@ARTICLE{TreeBASE2Ref22128,
author = {Luis Norberto Piacentini and M. J. Ramirez and Diana Silva},
title = {Description and relationships of Cauquenia (Araneae: Zoropsidae), with comments on the patterns of evolution of cribellum and male tibial crack on Lycosoidea.},
year = {2013},
keywords = {cladistics, morphology, phylogeny, systematics, taxonomy.},
doi = {10.1071/IS13031},
url = {http://},
pmid = {},
journal = {Invertebrate Systematics},
volume = {27},
number = {},
pages = {567?577},
abstract = {A new genus of the spider family Zoropsidae, Cauquenia, gen. nov., is proposed for Cauquenia maule, sp. nov., from the Maule region in central Chile. The familial placement is tested through the inclusion of Cauquenia in the lastest major morphological analyses of the superfamily Lycosoidea published to date, the subfamily placement of the South American zoropsid genus Itatiaya Mello-Leitão is also tested including them in the Raven and Stumkat (2005) analysis. Cauquenia and Itatiaya are closely related to the African genera Griswoldia Dippenaar-Schoeman & Jocqué and Phanotea Simon, with which it shares a cup-shaped median apophysis on the male pedipalp and tooth-like projections on the lateral lobes of the epigyne in females. The patterns of evolution of cribellum and male tibial crack in Lycosoidea are explored, the cribellum shows up as primitive present, with tree losses and four independt adquicitions, the mail tibial crack is lost twice. Asymmetric cost in cribellum gain:loss of 6:1 produce a primitive cribellum with 12 losses. }
}
Matrix 52024 of Study 14392
Citation title:
"Description and relationships of Cauquenia (Araneae: Zoropsidae), with comments on the patterns of evolution of cribellum and male tibial crack on Lycosoidea.".
Study name:
"Description and relationships of Cauquenia (Araneae: Zoropsidae), with comments on the patterns of evolution of cribellum and male tibial crack on Lycosoidea.".
This study is part of submission 14392
(Status: Published).
Matrices
Title: Lycosoidea
Rows
|
Taxon Label |
Row Segments |
Characters 1?–30 |
| Tengella |
(none)
|
0000000000000?000000000?0000?0 |
| Psechrus |
(none)
|
00001000001??????000000?0100?1 |
| Fecenia |
(none)
|
0000000010000?000001000?0100?0 |
| Udubada |
(none)
|
111?001011000?2?11000110111??? |
| Uduba3 |
(none)
|
111?001011000?2?11000110111??? |
| Zorocrates |
(none)
|
001?000010000?00000100100100?0 |
| Devendra seriatus |
(none)
|
101?0010000010??0000000?000201 |
| Devendra pardalis |
(none)
|
101?0010000010??0000000?000200 |
| Campostichomma |
(none)
|
10110010100010??000000100100?0 |
| Raecius |
(none)
|
111?0010000010??0001000?0000?0 |
| Zorodictyna |
(none)
|
110?001100000?000000100?000200 |
| Takeoa |
(none)
|
011?000000000?000000100?0200?0 |
| Acanthoctenus |
(none)
|
01000000000010??0000000?0100?1 |
| Zoropsis France |
(none)
|
1111000000000?010000100?0301?0 |
| Zoroposis Canary I |
(none)
|
1111000000000?010000100?0001?0 |
| Kilyana hendersoni |
(none)
|
1111000001000?0100000110011??? |
| Birrana bulburin |
(none)
|
1111000001000?010001100?011??? |
| Krukt piligyna |
(none)
|
1111000101000?010000100?0101?0 |
| Megateg elegans |
(none)
|
1111000001000?010001100?0401?1 |
| Huntia deepensis |
(none)
|
11110001000011??0000010?0000?0 |
| Uliodon NZ |
(none)
|
01110000001??????000100?0000?0 |
| Amauropelma truel. |
(none)
|
00000000000110??1000000?000210 |
| Griswoldia robusta |
(none)
|
100?0000000011??0000000?000201 |
| Griswoldia urbensis |
(none)
|
11010000000011??0000000?000200 |
| Phanotea pering |
(none)
|
11100000100011??0000000?020200 |
| Phanotea spX |
(none)
|
11100000000111??0000000?020210 |
| Phanotea spA |
(none)
|
11100000000111??0000000?030210 |
| Senoculus |
(none)
|
000?000010000?000000000?011??? |
| Mituliodon tarantul. |
(none)
|
0000000010000?00000001110100?1 |
| Dolomedes |
(none)
|
0000000010000?00001001101100?1 |
| Pisaura |
(none)
|
0000000010000?00001101101100?1 |
| Trechalea |
(none)
|
0000010011000?1?021001101100?0 |
| Rhoicinus |
(none)
|
0000110010000?1?02100110110221 |
| Hogna |
(none)
|
0000100010000?2?121001101100?0 |
| Sosippus |
(none)
|
0000100010000?2?121001111100?0 |
| Phoneutria |
(none)
|
00000000000011??0000000?000200 |
| Ctenus |
(none)
|
00000001000011??0001000?020200 |
| Stiphidion |
(none)
|
00000010101??????000000?0100?1 |
| Tapinillus |
(none)
|
0000000010000?000000000?011??? |
| Austrotengella |
(none)
|
0000001000000?000000000?000200 |
| Bengalla spV |
(none)
|
0010001000000?2?00000111010201 |
| Miturga |
(none)
|
0000000010000?11120100110101?1 |
| Diaprograpta |
(none)
|
0000000010000?11000001100100?1 |
| Argoctenus Q4 |
(none)
|
0000000010000?00000000110100?1 |
| Cauquenia maule |
(none)
|
11110011000011??0200000?000200 |
| Pseudoctenus |
(none)
|
11110000000010??0001000?010200 |
| Itatiaya modesta |
(none)
|
111?0010000011??0000000?000220 |
Columns
| Column |
Character Description |
|
1
|
Male tibial crack
|
|
2
|
Cymbium dorsally with dense scopulate patch
|
|
3
|
Apical cymbium
|
|
4
|
Male abdominal shield
|
|
5
|
Male palpal tibia with retrolateral apophysis
|
|
6
|
Male palpal tibia with retroapical cuticle unsclerotised
|
|
7
|
Male palpal tibia with ventral apophysis in addition to retrolateral
|
|
8
|
Cymbial dorsobasal projection
|
|
9
|
Subtegulum\tegulum interlocking lobe
|
|
10
|
Separate tegular conductor
|
|
11
|
Median apophysis
|
|
12
|
Median apophysis, position on tegulum
|
|
13
|
Median apophysis shape
|
|
14
|
Concave median apophysis
|
|
15
|
Convex median apophysis
|
|
16
|
Hooked median apophysis
|
|
17
|
Median apophysis angle
|
|
18
|
Tegulum
|
|
19
|
Distar tegular process
|
|
20
|
Tegular lobe process arising near embolous base
|
|
21
|
Paraembolic vane or lamina
|
|
22
|
Embolous base
|
|
23
|
Embolous araising from basal lobe
|
|
24
|
Basal lobe of embolus with process
|
|
25
|
Embolus, direction of curve
|
|
26
|
Embolus
|
|
27
|
Epigyne configuration
|
|
28
|
Lateral lobes, shape
|
|
29
|
Lateral lobes teeth, kind
|
|
30
|
Median sector of epigynum
|
|
31
|
Median lobe
|
|
32
|
Median lobe scape
|
|
33
|
Posterior divot or fossa on scape
|
|
34
|
Shape of copulatory ducts
|
|
35
|
Inner margin of epigynal groove
|
|
36
|
Head of spermatheca
|
|
37
|
Base of spermatheca chambered
|
|
38
|
Base of spermatheca with pronounced lobe
|
|
39
|
Fertilization ducts position
|
|
40
|
PER, shape
|
|
41
|
ALE and PME in line
|
|
42
|
PLE behind PME ratio of PER to OQP less thaan 1.6
|
|
43
|
ALE relative to AME
|
|
44
|
Tapetum
|
|
45
|
Ratio of male tibia I with carapaca width
|
|
46
|
Tarsus, dorsal tricobotrial rows
|
|
47
|
Dense claw tufts obscuring the pretarsus
|
|
48
|
Diamond-Shaped hair cluster below tufts
|
|
49
|
Claws on leg I
|
|
50
|
Scopula on leg I
|
|
51
|
Tarsal organ form
|
|
52
|
trichobothrial base, texture of the hood
|
|
53
|
Femur I with proventral spines
|
|
54
|
Female tibia I, lateral spines
|
|
55
|
Spines on tibia I, female, on raised bases
|
|
56
|
Pairs of ventral spines on tibia I of both sexes
|
|
57
|
Metatarsi I, II no. of ventral spines pairs
|
|
58
|
Male tibia I, dorsal spines
|
|
59
|
Female tibia I, dorsal spines
|
|
60
|
Male metatarsus I or II lateroapical pair of spines
|
|
61
|
Nursery web
|
|
62
|
Egg sac carried on the spinnerets
|
|
63
|
Retrocoxal hymen
|
|
64
|
Female with dorsal spigots on PMS
|
|
65
|
cribellum colulus
|
|
66
|
Trochanter notches
|
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