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Matrix 16190 of Study 13966

About Citation title: "Systematics and Phylogenetics of Indo-Pacific Luciolinae Fireflies (Coleoptera: Lampyridae) and the Description of new Genera".
About Study name: "Systematics and Phylogenetics of Indo-Pacific Luciolinae Fireflies (Coleoptera: Lampyridae) and the Description of new Genera".
About This study is part of submission 13966 (Status: Published).

Matrices

Title: Ballantyne & Lambkin IndoPacific Luciolinae 2013

Description: Mesquite matrix 436 morphological characters of males, females, and associated larvae, includes 142 Luciolinae species and 1 outgroup taxon

Rows

Taxon Label Row Segments Characters 1?–30
Photuris trivittata  (none) 000{01}0000-10090-00000ca01100
Missimia flavida  (none) 00002110-1030e00001e1211110100
Aquatica ficta  (none) 0000111{01}01030-000000e-00010
Aquatica hydrophila  (none) 00012110c10306000001--01110100
Aquatica lateralis  (none) 00001110-1030-000001--01110100
Aquatica leii  (none) 0000111{01}000{01}-5-00000--00
Aquatica wuhana  (none) 00001110-10{13}0-000000--00010
Curtos costipennis  (none) 000131110uvwjyu0001w0000110100
Curtos okinawanus  (none) 0001311{01}0{01}0{01}-0t000003
Photuroluciola deplanata  (none) 01000110a10210c10000-b00011100
Colophotia concolor  (none) 00001110-100-0-0001u000001010-
Colophotia plagiata  (none) 00001110z100a0-0001v000001010-
Colophotia praeusta  (none) 00001110-100b0s0001-000001010-
Poluninius selangoriensis  (none) 00003111-------0001-110001010-
Pyrophanes appendiculata  (none) 00003110-000---00000-200010101
Pyrophanes beccarii  (none) 00003110w{01}00---0001-0000010
Pyrophanes similis  (none) 000031103000f-p00000y-00010101
Pyrophanes quadrimaculata  (none) 00003110-000---00000-400010101
Pteroptyx gelasina  (none) 00003110-000d-q0001y000001010-
Pteroptyx macdermotti  (none) 00103110-{01}00-0-00000z-00010
Pteroptyx maipo  (none) 00003110-000---00000--0101010-
Pteroptyx malaccae  (none) 000031102000---0001-000001010-
Pteroptyx sp ML  (none) 00003110-100c0-00012000001010-
Pteroptyx sp MFC  (none) 000031110trv-x-000002-0001010-
Pteroptyx tener  (none) 00003110-000--r0001z000001010-
Pteroptyx truncata  (none) 00003110y000---0001-000001010-
Medeopteryx cribellata  (none) 00003110-000e--000{01}00000010
Medeopteryx corusca  (none) 00003110-000---0001-000001010-
Medeopteryx effulgens  (none) 00003110x000---00013000001010-
Medeopteryx platygaster  (none) 0000311{01}1000---0001-0000010
Medeopteryx pupilla  (none) 00003110-10{13}0-00001k0000010
Medeopteryx similispupillae  (none) 00003110-000-2-0001-000001010-
Trisinuata caudabifurca  (none) 000031110heg-w-0001c000001010-
Trisinuata dimidiata  (none) 00003110-000---0001-000001010-
Trisinuata microthorax  (none) 000031111------0001-000101010-
Trisinuata minor  (none) 000031110------0001-000001010-
Trisinuata papuae  (none) 000031110------0001-100101010-
Trisinuata papuana  (none) 00003110w1030s00001-000001010-
Trisinuata similispapuae  (none) 00003120-100-0-0001-000101010-
Australoluciola anthracina  (none) 000031211ehf-a-0001-0000010101
Australoluciola aspera  (none) 00003120-1030-00001-0000010101
Australoluciola australis  (none) 000031111gfh-p-00000xz0{01}{01
Australoluciola flavicollis  (none) 0000311{01}121q--n0001-000{01}
Australoluciola foveicollis  (none) 000031{12}10jkm---0001t0000010
Australoluciola fuscamagna  (none) 000031211kmnh--0001-0000010001
Australoluciola fuscaparva  (none) 000031111mnp---0001-0000010101
Australoluciola nigra  (none) 0000311{01}121r-q-0001s0000010
Australoluciola orapallida  (none) 000031111nps-r-0001-0000010101
Australoluciola pharusaurea  (none) 000031211pqtg--0001-0000010101
Luciola italica  (none) 00002110-10080-00000d-0101011f
Luciola antipodum  (none) 00013110-1030c00001b1001010100
Luciola aquilaclara  (none) 00013110b10300400000a-00010100
Luciola hypocrita  (none) 00010110-100-0-00000-d0{01}010
Luciola kagiana  (none) 00011110d100-0-00000--01110100
Luciola oculofissa  (none) 00013110-1030300001a1001010100
Luciola parvula  (none) 00012110910070300000b-0{01}010
Luciola syriaca  (none) 00001110-1030b000000-e0001011m
Emeia pseudosauteri  (none) 000011110vuayvv00001--02110000
Luciola aegrota  (none) 00003110-1030t-0001x1102110100
Luciola aquatilis  (none) 0000{12}110-101---00000h-01110
Luciola brahmina  (none) 00012110-100-0-00000--02110100
Luciola carinata  (none) 0000{12}110-101---00000--01110
Luciola cerata  (none) 00103110-10304000000--02110100
Luciola cingulata  (none) 00002110-101---00000f-02110100
Luciola cruciata  (none) 00002110-1030-000001-g0211011q
Luciola dejeani  (none) 00013110-10060-00000-f0101011s
Luciola dubia  (none) 00012110-1030-000001--02110100
Luciola owadai  (none) 00001110d10308000001--0111011r
Luciola praeusta  (none) 000031110abd5-200000--01110100
Luciola seriata  (none) 0000{12}110-101---00000--01110
Luciola striata  (none) 00020110-10200-00000--02110100
Luciola substriata J  (none) 0000{12}110e101--w00000g-01110
Luciola substriata F  (none) 0000{12}110-101-7-00000-h0{12}
Jeng matalanga  (none) 00000120-1030-00001-1102110100
Pygoluciola cowleyi  (none) 000031111bac4d50001d0001110000
Pygoluciola guigliae  (none) 10011120-1030-100001k-01110100
Pygoluciola hamulata  (none) 10011120f1030g000001-k02110100
Pygoluciola kinabalua  (none) 10011120-1030f000001--02110100
Pygoluciola qingyu  (none) 00011120g1030-000001-j02110100
Pygoluciola satoi  (none) 10011120-1030h00001f1101110100
Pygoluciola stylifer  (none) 10011120-1030-000001j-02110100
Pygoluciola wittmeri  (none) 10011120-1030-000001m-02110100
Convexa wolfi  (none) 00120110k100-0-00000-t02110100
Pacifica limbatifusca  (none) 000201105100p0z00100-801110100
Pacifica limbatipennis  (none) 00020110-100-0-001005701110100
Pacifica plagiata  (none) 00020110s100-0-001004502110100
Pacifica russellia  (none) 00020110-100-0x00100--02110100
Pacifica salomonis  (none) 00020110t100-0y00100-602110100
Lloydiella japenensis  (none) 00010110-1030900001h1102110100
Lloydiella majuscula  (none) 00010110j100-0-0001-1001110100
Lloydiella uberia  (none) 00010110-100-0800000-v02110100
Lloydiella wareo  (none) 00010110-10{03}0-00001j1102110
Asymmetricata circumdata  (none) 00020110-100-0-00000-n01110100
Asymmetricata ovalis  (none) 00020110-10030600000n-02110100
Pygatyphella eliptaminensis  (none) 00020110q100-1h0101-1002110100
Pygatyphella huonensis  (none) 00020110-100-1-0101m1002110100
Pygatyphella ignota  (none) 000201106100s1-010161002110100
Pygatyphella japenensis  (none) 00020110-100-{01}k0101-1002110
Pygatyphella karimui  (none) 00020110-100-{01}-0101-1{01}02
Pygatyphella kiunga  (none) 00020110r100-1-0{01}01n1002110
Pygatyphella marginata  (none) 00020110-100-1j0101-1002110100
Pygatyphella nabiria  (none) 00020110-100q{01}-0{01}0151001
Pygatyphella obsoleta  (none) 000201104100r{01}-0{01}0141102
Pygatyphella okapa  (none) 00020110-100-{01}m0101q1102110
Pygatyphella peculiaris  (none) 00020110-100-1-010171002110100
Pygatyphella pulcherrima  (none) 00020110-100n1-0101p1002110100
Pygatyphella tagensis  (none) 00020110-100m1-0101-1002110100
Pygatyphella tomba  (none) 00020110-100-{01}-0001g1002110
Pygatyphella uberia  (none) 000{12}0110u1030n00001r110{12}
Pygatyphella undulata  (none) 00020110-100k1-0101-1002110100
Pygatyphella wisselmerenia  (none) 00020110v100-0-0101-1002110100
Aquilonia costata  (none) 000111110cdb-j-00000--01110100
Gilvainsula messoria  (none) 000101108100-07000007-02110100
Gilvainsula similismessoria  (none) 000{12}0110-100-0-00000-u02110
Magnalata carolinae  (none) 00020110h10020-00000--02110100
Magnalata limbata  (none) 000{12}0110-100-0-000008s02010
Magnalata rennellia  (none) 00020110-100z0-00000-r02110000
Atyphella aphrogeneia  (none) 0001{01}110-10{02}0u-000009c00
Atyphella atra  (none) 00020110-100-0d00000--02110000
Atyphella brevis  (none) 00020110-100x0-00000t-02110000
Atyphella conspicua  (none) 00020110m100-0e00000-x01110000
Atyphella dalmatia  (none) 00020110-10{23}0-000001--02110
Atyphella ellioti  (none) 00020110-100-0-00000u-01110000
Atyphella flammans  (none) 000201107100w0-00000-902110000
Atyphella flammulans  (none) 00020110-100-0-00000v-02110000
Atyphella guerini  (none) 000{12}0110-100-0-00000s-02110
Atyphella immaculata  (none) 0002111{01}010{13}0m0000181001
Atyphella inconspicua  (none) 000131111dce---0001-0001110000
Atyphella kirakira  (none) 00020110-100-0900000q-02110100
Atyphella lamingtonia  (none) 00020110-100-0-00000-q02110100
Atyphella leucura  (none) 00020110-100v0a00000p-02110100
Atyphella lewisi  (none) 00010110-100-0-00000-w01110000
Atyphella lychnus  (none) 00020110n100-0f000006-01110000
Atyphella monteithi  (none) 00020110-100u0-00000-y01110000
Atyphella olivieri  (none) 00020110-100-0-00000--02110000
Atyphella palauensis  (none) 00020110p100-0-00000r-02110100
Atyphella scabra  (none) 00020110-100-0g00000--02110000
Atyphella scintillans  (none) 00020110-100t0-00000w-01110000
Atyphella similis  (none) 00010110-1030k0000191001110000
Atyphella testaceolineata  (none) 00020110-100-0b00000-p02110100
Luciola indica  (none) 00003110-000---0001-000001010-
Luciola trilucida  (none) 00003110-100---0001-000001010-
Luciola sp. 8  (none) 00011110-1030-000000--01110101

Columns

Column Character Description
1 PRONOTUM Irregularities in posterolateral areas (Ballantyne and Lambkin 2006, Figs 1, 3 ? 5)
2 Lateral margins: with longitudinal groove delimiting edges
3 Lateral areas of disc: density of punctation
4 Dimensions: Width across posterior portion of pronotum relative to width across elytral humeri
5 Dimensions: PN width/GHW index from below
6 Anterior margin: explanation
7 Anterolateral corners: shape
8 Lateral margins: subparallel sided
9 Lateral margins dimensions : of subparallel sided pronota
10 Lateral margins dimensions : of non subparallel sided pronota
11 Lateral margins: inclination lateral pronotal margins (anterior half) when margins are not subparallel sided
12 Lateral margins: nature of posterior half of margins that diverge in anterior half
13 Lateral margins: nature of divergence in posterior half
14 Lateral margins: where C > A or B, nature of posterior convergence of lateral margins that diverge in anterior half or more
15 Lateral margins: nature of lateral margins that diverge along their length
16 Lateral margins: sinuousity of lateral margins (vertical plane)
17 Lateral margins: indentation
18 Posterolateral corners: irregularities
19 Posterolateral corners: shape
20 Posterolateral corners: angle of rounded posterolateral corners
21 Posterolateral corners: inclination of angulate corners
22 Posterolateral corners: angle of angulate corners
23 Posterolateral corners: projection of posterolateral pronotal corners laterally beyond width of pronotum at B or C
24 Posterolateral corners: extension relative to median posterior margin
25 Posterior margin: emargination
26 Hypomera: nature of hypomera
27 Hypomera: elevation of median hypomeron
28 Hypomera: flattening of anterior hypomeron
29 Hypomera: flattening of posterior hypomeron
30 Hypomera: nature of more widely flattened posterior area of hypomeron
31 Hypomera: flattening of median area
32 Hypomera: nature of entire area
33 Hypomera: nature of widened flattened posterior part of hypomeron
34 ELYTRON Punctation: size
35 Elytron punctation: density
36 Elytron punctation: linearity
37 Elytron apex: dorsal surface elytral apex dimpled
38 Elytron apex: elytral apex deflexed
39 Elytron apex: shape outline deflexed apex
40 Elytron apex: length of truncate elytral apex
41 nature of inner margin (C) of truncate elytral apex
42 Nature of inner junction between truncate elytral apex margins B and C
43 Elytron apex: Posterior end of non deflexed elytral apex appearing emarginate with small hook along outer edge at end of epipleuron
44 Elytron Ventral aspects: extent of epipleuron
45 Elytron Ventral aspects: extent of sutural ridge
46 Elytron Ventral aspects: visibility of humerus from below (in horizontal specimen)
47 Elytron Ventral aspects: epipleuron at elytral base narrow, not widely expanded
48 Elytron visibility of anterior lateral margin of epipleuron relative to posterior edge of MS (specimen horizontal)
49 Elytron Ventral aspects: Lateral margin of epipleuron developed as a ridge
50 Elytron Ventral aspects: where elytral apex is not deflexed epipleuron and sutural ridge continue around elytral apex (view from above)
51 Elytron apex: nature of epipleural ridge at apex (view from beneath)
52 Elytron apex: nature of epipleural ridge at apex (view from beneath)
53 Elytron apex: thickening epipleuron and suture in apical half
54 Elytron interstitial line: development
55 Elytron interstitial lines exceed suture
56 Elytron Margins: approximation of sutural margins in closed elytra
57 Elytron margins: shape
58 HEAD: depression of vertex
59 Head: exposure as retracted into prothoracic cavity
60 Head: approximation of eyes ventrally
61 Head: eye excavation (visible when head is retracted)
62 Head: extent of emargination
63 Head: proximity antennal sockets
64 Head: proximity of antennal sockets that are not contiguous or almost so
65 Head: frons vertex junction
66 Head: median area frons vertex junction
67 Head: position of antennal sockets
68 Head: presence of clypeolabral suture
69 Head: position of clypeolabral suture
70 Head: nature of clypeolabral suture
71 Head: nature of sclerotisation of labrum
72 Head: nature of anterior margin of inflexible labrum
73 Head: length/width labrum
74 Head: proximity of eyes above labrum SIW/GHW
75 Mouthparts: functionality
76 Mouthparts: shape apical segment labial palp
77 Mouthparts: nature of non lunate labial palp apical segment -lateral compression
78 Mouthparts: shape of laterally compressed apical segment labial palp
79 Mouthparts: length of narrow triangular labial palp apical segment
80 Mouthparts: length of triangular labial palp apical segment/length of maxillary palp apical segment
81 Mouthparts: nature of inner margin of strongly flattened labial palp apical segment
82 Mouthparts: nature of inner margin of labial palpi that are irregular or dentate
83 Mouthparts: nature of apical segment of labial palp that is not strongly flattened
84 Mouthparts: Length of ovoid/fusiform palp
85 Antennae: length
86 Antennae: length FS 1
87 Antennae: expansion FS1
88 Antennae: median area FS1
89 Antennae: expansion FS4 at anterior apical angle
90 Antennae: FS 2 - 8
91 Antennae: FS 7 - 9
92 Antennae: number of segments
93 Antennae: antennal segments
94 Antennae: pedicel shape
95 LEGS: inner tarsal claw
96 Legs: MFC
97 Legs: femora 2 swollen
98 Legs: femora 3 swollen
99 Legs: femora 3 curved
100 Legs: basitarsus 2 excavated
101 Legs: swelling tibiae 3
102 Legs: curvature tibiae 1
103 Legs: curvature tibiae 2
104 Legs: curvature of tibiae 3
105 ABDOMEN VENTRITES V8: presence
106 V 3: shape posterior margin
107 V 4: shape posterior margin
108 V3, V4: extension anterior margin V4 into V3
109 V7: nature of posterior half
110 V7: nature of corners in arched or swollen posterior half
111 V 7: shape of corners in arched or swollen posterior half
112 V 7: intrusion arched or swollen posterior half of V7 into light organ
113 V 7: depth of intrusion of swollen posterior part of V7 into light organ
114 V7: presence median carina
115 V7: width median carina
116 V7: height median carina
117 V7: median longt trough ventral face of
118 V7: nature of surface
119 V7: Posterior margin trisinuate
120 V7: emarginations of trisinuate posterior margin with further developments
121 V7: presence incurving lobes along V7 posterior margin in emarginations of trisinuate posterior margin of V7
122 V7: Nature of apex of incurving lobes
123 V7: shape of incurving lobes
124 V7: incurving lobes club like (apex wider than base)
125 V7: nature of anterior margin of incurving lobes
126 V7: nature of posterior margin of incurving lobes
127 V7: presence pointed projections along V7 posterior margin in emarginations of trisinuate V7
128 V7: presence dimple
129 V7: dorsal surface reflexed into paired lobes
130 V7: lateral margins dorsal surface reflexed and enclosing sides of T8
131 cuticular remnants dorsal to sheath
132 cuticular remnants ventral to sheath
133 V7: apex symmetrical tergite 8 engulfing posterior margin
134 V7: lateral margins of symmetrical tergite 8 enfold V7 at sides
135 LO: Presence of light organ
136 LO V7: entire or bipartite
137 LO V7: nature posterior margin entire LO
138 LO V7: depth of posterior emargination of entire light organ
139 LO V7: shape of entire LO
140 LO V7: shape of entire LO
141 LO: bipartite light organ in 7 reaches sides of V7
142 LO: bipartite light organ in V7 reaches into PLP
143 LO: inner margins of bipartite light organ contiguous
144 LO: entire light organ in V7 reaches sides
145 LO: entire light organ in V7 reaches posterior margin (may be narrow clear area)
146 LO: entire light organ in V7 reaches into MPP
147 LO: entire light organ in V7 reaches into PLP
148 LO V7: nature anterior margin entire light organ V7
149 LO V7: emarginate anterior margin due to muscle attachment on sclerties surrounding aedeagal sheath
150 LO V7: depth of emargination
151 LO: width of emargination
152 LO: presence in V6
153 LO: size light organs V6
154 LO: size bipartite light organs V7
155 LO: size entire light organs V7
156 LO: median depression on anterior face of entire light organ in V7
157 MPP: presence
158 MPP: symmetry of apex
159 MPP: medial emargination apex (visible from below in horizontal specimen)
160 MPP: depth emargination
161 MPP: shape non medianly emarginate apex
162 MPP strongly laterally compressed (length greatly exceeds width)
163 MPP posterior margin bisinuate
164 MPP: L/W
165 MPP: length MPP in relation to PLP
166 MPP: width MPP in relation to PLP
167 MPP: dorsal inclination narrowed MPP (L > W)
168 MPP: dorsally inclined MPP bifurcate into well defined projections
169 MPP: relation of MPP or posterior margin of V7 to apex of symmetrical tergite 8
170 MPP: basal emarginations
171 MPP: with 2 hooks at apex
172 MPP: presence of median dorsal ridge
173 MPP: nature of dorsal ridge
174 MPP: width dorsal ridge
175 MPP: Dorsal ridge with 2 elevated median projections
176 MPP: median longt trough V surface
177 PLP: development
178 PLP: length
179 PLP: width (measured brom below)
180 PLP: L/W (width measured from below)
181 PLP: width of PLP that are considerably produced
182 PLP: inclination
183 PLP: width of obliquely inclined PLP (view from side if necessary)
184 ABDOMEN TERGITES. T8: symmetry
185 T8: ventral inclination of symmetrical apex
186 T8: width of ventral inclination of symmetrical apex that is strongly inclined ventrally
187 T8: width of ventral inclination of symmetrical apex that is not strongly narowed in posterior half
188 T8: thickness of ventral inclination of symmetrical apex that is not strongly narrowed in posterior 1/2
189 T8: Lateral margins enfold lateral margins of apex of V7
190 T7: Posterolateral corners of T7 slightly prolonged and overlap V7 at their edges
191 T8: prolongation posterolateral corners
192 T8: angle of prolongation of posterolateral corners
193 T8: length of prolongations of posterolateral corners
194 T8: presence of median trough
195 T8: length median trough
196 T8: symmetry of margins ventral trough
197 T8:nature of margins of median groove ventral surface
198 T8: Nature of anterior end of well defined ridged margins of median groove on tergite 8
199 T8: symmetry of flanges
200 T8: nature of anterior end of ridged margins that is not produced
201 T8: Nature of anterior end of ridged margins if produced (=flanges)
202 T8: width of flanges
203 T8: length/width flanges
204 T8: nature of tip of flanges
205 T8: orientation flanges
206 T8: presence lateral depressed lateral spiny and hairy troughs
207 T8: ventral face with asymmetrical projections
208 T8: presence of median posterior ventral ridge
209 T8: curvature of median posterior ventral ridge
210 T8: width (Posterior visible portion)
211 T8: POSTERIOR, ENTIRE (visible) PORTION OF T8 aubparallelsided along its length
212 T8: nature of margins of visible posterior area symmetrical tergite 8
213 T8: length concealed anterior portions
214 T8: narrowing (appears laterally emarginate) before origin of elongate anterolateral projections
215 T8: width concealed anterior portions in horizontal plane
216 T8: width anterior prolongations in vertical plane
217 T8: inner area of anterior area of prolongations with bifurcation
218 T8: emargination of median posterior margin
219 T8: paired lobes AT SIDE OF MEDIAN EMARGINATION
220 AEDEAGAL SHEATH. Sheath sternite: with elongate darkened area in posterior median area
221 Sheath sternite: inclination of dorsal ridge/ darkened area
222 Sheath sternite: length of dorsal ridge/darkened area
223 Symmetrical sheath sternite: posterior to tergite articulations tapering evenly on both sides to narrow apex
224 Aedeagal sheath symmetry (posterior to tergite articulations) (due to emarginations of sternite)
225 Sheath sternite: posterior to tergite articulations with lateral emargination/s
226 Sheath sternite: extent of lateral emarginations
227 Sheath sternite: level of emargination on right side
228 Sheath sternite: nature of lateral emargination on L side
229 Sheath sternite: emargination of both sides of lateral margins posterior 1/3
230 Sheath sternite: nature of lateral emarginations on R side beginning at tergite articulations
231 Sheath sternite: Nature posterior to tergite articulations
232 Sheath sternite: extent of subparallel sided sheath sternite
233 Sheath sternite: terminates in separate hairy lobe/s
234 Sheath sternite: posterior margin emarginate
235 Sheath sternite: emargination occurs between separate apical lobes
236 Sheath sternite: position of posterior emargination
237 Sheath sternite: width of emargination of posterior margin
238 Sheath sternite: width of emargination of posterior margin
239 Sheath sternite: depth of emargination of posterior margin
240 Sheath sternite:position of narrow emargination of posterior margin
241 Sheath sternite: presence of pointed projections to either side of median emargination of posterior margin
242 Sheath sternite: nature of posterior margin that is not medially emarginate
243 Sheath sternite: anterior portion (anterior to lateral tergite articulations) bent to right
244 Sheath sternite: posterior portion (posterior to lateral tergite articulations) bent to right
245 Sheath sternite: maximum width occurs at tergite articulations
246 Sheath sternite:nature of maximum width at tergite articulations
247 Sheath sternite:nature of sternite that reaches its maximum width posterior to tergite articulatios
248 Sheath sternite: nature of expansion in posterior 1/2 - 2/3
249 Sheath sternite: nature of anterior half
250 Tergite 9: presence of bulbous paraprocts
251 AEDEAGAL SHEATH: dimensions: length/width
252 tergite 9: elongate anterolateral arms of tergite 9 visible at sides of symmetrical sternite
253 tergite 9: extent of visibility of narrowed arms
254 Tergite 9: split into two pieces
255 Tergite 9: posterior margin with downwardly projecting pieces
256 Tergite 9: posterolateral areas narrowly project beyond the posterior margin
257 Tergite 9: presence of transverse band anterior to sheath tergite
258 Tergite 9: shape of median area anterior margin
259 Tergite 9: shape of median area anterior margin
260 AEDEAGUS: dimensions: L/W
261 Aedeagus dimensions: width across LL/ML
262 Aedeagus: apices of LL visible at sides of ML from above or below
263 Aedeagus dimensions: length LL where apices can be seen from beneath relative to ML measured from beneath
264 Aedeagus dimensions: equality of length LL
265 Aedeagus dimensions: length LL where apices cannot be seen from below
266 Aedeagus: symmetry
267 Aedeagus: asymmetry due to curvature of ML in horizontal plane
268 Aedeagus: asymmetry due in part to curvature of LL
269 ML: asymmetry due to asymmetrical projections of ML
270 ML: nature of ventral face of asymmetrical ML
271 ML: swollen along at more than half its length
272 ML: presence of lateral teeth
273 ML: width of lateral teeth
274 ML: shape of lateral teeth
275 ML: inclination apex
276 ML: strongly arched
277 ML: extent preapical ventral area
278 ML: Asymmetrical flap likeprojection to left side
279 ML: width of symmetrical ML at apex
280 ML: apex with arrowhead shape
281 ML: bulbous tip
282 ML: base bulbous, at least twice as wide as remainder
283 LL: separation of LL visible from beneath
284 LL: separation of LL along their inner dorsal length
285 LL: extent of separation of LL that cannot be seen from below as a proportion of their dorsal length (measured from base of LL)
286 LL: extent of separation of LL which are separated by < 1/2
287 LL: visibility from beneath of LL at sides of symmetrical ML
288 LL: visibility from beneath of LL at sides of asymmetrical ML
289 Aedeagus dimensions: length of LL that cannot be seen from below as a proportion of the median lobe length (measured along dorsal side from base of LL)
290 LL: length of LL as proportion of ML length (Measured from above from base of LL)
291 LL: extent of divergence along inner dorsal margins LL
292 LL: width LL at apices relative to ML apex
293 LL: separation at their apices of LL which are not visible from beneath
294 LL: Apices visible from beneath broadly and evenly expanded in horizontal plane
295 LL: visible at sides of ML with broad basal portion and narrowed apical portion
296 LL: width of apical pieces
297 LL: length of apical pieces
298 LL: extent of separation of basal portion of character 295
299 LL: Base of LL (ventral aspect) produced into lobes that may overlap base of ML
300 LL: anterior margin of dorsal base of LL produced
301 LL: anterior margin of dorsal base of LL asymmetrically produced
302 LL: excavation of anterior margin of base
303 LL: nature of anterior projection of base of LL
304 LL: elongate sausage shaped lobes on ventral surface
305 LL: ANTERIOR END INNER VENTRAL SURFACE LL EXTENDING INTO FLATTENED AND APICALLY ACUTE PROJECTION
306 LL: lateral appendages of LL
307 LL: preapical lateral margin LL produced
308 LL: shape of expanded preapical lateral margin
309 LL production of inner apical margin LL REWORD FOR CLARITY
310 LL length inner apical margin LL
311 LL: width of inner apical production
312 LL: prepical inner margin LL obliquely truncate (inner margin is shorter)
313 LL: inner preapical margin of LL that are obliquely truncate on theior inner margins with tooth
314 LL: whole of apices of LL outturned (Not narrowed inner portion)
315 LL: shape of outer margin of outturned apices of LL
316 EXPAND LL: apices of LL inturned (meet or nearly so in midline)
317 LL: apices incline inwards and hooked
318 LL: nature of inturned hooked apices
319 LL: projection on left LL
320 LL: Inner margins of LL approach closely at half their length from base
321 LL: longitudinal emarginate hair bearing area on outer margin LL
322 LL: inner dorsal margins of separated LL toothed
323 BP: strongly sclerotised
324 BP: shape
325 BP: with emarginated anterior margin
326 BP: width when viewed from beneath
327 FEMALE Pronotum: pronotal irregularities
328 Pronotum: punctation
329 Pronotum: dimensions
330 Pronotum: width across posterior PN margin/ width across elytral humeri
331 Pronotum: Indentation of posterolateral margin
332 Pronotum: irregularities at posterolateral corner
333 Elytra: punctation
334 Elytra: density of punctation
335 Elytra: development of fore wings
336 Wings: development of hind wings
337 Elytra: Interstitial line development
338 Elytra: carina
339 Head: size of eyes
340 Head: position of antennae
341 Head: nature of anterior prolongation of head
342 Head: clypeolabral suture
343 Legs: tibiae 3 curved
344 Legs: tibiae 2 curved
345 Legs tibiae 1 curved
346 Light organ: extent
347 Light organ: extent of LO that is confined to V6
348 elevations on tergite 8
349 median ridge V7
350 lateral elevated areas V7
351 lateral depressed areas V7
352 Bursa plates: presence
353 Bursa plates; number of pairs of plates
354 single pair of bursa plates appear as fine strips
355 single pair of bursa plates appear as hooks
356 bursa plates appear as wide (paired) plates
357 Presence of plate at base of median oviduct
358 Nature of SDG
359 Female genitalia: sizo of apical segment of styli to preapical segment
360 LARVA nature of lateral tergal margins
361 Nature of non explanate tergal margins
362 Lateral tergal margins covering laterotergites from above
363 laterotergites visible from above
364 lateral margins of thoracic and abdominal terga of differing shapes
365 Posterolateral corners of abdominal terga prolonged
366 Prolonged posterolateral corners of abdominal terga single or bifurcate
367 Angle of single prolonged corners
368 L/W protergum
369 nature of tergal margins
370 antennal segment 3 surmounted by hairs
371 nature of sense cone
372 length of sense cone
373 single pair of tubercles along anterior margin protergum
374 Protergum with tubercle like projections along posteriormargin only
375 dorsal surface of protergum with tubercles along anterior, lateral and posterior margins
376 shape of posterolateral corners protergum
377 shape of posterolateral pronotal corners that are not produced
378 shape posterolateral corners protergum that are narrowly produced
379 shape posterolateral corners pronotum that are narrowly produced
380 shape posterolateral corners meso and metaterga
381 inclination of narrowly produced posterolateral corners of meso andmetaterga
382 Median line with lateral margins
383 presence of ridged margins of median line
384 Median line extending to anterior and posterior margins of most terga
385 size punctures in anterior half terga 2 - 10
386 extent posterolateral corners tergum 12
387 posterior margins of abdominal terga 1 - 7 or 8 with projections to either side of median line
388 width of projections
389 nature of projections along posterior margins of terga 1 - 7 or 8
390 presence of brush hairs around tarsal claw
391 presence thickened lateral margin protergum
392 mandibles with inner teeth
393 Position of sensory organs on apical palpomere
394 L/W apical maxillary palpomere
395 tergal plates sclerotised or atleast coloured to lateral margins
396 Tergal plates heavily sclerotised
397 Ventral body lacking membranous areas
398 posterolateral corners of penultimate tergite broadly produced posteriorly
399 mode of life
400 tracheal gills
401 presence of tergum 8 spiracles
402 Presence of glands at sides of abdomen on eversible sacs
403 Presence of laterosternal plates to either side of median sternal plate in abdominal segments 1 - 8
404 filaments along posterior margins most terga
405 Ventral surface abdomen: posterior margin sternite with pried projections
406 Ventral surface abdomen: posterior margin laterosternites narrowly prolonged
407 ventral surface abdomen: posterolateral areas of laterotergites prolonged
408 COLOUR MALE: dorsal colouration male (pronotum and elytra only)
409 nature of concolourous dorsal colour
410 Elytral colour male: concolourous
411 elytral colour: nature of concolourous elytra
412 colour of pronotum male
413 nature of concolourous pronotum
414 Nature of colour on non concolorous pronotum
415 elytra dark with paler randomly scattered spots
416 elytra striped male (pale stripes in otherwise dark elytra))
417 number of pale interstitial lines in striped elytra
418 extent of pale lateral margin in striped elytra
419 fat body contributes to pale line colour in striped elytra
420 striped elytra with additional stripes due to lines of fat body
421 Base of largely pale elytra with dark area over humerus
422 extent of dark elytral markiing adjacent to MS
423 base of largely pale elytra with dark area adjacent to MS
424 base of largely pale elytra with dark area extending across base from humerus to MS
425 pale elytra with single apical dark area
426 Elytra with pale margin/s (different colour)
427 extent of pale elytral margins
428 Elytra with base much paler than rest
429 colour tergite 8 male
430 COLOUR FEMALE colour elytra female
431 Elytra with base much paler than rest
432 Nature of colour on non concolourous pronotum
433 nature of dorsal colouration female
434 type of crypsis female
435 dorsal colouration pattern male
436 types of crypsis male