TreeBASE Search-Matrix 17099 of study 14253

@ARTICLE{TreeBASE2Ref22012, author = {Daniel Whitmore and Thomas Pape and Pierfilippo Cerretti}, title = {Phylogeny of Heteronychia: the largest lineage of Sarcophaga (Diptera: Sarcophagidae)}, year = {2013}, keywords = {character analysis ? cladistics ? flesh flies ? male terminalia ? morphology ? Palaearctic ? terminology}, doi = {10.1111/zoj.12070}, url = {http://}, pmid = {}, journal = {Zoological Journal of the Linnean Society}, volume = {169}, number = {3}, pages = {604--639}, abstract = {Sarcophaga Meigen is one of the megadiverse genera of true flies, with approximately 850 valid species worldwide. The genus is divided into about 160 subgenera, the validity of a vast majority of which has never been verified using cladistic methods. This paper deals with the mainly Palaearctic subgenus Heteronychia Brauer & Bergenstamm, which comprises 89 species and is thus the largest subunit of Sarcophaga. We performed a cladistic analysis of the group based exclusively on male morphological characters. Parsimony analyses were run on a matrix of 84 characters for 88 species. Species of the subgenera Discachaeta Enderlein and Notoecus Stein were also included in the matrix. A further analysis was carried out using a subset of characters from the terminalia alone (70 characters). The results show that the clade formed by Heteronychia, Discachaeta, and Notoecus is monophyletic, with Discachaeta emerging as polyphyletic whereas Sarcophaga (Notoecus) longestylata Strobl is nested within the Sarcophaga filia-group. Character states supporting Heteronychia and the few well-supported species-groups are discussed in detail. The following synonymies are proposed: Discachaeta = Heteronychia (syn. nov.) and Notoecus = Heteronychia (syn. nov.). The paper also includes a historical background of the taxon in relation to the classfication of the genus Sarcophaga over the past two centuries, as well as a terminological review of the male terminalia, particularly of the distiphallus.} }

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Matrix 17099 of Study 14253

Citation title: "Phylogeny of Heteronychia: the largest lineage of Sarcophaga (Diptera: Sarcophagidae)".

Study name: "Phylogeny of Heteronychia: the largest lineage of Sarcophaga (Diptera: Sarcophagidae)".

This study is part of submission 14253 (Status: Published).

Matrices

Title: Genitalia matrix

Rows

Taxon Label	Row Segments	Characters 1?–30
Sarcophaga variegata	(none)	040000002010000000015001230000
Sarcophaga agnata	(none)	24102000210000101?235101105110
Sarcophaga longestylata	(none)	1010000011003010101421101141D0
Sarcophaga amita	(none)	221001001100101000110000101140
Sarcophaga amputata	(none)	2?1112111010001011110000001120
Sarcophaga arcipes	(none)	221100010200001000110000221141
Sarcophaga bezziana	(none)	221001010100101000111000101160
Sarcophaga cucullans	(none)	211112001100001000110000221131
Sarcophaga esthera	(none)	211112001100001000110000221131
Sarcophaga kunonis	(none)	201002000010201011020101001120
Sarcophaga pumila	(none)	211110?01200001000110000121161
Sarcophaga abramovi	(none)	1?1000010200001110111000201130
Sarcophaga anatolica	(none)	101001001100201011111000201140
Sarcophaga ancilla	(none)	1010010012000{01}1011111000201
Sarcophaga armeniaca	(none)	2?1001001200001001111000201140
Sarcophaga balanina	(none)	2111020012002010112200000011A0
Sarcophaga belanovskyi	(none)	111001001200001001111000201140
Sarcophaga benaci	(none)	221001011200201011111000202150
Sarcophaga boettcheri	(none)	121011011200301010142100214180
Sarcophaga bulgarica	(none)	221001010200001001111000201160
Sarcophaga chaetoneura	(none)	10101001120{01}001011213100202
Sarcophaga chiquita	(none)	201001010200201000011000201140
Sarcophaga clarahenae	(none)	101001001100001000111000201140
Sarcophaga consanguinea	(none)	221001001201001000111000202140
Sarcophaga croca	(none)	121001010200001011210100201140
Sarcophaga curvifemoralis	(none)	211001001000201011134100201140
Sarcophaga depressifrons	(none)	{12}21001000200001000111000201
Sarcophaga desertorum	(none)	??11?2001100????1121?0002011A?
Sarcophaga dissimilis	(none)	101010011200001011213100202140
Sarcophaga enderleini	(none)	211010011200201000233000202170
Sarcophaga ferox	(none)	22111201{12}200001000220000201
Sarcophaga filia	(none)	211001011100301010142110214180
Sarcophaga filiola	(none)	??1001002000?0????142??0214???
Sarcophaga gabrielei	(none)	211011002100201010103100201170
Sarcophaga gallica	(none)	221001000201001011211000201140
Sarcophaga giganta	(none)	211000002200201010233100302140
Sarcophaga haemorrhoa	(none)	2210010{01}0{23}00001000111000
Sarcophaga haemorrhoides	(none)	2210010{01}0200001000111000201
Sarcophaga helenae	(none)	2210010{01}0200001000111000102
Sarcophaga hellenica	(none)	221010002010001001122000202140
Sarcophaga heptapotamica	(none)	221001001200001000{01}11000201
Sarcophaga infantilis	(none)	121001001100001000111000201150
Sarcophaga infixa	(none)	22100100{01}201001011211000201
Sarcophaga javita	(none)	22100100{01}200011000212000201
Sarcophaga kataphygionis	(none)	221001000200001000111000201160
Sarcophaga kerteszi	(none)	2211120102001010001330002211B0
Sarcophaga kozlovi	(none)	2?10?000200?20???02330002021??
Sarcophaga lacrymans	(none)	101001001201001010050100202110
Sarcophaga lederbergi	(none)	22100100{01}200001000111000101
Sarcophaga lejlekensis	(none)	221001002010011000233000202170
Sarcophaga mediterranea	(none)	211003112000001110140010101120
Sarcophaga metopina	(none)	101001001200001000111000101140
Sarcophaga minima	(none)	2011000{01}1200001000141010101
Sarcophaga monspellensia	(none)	201013112000?01011220100101110
Sarcophaga mutila	(none)	221110011100001000112100102140
Sarcophaga nanula	(none)	121001000200001000111000201160
Sarcophaga nipponensis	(none)	2010100020010{01}1000110000202
Sarcophaga obvia	(none)	2210010002010110112100002021C0
Sarcophaga pandellei	(none)	2210010002000110112100002021C0
Sarcophaga pauciseta	(none)	211021000201201000201000202140
Sarcophaga penicillata	(none)	211003112000001110140110101120
Sarcophaga tangerensis	(none)	2?1001001200011010200000201120
Sarcophaga plotnikovi	(none)	221001000200201001111000202160
Sarcophaga pontica	(none)	121001001100001000111000201140
Sarcophaga porrecta	(none)	20101001{01}200201000233000202
Sarcophaga proxima	(none)	221021001200221000020000202120
Sarcophaga pseudobenaci	(none)	1210110021000{01}1000112000202
Sarcophaga recta	(none)	221021011200001010000100202120
Sarcophaga rondaniana	(none)	{12}{01}1000001201001000050000
Sarcophaga rosellensis	(none)	221021001201201000100000202120
Sarcophaga santospintosi	(none)	211001111200001010140110001130
Sarcophaga schineri	(none)	111000002200011000103000202160
Sarcophaga setinervis	(none)	2211110{01}1{12}000010{01}0112
Sarcophaga shnitnikovi	(none)	211000001100201000233000202170
Sarcophaga sicilia	(none)	101000011100301010142110214180
Sarcophaga siciliana	(none)	221010000010001000202000202140
Sarcophaga siciliensis	(none)	231111002010002110140110101120
Sarcophaga slovaca	(none)	121001000200001000111000201160
Sarcophaga tetrix	(none)	211003110000001100141010101190
Sarcophaga thirionae	(none)	211003112000001110140010101120
Sarcophaga tricolor	(none)	201001012000001010140010101120
Sarcophaga tunisiae	(none)	211001002100201000222000202140
Sarcophaga turana	(none)	1{13}1001011100301010142110214
Sarcophaga uncicurva	(none)	2{01}1003112000001110140110101
Sarcophaga vagans	(none)	211020001101201000211000203140
Sarcophaga vicina	(none)	2110100{01}2200201000233000302
Sarcophaga villeneuveana	(none)	2{01}10031120000011001400{01}0
Sarcophaga violovitshi	(none)	121001000200001011231000201140

Columns

Column	Character Description
1	Sternite 5, setae
2	Syntergosternite 7+8, microtrichosity
3	Epandrium, ventral elongation
4	Cercus, dorsal excavation
5	Cercus, median keel
6	Cercus, subapical hump
7	Cercus, dorsal saddle
8	Distal part of cercus, direction
9	Distal part of cercus, direction
10	Cercus tip, shape
11	Cercus, subapical ventral indentation
12	Cercus, subapical lateral keel
13	Cercus, distribution of setae
14	Cercus, setulae distal ventral surface
15	Cercus, microtrichia
16	Surstylus, microtrichia
17	Pregonite, orientation compared to postgonite
18	Pregonite, preapical twist
19	Pregonite tip, orientation
20	Pregonite tip, shape
21	Pregonite, type and distribution of setae
22	Pregonite, middle section
23	Pregonite, sclerotization
24	Postgonite, tip
25	Phallic tube, basal section
26	Paraphallus, basal part
27	Membrane
28	Vesica, number of lobes
29	Vesica, shape
30	Vesica, microtrichia
31	Harpes, structure
32	Harpes, proximal part (lateral view)
33	Harpes, proximal part - sclerotization
34	Harpes, proximal part - inner surface
35	Harpes, apical process
36	Harpes, apical process - position
37	Harpes, apical process - shape
38	Harpes, apical process - direction
39	Harpes, apical process - median expansion
40	Harpes, apical process - denticles
41	Harpes, microtrichia
42	Lateral styli, volume
43	Lateral styli, spines - number and arrangement
44	Lateral styli, spines - type
45	Lateral styli, length
46	Lateral styli, position with respect to juxta
47	Lateral styli, tip
48	Juxta, general aspect
49	Juxta, length
50	Juxta, orientation of main axis
51	Juxta, dorsal surface lateral view
52	Juxta, structure
53	Juxta, median process - preapical part
54	Juxta, median process - sclerotization
55	Juxta, median process - median suture
56	Juxta, median process - median keel
57	Juxta, median process - general shape and orientation
58	Juxta, median process - lateral margins
59	Juxta, median process - direction of tip
60	Juxta, median process - shape of tip in lateral view
61	Juxta, median process - preapical lateral indentation
62	Juxta, median process - apical tooth-like protuberances
63	Juxta, median process - median depression on tip
64	Juxta, median process - preapical widening
65	Juxta, median process - wide constriction near base
66	Juxta, median process - longitudinal furrows
67	Juxta, median process - tip in dorsal/apical view
68	Juxta, lateral process
69	Juxta, basal process
70	Juxta, connective part