauridae).
Journal of Vertebrate Paleontology, 29(3): 771-786.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S13241]

[ The following blocks are input data for analysis step 9932 ]
BEGIN TAXA;
        TITLE  TaxaForAnalysisStep9932;
        DIMENSIONS NTAX=21;
        TAXLABELS
            Goyocephale_lattimorei
            Homalocephale_calathocercos
            Pachycephalosauridae_indet_Ornatotholus_browni_holotype_AMNH_5450
            Pachycephalosauridae_indet_SMP_VP_1084
            Pachycephalosaurus_wyomingensis
            Prenocephale_prenes
            Psittacosaurus_mongoliensis
            Sphaerotholus_buchholtzae_holotype_TMP_87.113.3
            Sphaerotholus_goodwini_holotype_NMMNH_P_27403
            Stegoceras_breve_holotype_CMN_1423
            Stegoceras_edmontonense_holotype_CMN_8830
            Stegoceras_lambei_holotype_CMN_8818
            Stegoceras_sp._Gravitholus_albertae_holotype_TMP_72.27.1
            Stegoceras_sp._UCMP_130051
            Stegoceras_sternbergi_holotype_CMN_8817
            Stegoceras_validum_UA_2
            Stenopelix_valdensis
            Stygimoloch_spinifer
            Tylocephale_gilmorei
            Wannanosaurus_yanshiensis
            Yaverlandia_bitholus
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M14022]

	TITLE Character_Matrix_B;
	LINK TAXA = TaxaForAnalysisStep9932;
	DIMENSIONS NCHAR=53;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3" MISSING=? GAP= -;

MATRIX
[                                                                             10        20        30        40        50   ]
[                                                                              .         .         .         .         .   ]
Goyocephale_lattimorei                                                1?111111111?1111011110??100??0??001100100010011110011
Homalocephale_calathocercos                                           1?11?111???1111?11111111111110000011001000110111100??
Pachycephalosauridae_indet_Ornatotholus_browni_holotype_AMNH_5450     ???1???1????1????????1???????1??00?????11010?????00??
Pachycephalosauridae_indet_SMP_VP_1084                                ???1?111????1????????1???????1???2?????12????????01??
Pachycephalosaurus_wyomingensis                                       ???1?111????11???????11??????11002111?212110100111122
Prenocephale_prenes                                                   1?111111????11?111?1111111111111021202112111102120111
Psittacosaurus_mongoliensis                                           00000000000000000000000000000000000000000000000000000
Sphaerotholus_buchholtzae_holotype_TMP_87.113.3                       ???1?111????1????????1???????1??0212111121101??1301??
Sphaerotholus_goodwini_holotype_NMMNH_P_27403                         ???1?111????11?1?????1??1????1??0212111121111021301??
Stegoceras_breve_holotype_CMN_1423                                    ???1???1????1????????1???????1??121?1?111110???1?01??
Stegoceras_edmontonense_holotype_CMN_8830                             ???1???1????1????????1???????1??021???112110???1?01??
Stegoceras_lambei_holotype_CMN_8818                                   ???1???1????1????????1???????1??11?????11220???0?02??
Stegoceras_sp._Gravitholus_albertae_holotype_TMP_72.27.1              ???1???1????1????????1???????1??11?????112?0???1?02??
Stegoceras_sp._UCMP_130051                                            ???1?111????1????????1???????1??1100000112200??1102??
Stegoceras_sternbergi_holotype_CMN_8817                               ???1???1????1????????1???????1??11?????11220???1?02??
Stegoceras_validum_UA_2                                               ?1111111111111111111111111111100111200111220011120200
Stenopelix_valdensis                                                  111?????00?0?????0?00????0000????????????????????????
Stygimoloch_spinifer                                                  ???1??11????1????????1???????1??0211??2121111????1122
Tylocephale_gilmorei                                                  ???1?111????1111?????11??????1110212011121111??1101??
Wannanosaurus_yanshiensis                                             ???11111111100?00????0???????0???00000?00??0???1??0??
Yaverlandia_bitholus                                                  ???1?1?0????1???????????????????00???????00???????0??
;
END;

[ The following blocks are output data for analysis step 9932 ]
BEGIN TREES;
      TITLE Analysis_results_2;
      LINK TAXA = TaxaForAnalysisStep9932;
      TREE Figure_8B = [&R] (Psittacosaurus_mongoliensis,(Stenopelix_valdensis,Wannanosaurus_yanshiensis,Yaverlandia_bitholus,(Goyocephale_lattimorei,(Homalocephale_calathocercos,(Pachycephalosauridae_indet_Ornatotholus_browni_holotype_AMNH_5450,((Tylocephale_gilmorei,Prenocephale_prenes,Pachycephalosauridae_indet_SMP_VP_1084,Stegoceras_edmontonense_holotype_CMN_8830,Sphaerotholus_goodwini_holotype_NMMNH_P_27403,Sphaerotholus_buchholtzae_holotype_TMP_87.113.3,(Stygimoloch_spinifer,Pachycephalosaurus_wyomingensis)),(Stegoceras_breve_holotype_CMN_1423,(Stegoceras_sternbergi_holotype_CMN_8817,Stegoceras_lambei_holotype_CMN_8818,Stegoceras_validum_UA_2,Stegoceras_sp._Gravitholus_albertae_holotype_TMP_72.27.1,Stegoceras_sp._UCMP_130051))))))));
[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr55993]


END;