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Matrix 5975 of Study 10671

About Citation title: "Systematics of grammitid ferns (Polypodiaceae): using morphology and plastid sequence data to resolve the circumscriptions of Melpomene and the polyphyletic genera Lellingeria and Terpsichore.".
About Study name: "Systematics of grammitid ferns (Polypodiaceae): using morphology and plastid sequence data to resolve the circumscriptions of Melpomene and the polyphyletic genera Lellingeria and Terpsichore.".
About This study is part of submission 10661 (Status: Published).

Matrices

Title: Morphology and Indels

Rows

Taxon Label Row Segments Characters 1?–30
Calymmodon gracilis  (none) 00000?000100?1000{01}100011102
Calymmodon luerssenianus  (none) 0-000-0-0100?100001020????????
Campyloneurum angustifolium  (none) 021011101100?10001001000---000
Ceradenia aulaeifolia  (none) 02101?011100?10100100021010001
Ceradenia aulaeifolia 2  (none) 02101?011100?10100100021010001
Ceradenia fucoides  (none) 02000-01010100----------------
Ceradenia intonsa  (none) 02101?011100?10100100021001001
Ceradenia jungermannioides  (none) 00100-010100?0----------------
Ceradenia kalbreyeri Colombia  (none) 02101?011100?10100100021000001
Ceradenia kalbreyeri Costa Rica  (none) 02101?011100?10100100021000001
Ceradenia pilipes Costa Rica  (none) 02000?010100?001001000210{01}0
Ceradenia pilipes Ecuador  (none) 02000?010100?001001000210{01}0
Ceradenia sp. aff. kalbreyeri  (none) 02101?0??100?1010010002100000?
Ceradenia sp. ined. 1  (none) 02101?011100?101001000210{01}1
Ceradenia sp. ined. 1 2  (none) 02101?011100?101001000210{01}1
Ceradenia sp. moran 7520  (none) 02101-0??100?1010010002?????01
Ceradenia spixiana  (none) 02101?011100?101001000210{01}1
Chrysogrammitis musgraveana  (none) 0-001---0100?100-000201101----
Cochlidium punctatum  (none) 0?000?0?0100?1?00{01}100000---
Cochlidium rostratum  (none) 0?000-0-0100?1100{01}100000---
Cochlidium seminudum  (none) 00000?0?0100?10000100000---000
Ctenopteris heterophylla  (none) {0123}-001---0100?100-0102010-
Ctenopteris heterophylla 2  (none) {0123}-001---0100?100-0102010-
Ctenopteris lasiostipes  (none) 0-001---0100?100-0111010---0--
Ctenopteris nutans  (none) 0-001---1100?100-011101100----
Ctenopteris repandula  (none) {013}-000---0100?100-0103010--
Ctenopteris rhodocarpa  (none) 0-001---1100?100-011101100----
Enterosora enterosoroides  (none) 02101-011100?10000103010---100
Enterosora percrassa  (none) 0210110?1100?10100103011013011
Enterosora percrassa 2  (none) 0210110?1100?10100103011013011
Enterosora trichosora  (none) 02101?011100?10100103010---10?
Enterosora trifurcata Costa Rica  (none) 02101?011100?10100103011010011
Enterosora trifurcata Ecuador  (none) 02101?011100?10100103010---011
Grammitis billardieri  (none) {01}-1010000100?10000103000---
Grammitis ciliata  (none) {012}-000-0-0100?10000103010--
Grammitis deplanchei  (none) 001011001100?10100100010---100
Grammitis forbesiana  (none) 0-001-0-0100?10000103010---00-
Grammitis knutsfordiana  (none) 00001?100100?11000100110---000
Grammitis melanoloma  (none) 0?000?0?0100?10000103010---00?
Grammitis parva  (none) 0-000-0-0100?10000103010---00-
Grammitis poeppigiana  (none) 20001?010000?10000100010---000
Grammitis poeppigiana 2  (none) 20001?010000?10000100010---000
Grammitis pseudociliata  (none) 0-001-0-0100?10000103010---00-
Lellingeria hellwigii  (none) 00000?010100?11000011111102001
Lellingeria myosuroides 2  (none) 00000?010100?10001011111102101
Lellingeria prionodes  (none) 00000?010100?11000011111102001
Lellingeria apiculata Brazil  (none) 01101?000100?10000111011001011
Lellingeria apiculata Costa Rica  (none) 01101?000100?10000111011001011
Lellingeria bishopii  (none) 02101?0?0100?10100111001001001
Lellingeria humilis  (none) 11101?000100?1000011100100100?
Lellingeria limula  (none) 00000?010100?10001011110---001
Lellingeria limula 2  (none) 00000?010100?10001011110---001
Lellingeria limula 3  (none) 00000?010100?10001011110---001
Lellingeria major  (none) 01101?010100?10100111001001001
Lellingeria major 2  (none) 01101?010100?10100111001001001
Lellingeria major 3  (none) 01101?010100?10100111001001001
Lellingeria myosuroides  (none) 00000?010100?10001011111102101
Lellingeria pseudomitchellae  (none) 00000-0-0100?0----------------
Lellingeria saffordii  (none) 00000?000100?10001011111102001
Lellingeria schenckei  (none) 01000?0?010??1000011101100?00?
Lellingeria simacensis  (none) 01101?010100?10100111001001001
Lellingeria subimpressa  (none) 011011000100?10000111001001001
Lellingeria subsessilis Costa Rica  (none) 02101?000100?10100111001001001
Lellingeria subsessilis Ecuador  (none) 02101?000100?10100111001001001
Lellingeria suprasculpta  (none) 02001?000100?10100112000---00?
Lellingeria suspensa  (none) 02001?0?0100?10000111120---001
Lellingeria suspensa 2  (none) 02001?0?0100?10000111120---001
Melpomene anazalea ined.  (none) 13101?110000?10001112110---001
Melpomene firma  (none) 01101?0?0100?10011112120---001
Melpomene flabelliformis  (none) {02}0101?1?0000?1000{01}112120
Melpomene moniliformis  (none) {12}0101?110000?10001112120---
Melpomene occidentalis  (none) 001011010100?10001112110---001
Melpomene occidentalis 2  (none) 001011010100?10001112110---001
Melpomene personata ined.  (none) 01101-000100?10000112120---001
Melpomene pseudonutans  (none) {02}1101?0?0100?10001112120---
Melpomene vernicosa  (none) 01101?000100?10001112120---001
Microgramma percussa  (none) 001011101000?10001111000---100
Microgramma squamulosa  (none) 001011101000?10001111001001100
Micropolypodium achilleifolium  (none) {01}?000?000100?10000100000--0
Micropolypodium hyalinum  (none) 02000?1001111101011000210{01}2
Micropolypodium taenifolium  (none) 02000?000100?10100100021000001
Micropolypodium zurquinum  (none) 02000?000100?10100100020---001
Oreogrammitis hookeri  (none) 01001?010100?11000100110---000
Pecluma alfredii  (none) 011011001100?10000002000---000
Polypodium glycyrrhiza  (none) 1110111?1000?10000100000---100
Prosaptia alata  (none) 0210110?1100?10001111001002001
Prosaptia contigua  (none) 0?10110?1100?10000111011002001
Prosaptia obliquata  (none) 0100110?1100?10000113011002000
Prosaptia pubipes  (none) {02}?101?001100?10000111111002
Scleroglossum sulcatum  (none) 10000?0?0100?11000100000---000
Scleroglossum sulcatum 2  (none) 10000?0?0100?11000100000---000
Terpsichore aff. subtilis inded.  (none) 02101?010100?10100100021000001
Terpsichore alsopteris  (none) 02101?011100?10100100021000001
Terpsichore amphidasyon 2  (none) 011011001100?10100100121101001
Terpsichore amphidasyon 3  (none) 011011001100?10100100121101001
Terpsichore amphidasyon Moran 7646  (none) 011011001100?10100100121101001
Terpsichore anfractuosa Bolivia  (none) 00111?000100?10001011111002001
Terpsichore anfractuosa Bolivia 2  (none) ??????????????????????????????
Terpsichore anfractuosa Costa Rica  (none) 00111?000100?10001011111002001
Terpsichore anfractuosa Ecuador  (none) {01}0111?000100?10001011111102
Terpsichore anfractuosa Ecuador 2  (none) 00111?000100?10001011111002001
Terpsichore anfractuosa Mexico  (none) 00111?000100?10001011110-0-001
Terpsichore anfractuosa Mexico 2  (none) 00111?000100?10001011110-0-001
Terpsichore anfractuosa Peru  (none) 20111?000100?10001011111002001
Terpsichore anfractuosa Peru 2  (none) ??????????????????????????????
Terpsichore angustipes  (none) 02101?0?0100?10001111001002001
Terpsichore angustipes 2  (none) 02101?0?0100?10001111001002001
Terpsichore athyrioides  (none) 021010000100?10011111001002001
Terpsichore attenuatissima  (none) 02101?011100?10100100021000001
Terpsichore attenuatissima 2  (none) 02101-011100?10100100021000001
Terpsichore attenuatissima 3  (none) 02101?011100?10100100021000001
Terpsichore clathrata  (none) 02101?010100?10001111111002001
Terpsichore clavigera ined.  (none) 011011010100?10011111001002001
Terpsichore colombiense ined.  (none) 011011010100?10011111001002001
Terpsichore colombiense ined. 2  (none) ??????????????????????????????
Terpsichore cretata  (none) 02101?010100?10110101001100001
Terpsichore cuencana  (none) 00101?000100?10001111001002001
Terpsichore cultrata  (none) 02000?000100?10100100021001000
Terpsichore davidsmithii  (none) 00101?000100?1000111100100{12}
Terpsichore davidsmithii 2  (none) 00101?000100?1000111100100{12}
Terpsichore davidsmithii 3  (none) 00101?000100?1000111100100{12}
Terpsichore davidsmithii 4  (none) 00101?000100?1000111100100{12}
Terpsichore davidsmithii 5  (none) 00101?000100?1000111100100{12}
Terpsichore delicatula  (none) 021010000100?10100100001010011
Terpsichore dilatata  (none) 01101?000100?10001111001002001
Terpsichore dilatata 2  (none) 01101?000100?10001111001002001
Terpsichore hanekeana  (none) 02101?001110?101001000210{01}0
Terpsichore lanigera  (none) 021010000110?10100100021000010
Terpsichore lehmanniana  (none) 02100-000100?10000103001003010
Terpsichore leucosticta  (none) 021011011100?10100100021000001
Terpsichore leucosticta 2  (none) 021011011100?10100100021000001
Terpsichore loja ined.  (none) 01101?000100?10001111001002001
Terpsichore loja ined. 2  (none) 01101?000100?10001111001002001
Terpsichore loja ined. 3  (none) 01101?000100?10001111001002001
Terpsichore longicaulis  (none) 21101?110000?10101100021000001
Terpsichore longipilosa ined.  (none) 011011001100?10100100021101001
Terpsichore longipilosa ined. 2  (none) 011011001100?10100100021101001
Terpsichore longisetosa  (none) {01}?0011010100?10000100011010
Terpsichore mollissima  (none) 02000?0?0110?0----------------
Terpsichore nana  (none) 00101?000100?10011111001002001
Terpsichore pacifica  (none) 021011000100?10100100021100001
Terpsichore pichinchae  (none) 011011000100?10001111001002001
Terpsichore pichinchense  (none) 02101?000100?10001111011002001
Terpsichore pichinchense 2  (none) {01}2101?000100?10001111011002
Terpsichore pichinchense 3  (none) 02101?000100?10001111011002001
Terpsichore praeceps  (none) 1?101?010000?10100100021100001
Terpsichore semihirsuta  (none) 02101?011100?10100100021000001
Terpsichore semihirsuta Bolivia  (none) 021011011100?10100100021000001
Terpsichore semihirsuta Bolivia 2  (none) 021011011100?10100100021000001
Terpsichore semihirsuta Colombia  (none) 021011011100?10100100021000001
Terpsichore semihirsuta Colombia Choco  (none) 021011011100?10100100021000001
Terpsichore semihirsuta Ecuador  (none) 021011011100?10100100021000001
Terpsichore semihirsuta ined. 2 Costa Rica  (none) 02101?010100?10100100021000001
Terpsichore semihirsuta ined. 3 Mexico  (none) 02101?010100?10100100021000001
Terpsichore semihirsuta Peru  (none) 021011011100?10100100021000001
Terpsichore senilis  (none) 02000?000110?10100100001001010
Terpsichore smithii  (none) 020000000110?1010010002100001?
Terpsichore sp. aff. alsopteris ined. 1 Bolivia  (none) 02101?011100?10100100021000001
Terpsichore sp. aff. alsopteris ined. 1 Bolivia 2  (none) 02101?011100?10100100021000001
Terpsichore sp. aff. alsopteris ined. 1 Bolivia 3  (none) 02101?011100?10100100021000001
Terpsichore sp. aff. alsopteris ined. 1 Bolivia 4  (none) 02101?011100?10100100021000001
Terpsichore sp. aff. alsopteris ined. 2 Costa Rica  (none) 02101?011100?10100100021100001
Terpsichore sp. aff. taxifolia ined. Colombia  (none) 01101?010100?10100100021100001
Terpsichore sp. Sundue 1160  (none) 02101?0?0100?10101100021000001
Terpsichore sp. Sundue 1167  (none) 02101?0?0100?10110100021000001
Terpsichore subscabra  (none) 021011000100?10100100011010001
Terpsichore subscabra 2  (none) 021011000100?10100100011010001
Terpsichore subtilis  (none) 02101?0?0100?1011010102100100?
Terpsichore subtilis 2  (none) 02101?0?0100?1011010102100100?
Terpsichore taxifolia Brazil  (none) 0110110?0100?10100100021101001
Terpsichore taxifolia Ecuador  (none) 0110110?0100?10100100021101001
Terpsichore vascoi ined.  (none) 021011010100?10100100021000001
Terpsichore zeledoniana  (none) 011011011100?10100100020---000
Xiphopteris conjunctisora  (none) ?20011000100?10001110110---000
Zygophlebia mathewsii Costa Rica  (none) 021011011100?10100100020---001
Zygophlebia mathewsii Ecuador  (none) 021011011100?10100100020---001
Zygophlebia sectifrons  (none) 021011011100?10100100020---001

Columns

Column Character Description
1 habit
2 habit 2 position of fronds
3 root insertion
4 proliferous roots
5 rhizome symmetry
6 Stele perforations
7 rhizome phytomer internode length
8 branch buds
9 phyllopodia
10 rhizome branching
11 Rhizome setae
12 hairs on rhizome
13 rhizome hairs
14 rhizome scales
15 Rhizome scale attachment
16 rhizome scale cell shape
17 Rhizome scale cell lumen induration
18 scale base shape
19 scale shape
20 scales clathrate vs. concolorous [cell walls distinctly different color than lumen]
21 scale color [color of cell wall in clathrate scales]
22 scale iridescence
23 rhizome scale sheen
24 scale margin indument present vs. absent [distinct from apical gland]
25 distribution of marginal indument
26 scale margin indument type
27 color of marginal or apical setae
28 scale margin entire vs. denticulate [not inclusive of marginal indument, N/A for taxa with marginal indument]
29 scale surfaces
30 apical glandlike cell(s)
31 number of apical glandlike cells
32 fronds determinate vs. indeterminate
33 sterilefertile leaf dimorphy
34 Fertile portions of fronds more setose than sterile portions
35 leaf articulation
36 petiole shape in x section
37 petiole sclerenchyma outermost layer
38 Petiole stele number
39 petiole marginate or terete
40 blade base
41 blade apex
42 blade dissection at least at base
43 pinna base shape
44 pinna apex shape
45 Lamina texture
46 Leaf pallisade parenchyma layer
47 Leaf spongy parenchyma layer
48 Leaf vasculature position relative to palisade parenchyma
49 Lamina margin black and sclerenchymatous
50 Sclerenchyma present on pinna costae or first order veins
51 Sclerenchyma present on first order veins
52 sclerenchyma of midrib visible abaxially
53 sclerenchyma of midrib visible adaxially
54 sclerenchyma of pinna costae visible abaxially
55 sclerenchyma of pinna costae visible adaxially
56 sclerenchyma of veins visible abaxially
57 sclerenchyma of veins visible adaxially
58 vein order
59 secondary vein form [NA if 2ndary veins are unbranched]
60 vein fusion in sterile blades
61 hydathodes
62 Hydathodes cretaceous vs. glabrous [NA taxa without hydathodes]
63 blade scales
64 blade scale color pattern
65 setae present absent
66 Seta color
67 Setaphores
68 setae branching pattern
69 petiole setae
70 adaxial midrib setae
71 rachis setae uniform vs. long and short
72 marginal setae
73 marginal setae distribution
74 adaxial lamina setae
75 abaxial lamina setae
76 adaxial pinna costa setae
77 hairs [distinct from setae]
78 hair branching pattern
79 multicellular hair no of cells
80 hairs modification of cells
81 Gland cell color texture
82 Gland cells secretory
83 soral receptacle
84 sori distinct or confluent
85 soral position [applicable only when sori are uniseriate between axis and margin]
86 distribution of sori
87 mature sorus outline
88 sorus sunken vs. surficial
89 Wall angle of sunken sori [n/a for surficial sori]
90 sori embossed on adaxial side of blade
91 blade margin infolded over sori
92 tissue subtending receptacle
93 paraphyses
94 cells of paraphyses modification
95 receptacular setae
96 sporangial stalk width at midstalk
97 episporangial indument
98 position of episporangial indument
99 localization of episporangial indument
100 spore color
101 spore number
102 spore laesura
103 number of cells in newly shed spores
104 exospore (exine) surface
105 gametophyte form
106 rhizoid developed in first cell division
107 base number
108 Acrospermum ascomes
109 Aromatic odor when dry